treatments-xml/data/03/94/87/039487F2F3113E1CFF35F91FA01CDD79.xml

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<mods:title id="8B66EF46CC64047685C609354BA5727E">A second discovery of Diapontonia maranulus Bruce, 1986 (Crustacea: Decapoda: Palaemonidae) in the Caribbean</mods:title>
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<treatment id="039487F2F3113E1CFF35F91FA01CDD79" ID-DOI="http://doi.org/10.5281/zenodo.5593248" ID-GBIF-Taxon="189163598" ID-Zenodo-Dep="5593248" LSID="urn:lsid:plazi:treatment:039487F2F3113E1CFF35F91FA01CDD79" httpUri="http://treatment.plazi.org/id/039487F2F3113E1CFF35F91FA01CDD79" lastPageId="4" lastPageNumber="595" pageId="0" pageNumber="591">
<subSubSection id="C327656FF3113E18FF35F91FA12CDDCC" box="[151,597,1698,1725]" pageId="0" pageNumber="591" type="nomenclature">
<paragraph id="8B8236E4F3113E18FF35F91FA12CDDCC" blockId="0.[151,597,1698,1758]" box="[151,597,1698,1725]" pageId="0" pageNumber="591">
<heading id="D0CA8188F3113E18FF35F91FA12CDDCC" bold="true" box="[151,597,1698,1725]" fontSize="11" level="1" pageId="0" pageNumber="591" reason="1">
<taxonomicName id="4C3D4D67F3113E18FF35F91FA12CDDCC" ID-CoL="4725ec0f-03e8-4a3d-97c8-bb1658306f9d" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[151,597,1698,1725]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="591" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3113E18FF35F91FA12CDDCC" bold="true" box="[151,597,1698,1725]" pageId="0" pageNumber="591">
<emphasis id="B949EAF6F3113E18FF35F91FA2C3DDCD" bold="true" box="[151,442,1698,1724]" italics="true" pageId="0" pageNumber="591">Diapontonia maranulus</emphasis>
Bruce, 1986
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C327656FF3113E18FF35F978A23BDDAF" box="[151,322,1733,1758]" pageId="0" pageNumber="591" type="description">
<paragraph id="8B8236E4F3113E18FF35F978A23BDDAF" blockId="0.[151,597,1698,1758]" box="[151,322,1733,1758]" pageId="0" pageNumber="591">
(
<figureCitation id="13062A61F3113E18FF3DF978A395DDAF" box="[159,236,1733,1758]" captionStart="FIGURE 1" captionStartId="1.[151,249,1873,1895]" captionTargetBox="[156,1389,974,1737]" captionTargetId="figure-349@1.[151,1436,971,1852]" captionTargetPageId="1" captionText="FIGURE 1. Diapontonia maranulus Bruce, 1986, RMNH.CRUS.D.566905. A, carapace and anterior appendages, lateral view; B, anterior appendages, dorsal view; C, carapace, lateral view; D, tailfan, dorsal view, setae omitted. A,B, D, ovigerous female, pocl. 1.8; C, ovigerous female, pocl. 2.1. Scale bar A–C = 2; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4949967" httpUri="https://zenodo.org/record/4949967/files/figure.png" pageId="0" pageNumber="591">Figs. 1</figureCitation>
, 
<figureCitation id="13062A61F3113E18FF58F978A266DDAF" box="[250,287,1733,1758]" captionStart="FIGURE 2" captionStartId="2.[151,249,1876,1898]" captionTargetBox="[157,1436,220,1788]" captionTargetId="figure-80@2.[151,1436,215,2032]" captionTargetPageId="2" captionText="FIGURE 2. Dactylus right third pereiopod, median aspect. A, Diapontonia maranulus Bruce, 1986, ovigerous female pocl. 1.8, RMNH.CRUS.D. 566905; B, Periclimenes milleri Bruce, 1986, female pocl. 2.8, RMNH.CRUS.D.36342; C, Periclimenes ingressicolumbi Berggren &amp; Svane, 1989, ovigerous female, pocl. 2.3, RMNH.CRUS.D.37738; D, Altopontonia disparostris Bruce, 1990, ovigerous female paratype, pocl. 3.1, RMNH.CRUS.D.37690. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4949971" httpUri="https://zenodo.org/record/4949971/files/figure.png" pageId="0" pageNumber="591">2A</figureCitation>
, 
<figureCitation id="13062A61F3113E18FE89F978A240DDAF" box="[299,313,1733,1758]" captionStart="FIGURE 3" captionStartId="3.[151,249,1160,1182]" captionTargetBox="[318,1268,185,1139]" captionTargetId="figure-385@3.[318,1268,185,1139]" captionTargetPageId="3" captionText="FIGURE 3. Host echinoid Paleopneustes tholoformis Chesher, 1968, with Diapontonia maranulus Bruce, 1986 shrimps (indicated by circles) photographed with Curasub at 220 m depth off the SE coast of Curaçao. Photo by Adriaan (Dutch) Schrier." figureDoi="http://doi.org/10.5281/zenodo.4949973" httpUri="https://zenodo.org/record/4949973/files/figure.png" pageId="0" pageNumber="591">3</figureCitation>
)
</paragraph>
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<subSubSection id="C327656FF3113E18FF35F8B3A1C7DC52" box="[151,702,1805,1827]" pageId="0" pageNumber="591" type="reference_group">
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<taxonomicName id="4C3D4D67F3113E18FF35F8B3A122DC52" ID-CoL="4725ec0f-03e8-4a3d-97c8-bb1658306f9d" authority="Bruce, 1986: 57 - 68" authorityName="Bruce" authorityPageNumber="57 - 68" authorityYear="1986" box="[151,603,1805,1827]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="591" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3113E18FF35F8B3A2F0DC53" box="[151,393,1805,1827]" italics="true" pageId="0" pageNumber="591">Diapontonia maranulus</emphasis>
Bruce, 1986: 57–68
</taxonomicName>
, figs 1–7.
</paragraph>
</subSubSection>
<subSubSection id="C327656FF3113E1BFF35F8EDA11FDDDC" lastPageId="3" lastPageNumber="594" pageId="0" pageNumber="591" type="materials_examined">
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<emphasis id="B949EAF6F3113E18FF35F8EDA2F6DC18" bold="true" box="[151,399,1872,1897]" pageId="0" pageNumber="591">Material examined</emphasis>
.
</emphasis>
2 ovigerous females (pocl 1.8 and 2.1 (GenBank acc. nr. 
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<collectionCode id="ED2CAE21F3113E18FAAAF8EDA64FDC18" box="[1288,1334,1872,1897]" country="Russia" name="Kotel'nich Museum" pageId="0" pageNumber="591">KM</collectionCode>
921671
</accessionNumber>
)) 
<collectionCode id="ED2CAE21F3113E18FF35F8C9A388DCFC" box="[151,241,1908,1933]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="0" pageNumber="591" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3113E18FF56F8C9A239DCFC" box="[244,320,1908,1933]" pageId="0" pageNumber="591">CRUS</collectionCode>
.D.56690; stn 
<collectionCode id="ED2CAE21F3113E18FE5AF8C9A141DCFC" box="[504,568,1908,1933]" country="USA" httpUri="http://grbio.org/cool/s9xv-zncp" name="College of the Atlantic, Museum" pageId="0" pageNumber="591">COA</collectionCode>
.24, 
<collectingCountry id="F32A7674F3113E18FDD1F8C9A1A9DCFC" box="[627,720,1908,1933]" name="Curaçao" pageId="0" pageNumber="591">Curaçao</collectingCountry>
Substation, 
<collectingCountry id="F32A7674F3113E18FCD8F8C9A0A3DCFC" box="[890,986,1908,1933]" name="Curaçao" pageId="0" pageNumber="591">Curaçao</collectingCountry>
, 
<collectingCountry id="F32A7674F3113E18FC57F8C9A796DCFC" box="[1013,1263,1908,1933]" name="Curaçao" pageId="0" pageNumber="591">Netherlands Antilles</collectingCountry>
, 
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, 
<quantity id="4CC59B01F3113E18FEE6F825A2EDDCC1" box="[324,404,1944,1969]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="0" pageNumber="591" unit="m" value="220.0">220 m</quantity>
, 
<date id="FF831024F3113E18FE05F825A1FBDCC0" box="[423,642,1944,1969]" pageId="0" pageNumber="591" value="2013-11-08">8 November 2013</date>
, Curasub submersible, on 
<taxonomicName id="4C3D4D67F3113E18FC64F825A78EDCC1" authorityName="Chesher" authorityYear="1968" box="[966,1271,1944,1968]" class="Echinoidea" family="Paleopneustidae" genus="Paleopneustes" kingdom="Animalia" order="Spatangoida" pageId="0" pageNumber="591" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3113E18FC64F825A78EDCC1" box="[966,1271,1944,1968]" italics="true" pageId="0" pageNumber="591">Paleopneustes tholoformis</emphasis>
</taxonomicName>
, collected by Adriaan (Dutch) Schrier, Bruce Brandt, Charles H.J.M. Fransen, Sancia E. 
<collectionCode id="ED2CAE21F3113E18FBB9F801A757DCA2" box="[1051,1070,1980,2003]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="591">T</collectionCode>
. van der Meij and Bastian 
<collectionCode id="ED2CAE21F3113E18FA24F801A6E0DCA2" box="[1414,1433,1980,2003]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="591">T</collectionCode>
. Reijnen.— 6 ovigerous females (pocl. 1.75–2.30), 
<specimenCount id="9D3BFD6DF3103E19FCBFFF2AA0F9DBC1" box="[797,896,151,176]" pageId="1" pageNumber="592" type="male">4 males</specimenCount>
(pocl. 1.50–1.75), 
<specimenCount id="9D3BFD6DF3103E19FBDFFF2AA780DBC1" box="[1149,1273,151,176]" pageId="1" pageNumber="592" type="juvenile">1 juvenile</specimenCount>
(pocl. 1.43) 
<collectionCode id="ED2CAE21F3103E19FF35FF01A38BDBA4" box="[151,242,188,213]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="592" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3103E19FF57FF01A23BDBA4" box="[245,322,188,213]" pageId="1" pageNumber="592">CRUS</collectionCode>
.D.56691; 
<collectingCountry id="F32A7674F3103E19FE16FF01A168DBA4" box="[436,529,188,213]" name="Curaçao" pageId="1" pageNumber="592">Curaçao</collectingCountry>
Substation, 
<collectingCountry id="F32A7674F3103E19FD03FF01A078DBA4" box="[673,769,188,213]" name="Curaçao" pageId="1" pageNumber="592">Curaçao</collectingCountry>
, 
<collectingCountry id="F32A7674F3103E19FCACFF01A083DBA4" box="[782,1018,188,213]" name="Curaçao" pageId="1" pageNumber="592">Netherlands Antilles</collectingCountry>
, 
<geoCoordinate id="EE095023F3103E19FBAAFF01A7E2DBA4" box="[1032,1179,188,213]" degrees="12.084542" direction="north" orientation="latitude" pageId="1" pageNumber="592" precision="1" value="12.084542">12.084542ºN</geoCoordinate>
<geoCoordinate id="EE095023F3103E19FB07FF01A646DBA4" box="[1189,1343,188,213]" degrees="68.898222" direction="west" orientation="longitude" pageId="1" pageNumber="592" precision="1" value="-68.898224">68.898222ºW</geoCoordinate>
, 
<quantity id="4CC59B01F3103E19FAEEFF01A6EEDBA2" box="[1356,1431,188,213]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="1" pageNumber="592" unit="m" value="214.0">214 m</quantity>
, 
<date id="FF831024F3103E19FF35FF62A23EDB89" box="[151,327,223,248]" pageId="1" pageNumber="592" value="2014-03-31">31 March 2014</date>
, Curasub submersible, on 
<taxonomicName id="4C3D4D67F3103E19FDD3FF5DA0E2DB89" authorityName="Chesher" authorityYear="1968" box="[625,923,224,248]" class="Echinoidea" family="Paleopneustidae" genus="Paleopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3103E19FDD3FF5DA0E2DB89" box="[625,923,224,248]" italics="true" pageId="1" pageNumber="592">Paleopneustes tholoformis</emphasis>
</taxonomicName>
, collected by Adriaan (Dutch) Schrier, Cessa Rauch, Bert W. Hoeksema and Kaj van Tienderen.—1 non-ovigerous female 
<typeStatus id="54868846F3103E19FB07FEB9A67FDA6D" box="[1189,1286,260,284]" pageId="1" pageNumber="592" type="paratype">paratype</typeStatus>
(pocl 3.5) 
<collectionCode id="ED2CAE21F3103E19FF35FE9AA38BDA31" box="[151,242,295,320]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="592" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3103E19FF57FE9AA23BDA31" box="[245,322,295,320]" pageId="1" pageNumber="592">CRUS</collectionCode>
.D.36341; off Wood Cay, West End, Grand Bahama Island, 
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, 
<geoCoordinate id="EE095023F3103E19FBC6FE9AA79EDA31" box="[1124,1255,295,320]" degrees="79" direction="west" minutes="01.72" orientation="longitude" pageId="1" pageNumber="592" precision="9" value="-79.02866">79º01.72'W</geoCoordinate>
, 
<quantity id="4CC59B01F3103E19FB51FE9AA60DDA4E" box="[1267,1396,295,320]" metricMagnitude="2" metricUnit="m" metricValue="2.7649999999999997" metricValueMax="3.09" metricValueMin="2.44" pageId="1" pageNumber="592" unit="m" value="276.5" valueMax="309.0" valueMin="244.0">244–309 m</quantity>
, 
<date id="FF831024F3103E19FA22FE9AA269DA14" pageId="1" pageNumber="592" value="1983-06-14">14 June 1983</date>
, Johnson Sea Link Dive JSL-I-1357, on 
<taxonomicName id="4C3D4D67F3103E19FD4CFEF1A762DA15" authorityName="Chesher" authorityYear="1968" box="[750,1051,332,356]" class="Echinoidea" family="Paleopneustidae" genus="Paleopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3103E19FD4CFEF1A762DA15" box="[750,1051,332,356]" italics="true" pageId="1" pageNumber="592">Paleopneustes tholoformis</emphasis>
</taxonomicName>
(as 
<taxonomicName id="4C3D4D67F3103E19FBF2FEF1A6F2DA15" box="[1104,1419,332,356]" class="Echinoidea" family="Eurypatagidae" genus="Palaeopneustes" higherTaxonomySource="CoL" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3103E19FBF2FEF1A6F2DA15" box="[1104,1419,332,356]" italics="true" pageId="1" pageNumber="592">Palaeopneustes tholoformis</emphasis>
</taxonomicName>
), collected by Miller, Askew, Hendler, and Chulamanis.
</paragraph>
<paragraph id="8B8236E4F3103E19FF65FE29A7EAD846" blockId="1.[151,1436,151,896]" pageId="1" pageNumber="592">
<emphasis id="B949EAF6F3103E19FF65FE29A280DADC" bold="true" box="[199,505,404,429]" inLineHeading="true" pageId="1" pageNumber="592" reason="1">Material for comparison.</emphasis>
<taxonomicName id="4C3D4D67F3103E19FDA1FE29A7E1DADC" authority="Berggren &amp; Svane, 1989: 1" authorityName="Berggren &amp; Svane" authorityPageNumber="1" authorityYear="1989" box="[515,1176,404,429]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="ingressicolumbi">
<emphasis id="B949EAF6F3103E19FDA1FE29A028DADD" box="[515,849,404,428]" italics="true" pageId="1" pageNumber="592">Periclimenes ingressicolumbi</emphasis>
<bibRefCitation id="EFAC4B15F3103E19FCF9FE29A7E1DADC" author="Berggren, M. &amp; Svane, I." box="[859,1176,404,429]" pageId="1" pageNumber="592" pagination="432 - 444" refId="ref2814" refString="Berggren, M. &amp; Svane, I. (1989) Periclimenes ingressicolumbi, new species, a pontoniine shrimp associated with deep-water echinoids off San Salvador Island in the Bahamas, and a comparison with Periclimenes milleri. Journal of Crustacean Biology, 9 (3), 432 - 444. http: // dx. doi. org / 10.1163 / 193724089 X 00412" type="journal article" year="1989">
Berggren &amp; Svane, 1989: 
<specimenCount id="9D3BFD6DF3103E19FB28FE29A7E1DADC" box="[1162,1176,404,429]" pageId="1" pageNumber="592" type="male">1</specimenCount>
</bibRefCitation>
</taxonomicName>
male (pocl 2.4) and 1 ovigerous female (pocl 2.3) 
<typeStatus id="54868846F3103E19FE72FE05A145DAA1" box="[464,572,440,464]" pageId="1" pageNumber="592" type="paratype">paratypes</typeStatus>
<collectionCode id="ED2CAE21F3103E19FDE0FE0AA1E4DAA1" box="[578,669,439,464]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="592" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3103E19FD02FE0AA194DAA1" box="[672,749,439,464]" pageId="1" pageNumber="592">CRUS</collectionCode>
.D.37738; off San 
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, 
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<geoCoordinate id="EE095023F3103E19FB50FE0AA611DAA1" box="[1266,1384,439,464]" degrees="74" direction="west" minutes="32.7" orientation="longitude" pageId="1" pageNumber="592" precision="92" value="-74.545">74º32.7'W</geoCoordinate>
, 
<quantity id="4CC59B01F3103E19FAD0FE0AA3D6DA82" metricMagnitude="2" metricUnit="m" metricValue="5.79" pageId="1" pageNumber="592" unit="m" value="579.0">579 m</quantity>
, l 
<date id="FF831024F3103E19FF6FFE61A23DDA84" box="[205,324,476,501]" pageId="1" pageNumber="592" value="1987-05">May 1987</date>
, Johnson Sea Link Dive no. 2039, on 
<taxonomicName id="4C3D4D67F3103E19FD59FE61A75EDA85" authorityName="Chesher" authorityYear="1968" box="[763,1063,476,500]" class="Echinoidea" family="Paleopneustidae" genus="Paleopneustes" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3103E19FD59FE61A75EDA85" box="[763,1063,476,500]" italics="true" pageId="1" pageNumber="592">Paleopneustes tholoformis</emphasis>
</taxonomicName>
(as 
<taxonomicName id="4C3D4D67F3103E19FBFBFE61A6EBDA85" box="[1113,1426,476,500]" class="Echinoidea" family="Eurypatagidae" genus="Palaeopneustes" higherTaxonomySource="CoL" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3103E19FBFBFE61A6EBDA85" box="[1113,1426,476,500]" italics="true" pageId="1" pageNumber="592">Palaeopneustes tholoformis</emphasis>
</taxonomicName>
) and 
<taxonomicName id="4C3D4D67F3103E19FF65FDBDA1EFD969" authority="(A. Agassiz, 1880)" baseAuthorityName="A. Agassiz" baseAuthorityYear="1880" box="[199,662,511,536]" class="Echinoidea" genus="Heterobrissus" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="hystrix">
<emphasis id="B949EAF6F3103E19FF65FDBDA2C1D969" box="[199,440,512,536]" italics="true" pageId="1" pageNumber="592">Heterobrissus hystrix</emphasis>
(A. Agassiz, 1880)
</taxonomicName>
(as 
<taxonomicName id="4C3D4D67F3103E19FD65FDBDA7FDD969" authority="Agassiz, 1880" authorityName="Agassiz" authorityYear="1880" box="[711,1156,511,536]" class="Echinoidea" genus="Archaeopneustes" higherTaxonomySource="CoL" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="hystrix">
<emphasis id="B949EAF6F3103E19FD65FDBDA0A2D969" box="[711,987,512,536]" italics="true" pageId="1" pageNumber="592">Archaeopneustes hystrix</emphasis>
Agassiz, 1880
</taxonomicName>
).— 
<taxonomicName id="4C3D4D67F3103E19FB15FDBDA23FD94C" authority="Bruce, 1986: 1" authorityName="Bruce" authorityPageNumber="1" authorityYear="1986" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3103E19FB15FDBDA6E5D969" box="[1207,1436,512,536]" italics="true" pageId="1" pageNumber="592">Periclimenes milleri</emphasis>
Bruce, 1986: 
<specimenCount id="9D3BFD6DF3103E19FE9AFD99A23FD94C" box="[312,326,548,573]" pageId="1" pageNumber="592" type="female">1</specimenCount>
</taxonomicName>
female 
<typeStatus id="54868846F3103E19FE09FD99A172D94D" box="[427,523,548,572]" pageId="1" pageNumber="592" type="paratype">paratype</typeStatus>
(pocl 2.8) 
<collectionCode id="ED2CAE21F3103E19FD37FD99A196D94C" box="[661,751,548,573]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="592" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3103E19FD51FD99A039D94C" box="[755,832,548,573]" pageId="1" pageNumber="592">CRUS</collectionCode>
.D.36342; off San 
<collectingCountry id="F32A7674F3103E19FBB5FD99A7B7D94C" box="[1047,1230,548,573]" name="Salvador Island" pageId="1" pageNumber="592">Salvador Island</collectingCountry>
, Bahama Islands, 
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, 
<geoCoordinate id="EE095023F3103E19FE86FDFAA2D1D911" box="[292,424,583,608]" degrees="74" direction="west" minutes="32.53" orientation="longitude" pageId="1" pageNumber="592" precision="9" value="-74.54217">74º32.53'W</geoCoordinate>
, 
<quantity id="4CC59B01F3103E19FE17FDFAA178D92E" box="[437,513,583,608]" metricMagnitude="2" metricUnit="m" metricValue="5.27" pageId="1" pageNumber="592" unit="m" value="527.0">527 m</quantity>
, 
<date id="FF831024F3103E19FDADFDFAA1AAD911" box="[527,723,583,608]" pageId="1" pageNumber="592" value="1983-10-22">22 October 1983</date>
, Stn Johnson Sea Link SL-1500, on 
<taxonomicName id="4C3D4D67F3103E19FBDEFDF5A23FD9F4" authority="(A. Agassiz, 1880)" baseAuthorityName="A. Agassiz" baseAuthorityYear="1880" class="Echinoidea" genus="Heterobrissus" kingdom="Animalia" order="Spatangoida" pageId="1" pageNumber="592" phylum="Echinodermata" rank="species" species="hystrix">
<emphasis id="B949EAF6F3103E19FBDEFDF5A616D911" box="[1148,1391,584,608]" italics="true" pageId="1" pageNumber="592">Heterobrissus hystrix</emphasis>
(A. Agassiz, 1880)
</taxonomicName>
, collected by Askew, Kier, Hall and Miller, bottom temp 17.9°C.— 
<taxonomicName id="4C3D4D67F3103E19FBDBFDD1A232D9D9" authority="Bruce, 1990: 1" authorityName="Bruce" authorityPageNumber="1" authorityYear="1990" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="disparostris">
<emphasis id="B949EAF6F3103E19FBDBFDD1A6E5D9F5" box="[1145,1436,620,644]" italics="true" pageId="1" pageNumber="592">Altopontonia disparostris</emphasis>
<bibRefCitation id="EFAC4B15F3103E19FF35FD2DA232D9D9" author="Bruce, A. J." box="[151,331,655,680]" pageId="1" pageNumber="592" pagination="149 - 215" refId="ref2974" refString="Bruce, A. J. (1990) Crustacea Decapoda: 5. Deep-sea palaemonoid shrimps from New Caledonian waters. In: A. Crosnier (Ed.), Resultats des Campagnes MUSORSTOM, 6. Memoires du Museum National d'Histoire Naturelle, 145, 149 - 215. [figs. 1 - 39]" type="journal article" year="1990">Bruce, 1990: 1</bibRefCitation>
</taxonomicName>
ovigerous female 
<typeStatus id="54868846F3103E19FD93FD2DA1E8D9D9" box="[561,657,656,680]" pageId="1" pageNumber="592" type="paratype">paratype</typeStatus>
(pocl 3.1) 
<collectionCode id="ED2CAE21F3103E19FC80FD32A005D9D9" box="[802,892,655,680]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="592" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3103E19FC22FD32A0B4D9D9" box="[896,973,655,680]" pageId="1" pageNumber="592">CRUS</collectionCode>
.D.37690; Station CP/45 (Biocal), 
<collectingCountry id="F32A7674F3103E19FAC5FD2DA275D9BC" name="New Caledonia" pageId="1" pageNumber="592">New Caledonia</collectingCountry>
, 
<geoCoordinate id="EE095023F3103E19FEBEFD09A20DD9BC" box="[284,372,692,717]" degrees="22" direction="south" minutes="47" orientation="latitude" pageId="1" pageNumber="592" precision="925" value="-22.783333">22°47'S</geoCoordinate>
<geoCoordinate id="EE095023F3103E19FE22FD09A293D9BC" box="[384,490,692,717]" degrees="167" direction="east" minutes="15" orientation="longitude" pageId="1" pageNumber="592" precision="925" value="167.25">167°15'E</geoCoordinate>
, 
<quantity id="4CC59B01F3103E19FE58FD09A107D9BA" box="[506,638,692,717]" metricMagnitude="2" metricUnit="m" metricValue="4.575" metricValueMax="4.85" metricValueMin="4.3" pageId="1" pageNumber="592" unit="m" value="457.5" valueMax="485.0" valueMin="430.0">430–485 m</quantity>
depth, 
<date id="FF831024F3103E19FD7EFD09A01FD9BC" box="[732,870,692,717]" pageId="1" pageNumber="592" value="1985-08-30">30.viii.1985</date>
, collected by Richer de Forges, donated by the Museum National d'Histoire Naturelle, Paris through Alain Crosnier.— 1 ovigerous female (pocl 2.8 (GenBank acc. nr. 
<accessionNumber id="946EAB07F3103E19FF5AFD41A203D864" box="[248,378,764,789]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM921672" pageId="1" pageNumber="592">
<collectionCode id="ED2CAE21F3103E19FF5AFD41A25CD864" box="[248,293,764,789]" country="Russia" name="Kotel'nich Museum" pageId="1" pageNumber="592">KM</collectionCode>
921672
</accessionNumber>
)) 
<collectionCode id="ED2CAE21F3103E19FE3AFD41A28AD864" box="[408,499,764,789]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="592" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3103E19FE54FD41A13AD864" box="[502,579,764,789]" pageId="1" pageNumber="592">CRUS</collectionCode>
.D.51028; NORFANZ stn. 20/60 or 69, North Norfolk Ridge, 
<geoCoordinate id="EE095023F3103E19FA83FD41A6E5D864" box="[1313,1436,764,789]" degrees="29" direction="south" minutes="41.84" orientation="latitude" pageId="1" pageNumber="592" precision="9" value="-29.697334">29°41.84'S</geoCoordinate>
<geoCoordinate id="EE095023F3103E19FF35FCA2A25CD849" box="[151,293,799,824]" degrees="168" direction="east" minutes="02.61" orientation="longitude" pageId="1" pageNumber="592" precision="9" value="168.0435">168°02.61'E</geoCoordinate>
; 
<quantity id="4CC59B01F3103E19FE92FCA2A2D6D846" box="[304,431,799,824]" metricMagnitude="2" metricUnit="m" metricValue="3.295" metricValueMax="3.37" metricValueMin="3.22" pageId="1" pageNumber="592" unit="m" value="329.5" valueMax="337.0" valueMin="322.0">322–337 m</quantity>
depth, beam trawl, 
<date id="FF831024F3103E19FD2FFCA2A187D849" box="[653,766,799,824]" pageId="1" pageNumber="592" value="2003-05-14">14.v.2003</date>
, 
<collectionCode id="ED2CAE21F3103E19FCABFC9DA022D846" box="[777,859,800,823]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="1" pageNumber="592" type="Museum">NMNZ</collectionCode>
CR. 9991; don. A.J. Bruce.
</paragraph>
<paragraph id="8B8236E4F3103E19FF65FCF9A11BD8F1" blockId="1.[151,1436,151,896]" pageId="1" pageNumber="592">
<emphasis id="B949EAF6F3103E19FF65FCF9A146D82C" bold="true" box="[199,575,836,861]" pageId="1" pageNumber="592">Comparison with type-material</emphasis>
. 
<materialsCitation id="3B553CB9F3103E19FDEEFCF9A127D8F1" ID-GBIF-Occurrence="3391231302" country="Curaçao" location="Specimens" pageId="1" pageNumber="592" specimenCount="1" specimenCount-female="1">
<location id="8EE2603FF3103E19FDEEFCF9A1BCD82C" LSID="urn:lsid:plazi:treatment:039487F2F3113E1CFF35F91FA01CDD79:8EE2603FF3103E19FDEEFCF9A1BCD82C" box="[588,709,836,861]" country="Curaçao" name="Specimens" pageId="1" pageNumber="592">Specimens</location>
(pocl 1.4–2.3) from 
<collectingCountry id="F32A7674F3103E19FC0CFCF9A772D82C" box="[942,1035,836,861]" name="Curaçao" pageId="1" pageNumber="592">Curaçao</collectingCountry>
are somewhat smaller than the 
<specimenCount id="9D3BFD6DF3103E19FACFFCF9A397D8F1" pageId="1" pageNumber="592" type="female">four females</specimenCount>
of the type-series (pocl 3.5–5.1)
</materialsCitation>
.
</paragraph>
<caption id="DF42666CF3103E19FF35F8ECA00EDCD5" ID-DOI="http://doi.org/10.5281/zenodo.4949967" ID-Zenodo-Dep="4949967" httpUri="https://zenodo.org/record/4949967/files/figure.png" pageId="1" pageNumber="592" startId="1.[151,249,1873,1895]" targetBox="[156,1389,974,1737]" targetPageId="1">
<paragraph id="8B8236E4F3103E19FF35F8ECA00EDCD5" blockId="1.[151,1436,1873,1956]" pageId="1" pageNumber="592">
<emphasis id="B949EAF6F3103E19FF35F8ECA26BDC17" bold="true" box="[151,274,1873,1895]" pageId="1" pageNumber="592">FIGURE 1.</emphasis>
<taxonomicName id="4C3D4D67F3103E19FEB5F8EFA1F5DC16" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[279,652,1873,1895]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3103E19FEB5F8EFA173DC17" box="[279,522,1873,1895]" italics="true" pageId="1" pageNumber="592">Diapontonia maranulus</emphasis>
Bruce, 1986
</taxonomicName>
, RMNH.CRUS.D.566905. A, carapace and anterior appendages, lateral view; B, anterior appendages, dorsal view; C, carapace, lateral view; D, tailfan, dorsal view, setae omitted. A,B, D, ovigerous female, pocl. 1.8; C, ovigerous female, pocl. 2.1. Scale bar A–C = 2; D = 1 mm.
</paragraph>
</caption>
<caption id="DF42666CF3133E1AFF35F8E9A711DCB7" ID-DOI="http://doi.org/10.5281/zenodo.4949971" ID-Zenodo-Dep="4949971" httpUri="https://zenodo.org/record/4949971/files/figure.png" pageId="2" pageNumber="593" startId="2.[151,249,1876,1898]" targetBox="[157,1436,220,1788]" targetPageId="2">
<paragraph id="8B8236E4F3133E1AFF35F8E9A711DCB7" blockId="2.[151,1437,1876,1990]" pageId="2" pageNumber="593">
<emphasis id="B949EAF6F3133E1AFF35F8E9A26DDC18" bold="true" box="[151,276,1876,1898]" pageId="2" pageNumber="593">FIGURE 2.</emphasis>
Dactylus right third pereiopod, median aspect. A, 
<taxonomicName id="4C3D4D67F3133E1AFC84F8E9A7DDDC1B" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[806,1188,1876,1898]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3133E1AFC84F8E9A762DC18" box="[806,1051,1876,1897]" italics="true" pageId="2" pageNumber="593">Diapontonia maranulus</emphasis>
Bruce, 1986
</taxonomicName>
, ovigerous female pocl. 1.8, RMNH.CRUS.D. 566905; B, 
<taxonomicName id="4C3D4D67F3133E1AFE50F8CEA03ADCF9" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[498,835,1906,1928]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3133E1AFE50F8CEA1B9DCF9" box="[498,704,1907,1928]" italics="true" pageId="2" pageNumber="593">Periclimenes milleri</emphasis>
Bruce, 1986
</taxonomicName>
, female pocl. 2.8, RMNH.CRUS.D.36342; C, 
<taxonomicName id="4C3D4D67F3133E1AFABAF8CEA13BDCD6" authority="Berggren &amp; Svane, 1989" authorityName="Berggren &amp; Svane" authorityYear="1989" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="ingressicolumbi">
<emphasis id="B949EAF6F3133E1AFABAF8CEA241DCD7" italics="true" pageId="2" pageNumber="593">Periclimenes ingressicolumbi</emphasis>
<bibRefCitation id="EFAC4B15F3133E1AFE9DF82FA13BDCD6" author="Berggren, M. &amp; Svane, I." box="[319,578,1937,1959]" pageId="2" pageNumber="593" pagination="432 - 444" refId="ref2814" refString="Berggren, M. &amp; Svane, I. (1989) Periclimenes ingressicolumbi, new species, a pontoniine shrimp associated with deep-water echinoids off San Salvador Island in the Bahamas, and a comparison with Periclimenes milleri. Journal of Crustacean Biology, 9 (3), 432 - 444. http: // dx. doi. org / 10.1163 / 193724089 X 00412" type="journal article" year="1989">Berggren &amp; Svane, 1989</bibRefCitation>
</taxonomicName>
, ovigerous female, pocl. 2.3, RMNH.CRUS.D.37738; D, 
<taxonomicName id="4C3D4D67F3133E1AFB3AF82CA26CDCB7" authority="Bruce, 1990" authorityName="Bruce" authorityYear="1990" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="disparostris">
<emphasis id="B949EAF6F3133E1AFB3AF82CA6E5DCD7" box="[1176,1436,1937,1958]" italics="true" pageId="2" pageNumber="593">Altopontonia disparostris</emphasis>
<bibRefCitation id="EFAC4B15F3133E1AFF35F80DA26CDCB7" author="Bruce, A. J." box="[151,277,1968,1990]" pageId="2" pageNumber="593" pagination="149 - 215" refId="ref2974" refString="Bruce, A. J. (1990) Crustacea Decapoda: 5. Deep-sea palaemonoid shrimps from New Caledonian waters. In: A. Crosnier (Ed.), Resultats des Campagnes MUSORSTOM, 6. Memoires du Museum National d'Histoire Naturelle, 145, 149 - 215. [figs. 1 - 39]" type="journal article" year="1990">Bruce, 1990</bibRefCitation>
</taxonomicName>
, ovigerous female paratype, pocl. 3.1, RMNH.CRUS.D.37690. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF42666CF3123E1BFF35FB35A39FDFAA" ID-DOI="http://doi.org/10.5281/zenodo.4949973" ID-Zenodo-Dep="4949973" httpUri="https://zenodo.org/record/4949973/files/figure.png" pageId="3" pageNumber="594" startId="3.[151,249,1160,1182]" targetBox="[318,1268,185,1139]" targetPageId="3">
<paragraph id="8B8236E4F3123E1BFF35FB35A39FDFAA" blockId="3.[151,1436,1160,1243]" pageId="3" pageNumber="594">
<emphasis id="B949EAF6F3123E1BFF35FB35A26EDFEC" bold="true" box="[151,279,1160,1182]" pageId="3" pageNumber="594">FIGURE 3.</emphasis>
Host echinoid 
<taxonomicName id="4C3D4D67F3123E1BFE62FB35A00FDFEF" authority="Chesher, 1968" authorityName="Chesher" authorityYear="1968" box="[448,886,1160,1182]" class="Echinoidea" family="Paleopneustidae" genus="Paleopneustes" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="594" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3123E1BFE62FB35A1ABDFEC" box="[448,722,1160,1181]" italics="true" pageId="3" pageNumber="594">Paleopneustes tholoformis</emphasis>
Chesher, 1968
</taxonomicName>
, with 
<taxonomicName id="4C3D4D67F3123E1BFC1FFB34A638DFEF" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[957,1345,1160,1182]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3123E1BFC1FFB34A7CCDFEC" box="[957,1205,1160,1182]" italics="true" pageId="3" pageNumber="594">Diapontonia maranulus</emphasis>
Bruce, 1986
</taxonomicName>
shrimps (indicated by circles) photographed with Curasub at 220 m depth off the SE coast of Curaçao. Photo by Adriaan (Dutch) Schrier.
</paragraph>
</caption>
<paragraph id="8B8236E4F3123E1BFF65FAB5A016DEA4" blockId="3.[151,1437,1288,2033]" pageId="3" pageNumber="594">
<materialsCitation id="3B553CB9F3123E1BFF65FAB5A012DEA4" ID-GBIF-Occurrence="3391231301" collectionCode="RMNH, CRUS" country="Curaçao" location="The" municipality="The" pageId="3" pageNumber="594" specimenCount="1" typeStatus="paratype">
<location id="8EE2603FF3123E1BFF65FAB5A38ADE50" LSID="urn:lsid:plazi:treatment:039487F2F3113E1CFF35F91FA01CDD79:8EE2603FF3123E1BFF65FAB5A38ADE50" box="[199,243,1288,1313]" country="Curaçao" municipality="The" name="The" pageId="3" pageNumber="594">The</location>
specimens from 
<collectingCountry id="F32A7674F3123E1BFE17FAB5A16BDE50" box="[437,530,1288,1313]" name="Curaçao" pageId="3" pageNumber="594">Curaçao</collectingCountry>
have 7–10 (usually 8 or 9) small teeth on the dorsal margin of the rostrum (
<figureCitation id="13062A61F3123E1BFACDFAB5A3A2DE34" captionStart="FIGURE 1" captionStartId="1.[151,249,1873,1895]" captionTargetBox="[156,1389,974,1737]" captionTargetId="figure-349@1.[151,1436,971,1852]" captionTargetPageId="1" captionText="FIGURE 1. Diapontonia maranulus Bruce, 1986, RMNH.CRUS.D.566905. A, carapace and anterior appendages, lateral view; B, anterior appendages, dorsal view; C, carapace, lateral view; D, tailfan, dorsal view, setae omitted. A,B, D, ovigerous female, pocl. 1.8; C, ovigerous female, pocl. 2.1. Scale bar A–C = 2; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4949967" httpUri="https://zenodo.org/record/4949967/files/figure.png" pageId="3" pageNumber="594">Fig. 1A, B</figureCitation>
) of which 2 or 3 are situated postorbitally; the ventral margin of the rostrum has 1 or 2 subdistal teeth. 
<collectingMunicipality id="6BE6AC9EF3123E1BFAD3FA91A6E4DE34" box="[1393,1437,1324,1349]" pageId="3" pageNumber="594">The</collectingMunicipality>
smaller female (pocl 1.8) of 
<collectionCode id="ED2CAE21F3123E1BFE55FAEDA129DE18" box="[503,592,1360,1385]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="594" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED2CAE21F3123E1BFDF1FAEDA1E7DE18" box="[595,670,1360,1385]" pageId="3" pageNumber="594">CRUS</collectionCode>
.D.56690 has the ventral margin devoid of teeth (
<figureCitation id="13062A61F3123E1BFB55FAEDA62DDE18" box="[1271,1364,1360,1385]" captionStart="FIGURE 1" captionStartId="1.[151,249,1873,1895]" captionTargetBox="[156,1389,974,1737]" captionTargetId="figure-349@1.[151,1436,971,1852]" captionTargetPageId="1" captionText="FIGURE 1. Diapontonia maranulus Bruce, 1986, RMNH.CRUS.D.566905. A, carapace and anterior appendages, lateral view; B, anterior appendages, dorsal view; C, carapace, lateral view; D, tailfan, dorsal view, setae omitted. A,B, D, ovigerous female, pocl. 1.8; C, ovigerous female, pocl. 2.1. Scale bar A–C = 2; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4949967" httpUri="https://zenodo.org/record/4949967/files/figure.png" pageId="3" pageNumber="594">Fig. 1B</figureCitation>
). The 
<typeStatus id="54868846F3123E1BFF35FAC8A27BDEFC" box="[151,258,1397,1421]" pageId="3" pageNumber="594" type="paratype">paratypes</typeStatus>
have 7–9 teeth on the dorsal margin of the rostrum of which 0–3 are situated postorbitally; the ventral margin of these specimens always has a subdistal tooth although minute in the small males. Carapace and anterior appendages (
<figureCitation id="13062A61F3123E1BFE8BFA01A2D9DEA4" box="[297,416,1468,1493]" captionStart="FIGURE 1" captionStartId="1.[151,249,1873,1895]" captionTargetBox="[156,1389,974,1737]" captionTargetId="figure-349@1.[151,1436,971,1852]" captionTargetPageId="1" captionText="FIGURE 1. Diapontonia maranulus Bruce, 1986, RMNH.CRUS.D.566905. A, carapace and anterior appendages, lateral view; B, anterior appendages, dorsal view; C, carapace, lateral view; D, tailfan, dorsal view, setae omitted. A,B, D, ovigerous female, pocl. 1.8; C, ovigerous female, pocl. 2.1. Scale bar A–C = 2; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4949967" httpUri="https://zenodo.org/record/4949967/files/figure.png" pageId="3" pageNumber="594">Fig. 1B, C</figureCitation>
) are as described for the type-specimens
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236E4F3123E1BFF65FA5DA16BDDF8" blockId="3.[151,1437,1288,2033]" pageId="3" pageNumber="594">
Abdomen, caudal fan (
<figureCitation id="13062A61F3123E1BFE65FA5DA164DE88" box="[455,541,1504,1529]" captionStart="FIGURE 1" captionStartId="1.[151,249,1873,1895]" captionTargetBox="[156,1389,974,1737]" captionTargetId="figure-349@1.[151,1436,971,1852]" captionTargetPageId="1" captionText="FIGURE 1. Diapontonia maranulus Bruce, 1986, RMNH.CRUS.D.566905. A, carapace and anterior appendages, lateral view; B, anterior appendages, dorsal view; C, carapace, lateral view; D, tailfan, dorsal view, setae omitted. A,B, D, ovigerous female, pocl. 1.8; C, ovigerous female, pocl. 2.1. Scale bar A–C = 2; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4949967" httpUri="https://zenodo.org/record/4949967/files/figure.png" pageId="3" pageNumber="594">Fig. 1D</figureCitation>
) and pereiopods are as described for the type specimens. The configuration and ornamentation of the teeth on the flexor margin of the dactylus as well as the unguis and ventral and distoventral spines of the propodus of the third pereiopod (
<figureCitation id="13062A61F3123E1BFD0BF995A07ADD30" box="[681,771,1576,1601]" captionStart="FIGURE 2" captionStartId="2.[151,249,1876,1898]" captionTargetBox="[157,1436,220,1788]" captionTargetId="figure-80@2.[151,1436,215,2032]" captionTargetPageId="2" captionText="FIGURE 2. Dactylus right third pereiopod, median aspect. A, Diapontonia maranulus Bruce, 1986, ovigerous female pocl. 1.8, RMNH.CRUS.D. 566905; B, Periclimenes milleri Bruce, 1986, female pocl. 2.8, RMNH.CRUS.D.36342; C, Periclimenes ingressicolumbi Berggren &amp; Svane, 1989, ovigerous female, pocl. 2.3, RMNH.CRUS.D.37738; D, Altopontonia disparostris Bruce, 1990, ovigerous female paratype, pocl. 3.1, RMNH.CRUS.D.37690. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4949971" httpUri="https://zenodo.org/record/4949971/files/figure.png" pageId="3" pageNumber="594">Fig. 2A</figureCitation>
) is as in the female 
<typeStatus id="54868846F3123E1BFC4EF994A735DD30" box="[1004,1100,1577,1601]" pageId="3" pageNumber="594" type="paratype">paratype</typeStatus>
figured by Bruce (1986: Fig. 5A). Both third and fourth pereiopods with one distoventral spine on lateral surface and two on medial surface as figured by Bruce (1986: Fig. 5D).
</paragraph>
<paragraph id="8B8236E4F3123E1BFF65F929A11FDDDC" blockId="3.[151,1437,1288,2033]" box="[199,614,1684,1709]" pageId="3" pageNumber="594">
<emphasis id="B949EAF6F3123E1BFF65F929A236DDDD" box="[199,335,1684,1709]" inLineHeading="true" pageId="3" pageNumber="594" reason="1">
<emphasis id="B949EAF6F3123E1BFF65F929A231DDDC" bold="true" box="[199,328,1684,1709]" pageId="3" pageNumber="594">Coloration</emphasis>
.
</emphasis>
Uniform brown (
<figureCitation id="13062A61F3123E1BFDB6F929A12EDDDC" box="[532,599,1684,1709]" captionStart="FIGURE 3" captionStartId="3.[151,249,1160,1182]" captionTargetBox="[318,1268,185,1139]" captionTargetId="figure-385@3.[318,1268,185,1139]" captionTargetPageId="3" captionText="FIGURE 3. Host echinoid Paleopneustes tholoformis Chesher, 1968, with Diapontonia maranulus Bruce, 1986 shrimps (indicated by circles) photographed with Curasub at 220 m depth off the SE coast of Curaçao. Photo by Adriaan (Dutch) Schrier." figureDoi="http://doi.org/10.5281/zenodo.4949973" httpUri="https://zenodo.org/record/4949973/files/figure.png" pageId="3" pageNumber="594">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327656FF3123E1CFF65F905A01CDD79" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594" type="biology_ecology">
<paragraph id="8B8236E4F3123E1BFF65F905A732DD84" blockId="3.[151,1437,1288,2033]" pageId="3" pageNumber="594">
<emphasis id="B949EAF6F3123E1BFF65F905A386DDA0" bold="true" box="[199,255,1720,1745]" pageId="3" pageNumber="594">Host</emphasis>
. 
<taxonomicName id="4C3D4D67F3123E1BFEB2F905A195DDA0" authority="Chesher, 1968" authorityName="Chesher" authorityYear="1968" box="[272,748,1720,1745]" class="Echinoidea" family="Paleopneustidae" genus="Paleopneustes" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="594" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3123E1BFEB2F905A144DDA1" box="[272,573,1720,1744]" italics="true" pageId="3" pageNumber="594">Paleopneustes tholoformis</emphasis>
Chesher, 1968
</taxonomicName>
(Echinodermata: 
<taxonomicName id="4C3D4D67F3123E1BFC19F905A747DDA0" box="[955,1086,1720,1745]" class="Echinoidea" kingdom="Animalia" pageId="3" pageNumber="594" phylum="Echinodermata" rank="class">Echinoidea</taxonomicName>
: 
<taxonomicName id="4C3D4D67F3123E1BFBE9F905A668DDA0" authorityName="Pictet" authorityYear="1857" box="[1099,1297,1720,1745]" class="Echinoidea" family="Asterostomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="3" pageNumber="594" phylum="Echinodermata" rank="family">Asterostomatidae</taxonomicName>
) (
<figureCitation id="13062A61F3123E1BFA89F905A608DDA0" box="[1323,1393,1720,1745]" captionStart="FIGURE 3" captionStartId="3.[151,249,1160,1182]" captionTargetBox="[318,1268,185,1139]" captionTargetId="figure-385@3.[318,1268,185,1139]" captionTargetPageId="3" captionText="FIGURE 3. Host echinoid Paleopneustes tholoformis Chesher, 1968, with Diapontonia maranulus Bruce, 1986 shrimps (indicated by circles) photographed with Curasub at 220 m depth off the SE coast of Curaçao. Photo by Adriaan (Dutch) Schrier." figureDoi="http://doi.org/10.5281/zenodo.4949973" httpUri="https://zenodo.org/record/4949973/files/figure.png" pageId="3" pageNumber="594">Fig. 3</figureCitation>
). It was observed that one echinoid could host up to 
<specimenCount id="9D3BFD6DF3123E1BFD1AF961A029DD84" box="[696,848,1756,1781]" pageId="3" pageNumber="594" type="generic">10 specimens</specimenCount>
of the shrimp species.
</paragraph>
<paragraph id="8B8236E4F3123E1BFF65F8BDA1C7DC80" blockId="3.[151,1437,1288,2033]" pageId="3" pageNumber="594">
<emphasis id="B949EAF6F3123E1BFF65F8BDA2CCDC68" bold="true" box="[199,437,1792,1817]" inLineHeading="true" pageId="3" pageNumber="594" reason="1">Systematic position.</emphasis>
At present three species of deep water echinoid associated pontoniine shrimps have been described from the tropical West Atlantic: 
<taxonomicName id="4C3D4D67F3123E1BFD07F898A0C3DC4D" authorityName="Bruce" authorityYear="1986" box="[677,954,1828,1852]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3123E1BFD07F898A0C3DC4D" box="[677,954,1828,1852]" italics="true" pageId="3" pageNumber="594">Diapontonia maranulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3D4D67F3123E1BFC72F899A619DC4C" authority="Bruce, 1986" authorityName="Bruce" authorityYear="1986" box="[976,1376,1828,1853]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3123E1BFC72F899A7C4DC4D" box="[976,1213,1828,1852]" italics="true" pageId="3" pageNumber="594">Periclimenes milleri</emphasis>
Bruce, 1986
</taxonomicName>
, and 
<taxonomicName id="4C3D4D67F3123E1BFF35F8F5A079DC10" authority="Berggren &amp; Svane, 1989" authorityName="Berggren &amp; Svane" authorityYear="1989" box="[151,768,1864,1889]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="ingressicolumbi">
<emphasis id="B949EAF6F3123E1BFF35F8F5A298DC11" box="[151,481,1864,1888]" italics="true" pageId="3" pageNumber="594">Periclimenes ingressicolumbi</emphasis>
<bibRefCitation id="EFAC4B15F3123E1BFE4AF8F4A079DC10" author="Berggren, M. &amp; Svane, I." box="[488,768,1864,1889]" pageId="3" pageNumber="594" pagination="432 - 444" refId="ref2814" refString="Berggren, M. &amp; Svane, I. (1989) Periclimenes ingressicolumbi, new species, a pontoniine shrimp associated with deep-water echinoids off San Salvador Island in the Bahamas, and a comparison with Periclimenes milleri. Journal of Crustacean Biology, 9 (3), 432 - 444. http: // dx. doi. org / 10.1163 / 193724089 X 00412" type="journal article" year="1989">Berggren &amp; Svane, 1989</bibRefCitation>
</taxonomicName>
. These three species were until now only known from a few records in the vicinity of the Bahama Islands (
<bibRefCitation id="EFAC4B15F3123E1BFD0AF8D1A03CDCF4" author="Bruce, A. J." box="[680,837,1900,1925]" pageId="3" pageNumber="594" pagination="125 - 133" refId="ref2882" refString="Bruce, A. J. (1986 a) Diapontonia maranulus, new genus, new species, a pontoniine shrimp associate of a deep-water echinoid. Journal of the Crustacean Biology, 6 (1), 125 - 133. http: // dx. doi. org / 10.1163 / 193724086 X 00785" type="journal article" year="1986">Bruce, 1986a</bibRefCitation>
,b; 
<bibRefCitation id="EFAC4B15F3123E1BFCC4F8D0A7FCDCF4" author="Berggren, M. &amp; Svane, I." box="[870,1157,1900,1925]" pageId="3" pageNumber="594" pagination="432 - 444" refId="ref2814" refString="Berggren, M. &amp; Svane, I. (1989) Periclimenes ingressicolumbi, new species, a pontoniine shrimp associated with deep-water echinoids off San Salvador Island in the Bahamas, and a comparison with Periclimenes milleri. Journal of Crustacean Biology, 9 (3), 432 - 444. http: // dx. doi. org / 10.1163 / 193724089 X 00412" type="journal article" year="1989">Berggren &amp; Svane, 1989</bibRefCitation>
). The recently described deep water species 
<taxonomicName id="4C3D4D67F3123E1BFECDF82DA093DCD8" authority="Anker, Pachelle &amp; Tavares, 2014" authorityName="Anker, Pachelle &amp; Tavares" authorityYear="2014" box="[367,1002,1936,1961]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="bathyalis">
<emphasis id="B949EAF6F3123E1BFECDF82DA116DCD9" box="[367,623,1936,1960]" italics="true" pageId="3" pageNumber="594">Periclimenes bathyalis</emphasis>
<bibRefCitation id="EFAC4B15F3123E1BFDD4F82DA093DCD8" author="Anker, A. &amp; Pachelle, P. G. &amp; Tavares, M." box="[630,1002,1936,1961]" pageId="3" pageNumber="594" pagination="263 - 278" refId="ref2747" refString="Anker, A., Pachelle, P. G. &amp; Tavares, M. (2014) Two new species and two new records of deep-water caridean shrimps from Brazil (Decapoda: Pandalidae, Palaemonidae, Crangonidae). Zootaxa, 3815 (2), 263 - 278. http: // dx. doi. org / 10.11646 / zootaxa. 3815.2.6" type="journal article" year="2014">Anker, Pachelle &amp; Tavares, 2014</bibRefCitation>
</taxonomicName>
, collected off 
<collectingRegion id="49F9F806F3123E1BFB2AF82DA653DCD8" box="[1160,1322,1936,1961]" country="Brazil" name="Espirito Santo" pageId="3" pageNumber="594">Espírito Santo</collectingRegion>
, 
<collectingCountry id="F32A7674F3123E1BFA94F82DA604DCD8" box="[1334,1405,1936,1961]" name="Brazil" pageId="3" pageNumber="594">Brazil</collectingCountry>
, at a depth of 
<quantity id="4CC59B01F3123E1BFEBEF809A210DCBD" box="[284,361,1972,1997]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="594" unit="m" value="360.0">360 m</quantity>
, is morphologically similar to 
<taxonomicName id="4C3D4D67F3123E1BFD7AF808A03ADCBD" authorityName="Bruce" authorityYear="1986" box="[728,835,1972,1996]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3123E1BFD7AF808A03ADCBD" box="[728,835,1972,1996]" italics="true" pageId="3" pageNumber="594">P. milleri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D4D67F3123E1BFC21F808A72CDCBD" authorityName="Berggren &amp; Svane" authorityYear="1989" box="[899,1109,1972,1996]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="ingressicolumbi">
<emphasis id="B949EAF6F3123E1BFC21F808A72CDCBD" box="[899,1109,1972,1996]" italics="true" pageId="3" pageNumber="594">P. ingressicolumbi</emphasis>
</taxonomicName>
. It is only known from the 
<typeStatus id="54868846F3123E1BFF35F865A380DC80" box="[151,249,2008,2033]" pageId="3" pageNumber="594" type="holotype">holotype</typeStatus>
male specimen and its host is unknown.
</paragraph>
<paragraph id="8B8236E4F3153E1CFF65FF25A007DA14" blockId="4.[151,1440,151,1544]" pageId="4" pageNumber="595">
<taxonomicName id="4C3D4D67F3153E1CFF65FF25A2A1DBC1" authorityName="Bruce" authorityYear="1986" box="[199,472,152,176]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3153E1CFF65FF25A2A1DBC1" box="[199,472,152,176]" italics="true" pageId="4" pageNumber="595">Diapontonia maranulus</emphasis>
</taxonomicName>
differs from the three 
<taxonomicName id="4C3D4D67F3153E1CFD4FFF25A0A5DBC1" box="[749,988,151,176]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B949EAF6F3153E1CFD4FFF25A006DBC1" box="[749,895,152,176]" italics="true" pageId="4" pageNumber="595">Periclimenes</emphasis>
species
</taxonomicName>
in the absence of a hepatic tooth, an important character often used in the delimitation of genera in 
<taxonomicName id="4C3D4D67F3153E1CFCF0FF01A0A5DBA4" authorityName="Kingsley" authorityYear="1879" box="[850,988,188,213]" class="Malacostraca" family="Pontoniinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="family">Pontoniinae</taxonomicName>
. Morphological differences between the three species of 
<taxonomicName id="4C3D4D67F3153E1CFEF2FF5DA29BDB89" authorityName="O.G.Costa" authorityYear="1844" box="[336,482,224,248]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFEF2FF5DA29BDB89" box="[336,482,224,248]" italics="true" pageId="4" pageNumber="595">Periclimenes</emphasis>
</taxonomicName>
are mainly found in the ventral rostral dentition, antennular peduncle, scaphocerite, major second pereiopod, size and position of the hepatic tooth, propodi and dactyli of the third to fifth pereiopods, telson, number of setae and spines on the endopod of the first pleopod, and the size and number of spines on the appendix masculina (
<bibRefCitation id="EFAC4B15F3153E1CFE26FEF1A1E6DA14" author="Berggren, M. &amp; Svane, I." box="[388,671,332,357]" pageId="4" pageNumber="595" pagination="432 - 444" refId="ref2814" refString="Berggren, M. &amp; Svane, I. (1989) Periclimenes ingressicolumbi, new species, a pontoniine shrimp associated with deep-water echinoids off San Salvador Island in the Bahamas, and a comparison with Periclimenes milleri. Journal of Crustacean Biology, 9 (3), 432 - 444. http: // dx. doi. org / 10.1163 / 193724089 X 00412" type="journal article" year="1989">Berggren &amp; Svane, 1989</bibRefCitation>
; 
<bibRefCitation id="EFAC4B15F3153E1CFD09FEF1A00FDA14" author="Anker, A. &amp; Pachelle, P. G. &amp; Tavares, M." box="[683,886,332,357]" pageId="4" pageNumber="595" pagination="263 - 278" refId="ref2747" refString="Anker, A., Pachelle, P. G. &amp; Tavares, M. (2014) Two new species and two new records of deep-water caridean shrimps from Brazil (Decapoda: Pandalidae, Palaemonidae, Crangonidae). Zootaxa, 3815 (2), 263 - 278. http: // dx. doi. org / 10.11646 / zootaxa. 3815.2.6" type="journal article" year="2014">Anker et al., 2014</bibRefCitation>
)
</paragraph>
<paragraph id="8B8236E4F3153E1CFF77FECDA00EDABE" blockId="4.[151,1440,151,1544]" pageId="4" pageNumber="595">
It is interesting to note that 
<bibRefCitation id="EFAC4B15F3153E1CFD9AFECDA008DAF9" author="Berggren, M. &amp; Svane, I." box="[568,881,367,392]" pageId="4" pageNumber="595" pagination="432 - 444" refId="ref2814" refString="Berggren, M. &amp; Svane, I. (1989) Periclimenes ingressicolumbi, new species, a pontoniine shrimp associated with deep-water echinoids off San Salvador Island in the Bahamas, and a comparison with Periclimenes milleri. Journal of Crustacean Biology, 9 (3), 432 - 444. http: // dx. doi. org / 10.1163 / 193724089 X 00412" type="journal article" year="1989">Berggren &amp; Svane (1989)</bibRefCitation>
collected 
<specimenCount id="9D3BFD6DF3153E1CFC56FECDA7CEDAF9" box="[1012,1207,367,392]" pageId="4" pageNumber="595" type="generic">seven specimens</specimenCount>
from the echinoid 
<taxonomicName id="4C3D4D67F3153E1CFF35FE29A2BADADD" authorityName="Chesher" authorityYear="1968" box="[151,451,404,428]" class="Echinoidea" family="Paleopneustidae" genus="Paleopneustes" kingdom="Animalia" order="Spatangoida" pageId="4" pageNumber="595" phylum="Echinodermata" rank="species" species="tholoformis">
<emphasis id="B949EAF6F3153E1CFF35FE29A2BADADD" box="[151,451,404,428]" italics="true" pageId="4" pageNumber="595">Paleopneustes tholoformis</emphasis>
</taxonomicName>
, identified by them as belonging to 
<taxonomicName id="4C3D4D67F3153E1CFCCBFE28A0A8DADD" authorityName="Bruce" authorityYear="1986" box="[873,977,404,428]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3153E1CFCCBFE28A0A8DADD" box="[873,977,404,428]" italics="true" pageId="4" pageNumber="595">P. milleri</emphasis>
</taxonomicName>
, of which two were lacking the hepatic tooth and in 
<specimenCount id="9D3BFD6DF3153E1CFE86FE05A2C4DAA1" box="[292,445,439,464]" pageId="4" pageNumber="595" type="generic">one specimen</specimenCount>
a very small hepatic tooth was present.
</paragraph>
<paragraph id="8B8236E4F3153E1CFF72FE61A61BD99E" blockId="4.[151,1440,151,1544]" pageId="4" pageNumber="595">
The three Caribbean species are very similar and share several apomorphic characters, most conspicuous in the armature of the ambulatory propodi and dactyli (
<figureCitation id="13062A61F3153E1CFD4BFE42A01BD969" box="[745,866,511,536]" captionStart="FIGURE 2" captionStartId="2.[151,249,1876,1898]" captionTargetBox="[157,1436,220,1788]" captionTargetId="figure-80@2.[151,1436,215,2032]" captionTargetPageId="2" captionText="FIGURE 2. Dactylus right third pereiopod, median aspect. A, Diapontonia maranulus Bruce, 1986, ovigerous female pocl. 1.8, RMNH.CRUS.D. 566905; B, Periclimenes milleri Bruce, 1986, female pocl. 2.8, RMNH.CRUS.D.36342; C, Periclimenes ingressicolumbi Berggren &amp; Svane, 1989, ovigerous female, pocl. 2.3, RMNH.CRUS.D.37738; D, Altopontonia disparostris Bruce, 1990, ovigerous female paratype, pocl. 3.1, RMNH.CRUS.D.37690. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4949971" httpUri="https://zenodo.org/record/4949971/files/figure.png" pageId="4" pageNumber="595">Fig. 2A–C</figureCitation>
) like (1) the distally strongly serrulate ventral and distoventral spines of the propodi, (2) the presence of three distoventral spines on the propodi of the third and fourth pereiopods of which one situated laterally and two medially, (3) the presence of a series of 8–10 acute ventral teeth on the flexor margin of the dactylus, and (4) with the ventral border of the unguis minutely crenulate or denticulate. The presence of minute crenulation on the ventral margin of the unguis in 
<taxonomicName id="4C3D4D67F3153E1CFB6BFD2CA64CD9D9" authorityName="Bruce" authorityYear="1986" box="[1225,1333,656,680]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3153E1CFB6BFD2CA64CD9D9" box="[1225,1333,656,680]" italics="true" pageId="4" pageNumber="595">P. milleri</emphasis>
</taxonomicName>
was not described by 
<bibRefCitation id="EFAC4B15F3153E1CFEE7FD09A28DD9BC" author="Bruce, A. J." box="[325,500,692,717]" pageId="4" pageNumber="595" pagination="637 - 645" refId="ref2939" refString="Bruce, A. J. (1986 b) Periclimenes milleri new species, a bathyal echinoid-associated pontoniine shrimp from the Bahamas. Bulletin of Marine Science, 39, 637 - 645." type="journal article" year="1986">Bruce (1986b)</bibRefCitation>
nor by 
<bibRefCitation id="EFAC4B15F3153E1CFDD3FD09A0C1D9BC" author="Berggren, M. &amp; Svane, I." box="[625,952,692,717]" pageId="4" pageNumber="595" pagination="432 - 444" refId="ref2814" refString="Berggren, M. &amp; Svane, I. (1989) Periclimenes ingressicolumbi, new species, a pontoniine shrimp associated with deep-water echinoids off San Salvador Island in the Bahamas, and a comparison with Periclimenes milleri. Journal of Crustacean Biology, 9 (3), 432 - 444. http: // dx. doi. org / 10.1163 / 193724089 X 00412" type="journal article" year="1989">Berggren &amp; Svane (1989)</bibRefCitation>
but observed in the 
<typeStatus id="54868846F3153E1CFB7EFD09A644D9BD" box="[1244,1341,692,716]" pageId="4" pageNumber="595" type="paratype">paratype</typeStatus>
female (RMNH.CRUS.D.36342) as is shown in figure 2C. These characters have not been described for 
<taxonomicName id="4C3D4D67F3153E1CFB7BFD64A623D981" authorityName="Anker, Pachelle &amp; Tavares" authorityYear="2014" box="[1241,1370,728,752]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="bathyalis">
<emphasis id="B949EAF6F3153E1CFB7BFD64A623D981" box="[1241,1370,728,752]" italics="true" pageId="4" pageNumber="595">P. bathyalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8236E4F3153E1CFF65FD41A6E4DF61" blockId="4.[151,1440,151,1544]" pageId="4" pageNumber="595">
The most closely related species from the Indo-West Pacific seems 
<taxonomicName id="4C3D4D67F3153E1CFC7EFD41A6E0D864" authority="Bruce, 1990" authorityName="Bruce" authorityYear="1990" box="[988,1433,764,789]" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="disparostris">
<emphasis id="B949EAF6F3153E1CFC7EFD41A787D865" box="[988,1278,764,788]" italics="true" pageId="4" pageNumber="595">Altopontonia disparostris</emphasis>
<bibRefCitation id="EFAC4B15F3153E1CFAABFD41A6E0D864" author="Bruce, A. J." box="[1289,1433,764,789]" pageId="4" pageNumber="595" pagination="149 - 215" refId="ref2974" refString="Bruce, A. J. (1990) Crustacea Decapoda: 5. Deep-sea palaemonoid shrimps from New Caledonian waters. In: A. Crosnier (Ed.), Resultats des Campagnes MUSORSTOM, 6. Memoires du Museum National d'Histoire Naturelle, 145, 149 - 215. [figs. 1 - 39]" type="journal article" year="1990">Bruce, 1990</bibRefCitation>
</taxonomicName>
. Specimens of this species have been collected from depths of 
<quantity id="4CC59B01F3153E1CFCF4FCA2A0AFD846" box="[854,982,799,824]" metricMagnitude="2" metricUnit="m" metricValue="4.125" metricValueMax="5.029999999999999" metricValueMin="3.22" pageId="4" pageNumber="595" unit="m" value="412.5" valueMax="503.0" valueMin="322.0">322–503 m</quantity>
off 
<collectingCountry id="F32A7674F3153E1CFBA4FC9DA7CED849" box="[1030,1207,799,824]" name="New Caledonia" pageId="4" pageNumber="595">New Caledonia</collectingCountry>
, the 
<collectingRegion id="49F9F806F3153E1CFB4CFCA2A6E5D849" box="[1262,1436,799,824]" country="New Caledonia" name="Province des Iles" pageId="4" pageNumber="595">Loyalty Islands</collectingRegion>
and the North Norfolk Ridge (
<bibRefCitation id="EFAC4B15F3153E1CFE56FCF9A1FAD82C" author="Bruce, A. J." box="[500,643,836,861]" pageId="4" pageNumber="595" pagination="149 - 215" refId="ref2974" refString="Bruce, A. J. (1990) Crustacea Decapoda: 5. Deep-sea palaemonoid shrimps from New Caledonian waters. In: A. Crosnier (Ed.), Resultats des Campagnes MUSORSTOM, 6. Memoires du Museum National d'Histoire Naturelle, 145, 149 - 215. [figs. 1 - 39]" type="journal article" year="1990">Bruce, 1990</bibRefCitation>
, 
<bibRefCitation id="EFAC4B15F3153E1CFD32FCF9A1B3D82C" author="Bruce, A. J." box="[656,714,836,861]" pageId="4" pageNumber="595" pagination="299 - 411" refId="ref3033" refString="Bruce, A. J. (1991) Crustacea Decapoda: Further deep-sea palaemonoid shrimps from New Caledonian waters. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM, 9. Memoires du Museum National d'Histoire Naturelle, 152 (A), 299 - 411. [figs. 1 - 75]" type="journal article" year="1991">1991</bibRefCitation>
, 
<bibRefCitation id="EFAC4B15F3153E1CFD7AFCF9A06AD82C" author="Bruce, A. J." box="[728,787,836,861]" pageId="4" pageNumber="595" pagination="197 - 267" refId="ref3095" refString="Bruce, A. J. (1996) Crustacea Decapoda: Palaemonid shrimps from the Indo-West Pacific region, mainly from New Caledonia. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM, 15. Memoires du Museum National d'Histoire Naturelle, 168, 197 - 267. [figs 1 - 31]" type="journal article" year="1996">1996</bibRefCitation>
, 
<bibRefCitation id="EFAC4B15F3153E1CFC82FCF9A024D82C" author="Bruce, A. J." box="[800,861,836,861]" pageId="4" pageNumber="595" pagination="1 - 20" refId="ref3157" refString="Bruce, A. J. (2005) Pontoniine shrimps from the 2003 NORFANZ Expedition, 10 May - 16 June (Crustacea: Decapoda: Palaemonidae). Zootaxa, 981, 1 - 20." type="journal article" year="2005">2005</bibRefCitation>
). Its host is unknown, although its resemblance to 
<taxonomicName id="4C3D4D67F3153E1CFF35FCD5A25CD80E" authorityName="Bruce" authorityYear="1986" box="[151,293,872,895]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFF35FCD5A25CD80E" box="[151,293,872,895]" italics="true" pageId="4" pageNumber="595">Diapontonia</emphasis>
</taxonomicName>
suggests it might be associated with echinoids (
<bibRefCitation id="EFAC4B15F3153E1CFCEBFCDAA0A1D8F1" author="Bruce, A. J." box="[841,984,871,896]" pageId="4" pageNumber="595" pagination="149 - 215" refId="ref2974" refString="Bruce, A. J. (1990) Crustacea Decapoda: 5. Deep-sea palaemonoid shrimps from New Caledonian waters. In: A. Crosnier (Ed.), Resultats des Campagnes MUSORSTOM, 6. Memoires du Museum National d'Histoire Naturelle, 145, 149 - 215. [figs. 1 - 39]" type="journal article" year="1990">Bruce, 1990</bibRefCitation>
). The species is generally similar to the three Atlantic species mentioned above. It shares the presence of the strongly serrulate ventral and distoventral spines on the propodi (
<figureCitation id="13062A61F3153E1CFE3AFC12A289D8B9" box="[408,496,943,968]" captionStart="FIGURE 2" captionStartId="2.[151,249,1876,1898]" captionTargetBox="[157,1436,220,1788]" captionTargetId="figure-80@2.[151,1436,215,2032]" captionTargetPageId="2" captionText="FIGURE 2. Dactylus right third pereiopod, median aspect. A, Diapontonia maranulus Bruce, 1986, ovigerous female pocl. 1.8, RMNH.CRUS.D. 566905; B, Periclimenes milleri Bruce, 1986, female pocl. 2.8, RMNH.CRUS.D.36342; C, Periclimenes ingressicolumbi Berggren &amp; Svane, 1989, ovigerous female, pocl. 2.3, RMNH.CRUS.D.37738; D, Altopontonia disparostris Bruce, 1990, ovigerous female paratype, pocl. 3.1, RMNH.CRUS.D.37690. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4949971" httpUri="https://zenodo.org/record/4949971/files/figure.png" pageId="4" pageNumber="595">Fig. 2D</figureCitation>
) of the ambulatory pereiopods with the three Atlantic species. It has the absence of a hepatic tooth in common with 
<taxonomicName id="4C3D4D67F3153E1CFE5DFC68A075D89D" authorityName="Bruce" authorityYear="1986" box="[511,780,980,1004]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="maranulus">
<emphasis id="B949EAF6F3153E1CFE5DFC68A075D89D" box="[511,780,980,1004]" italics="true" pageId="4" pageNumber="595">Diapontonia maranulus</emphasis>
</taxonomicName>
. It differs from the three Atlantic species principally in the form of the major chela and in having the flexor margin of the dactyli of the ambulatory pereiopods devoid of teeth.
</paragraph>
<paragraph id="8B8236E4F3153E1CFF65FBA1A74DDE41" blockId="4.[151,1440,151,1544]" pageId="4" pageNumber="595">
Species of the recently described Indo-West Pacific deep-water, possibly echinoid associated genus 
<taxonomicName id="4C3D4D67F3153E1CFF35FBFDA1F4DF29" authority="(Marin &amp; Chan, 2014)" baseAuthorityName="Marin &amp; Chan" baseAuthorityYear="2014" box="[151,653,1087,1112]" class="Malacostraca" family="Palaemonidae" genus="Echinopericlimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFF35FBFDA20CDF29" box="[151,373,1088,1112]" italics="true" pageId="4" pageNumber="595">Echinopericlimenes</emphasis>
(
<bibRefCitation id="EFAC4B15F3153E1CFE2BFB82A1FFDF29" author="Marin, I. &amp; Chan, T. - Y." box="[393,646,1087,1112]" pageId="4" pageNumber="595" pagination="301 - 324" refId="ref3284" refString="Marin, I. &amp; Chan, T. - Y. (2014) Deep water echinoid-associated pontoniine shrimp &quot; Periclimenes hertwigi Balss, 1913 &quot; species group (Crustacea: Decapoda: Caridea: Palaemonidae): species review, description of a new genus and species from Philippines. Zootaxa, 3835 (3), 301 - 324. http: // dx. doi. org / 10.11646 / zootaxa. 3835.3.1" type="journal article" year="2014">Marin &amp; Chan, 2014</bibRefCitation>
)
</taxonomicName>
share several characters in the rostrum, chelae and dactyli of the ambulatory pereiopods with 
<taxonomicName id="4C3D4D67F3153E1CFE42FBD8A117DF0D" authorityName="Bruce" authorityYear="1986" box="[480,622,1125,1148]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFE42FBD8A117DF0D" box="[480,622,1125,1148]" italics="true" pageId="4" pageNumber="595">Diapontonia</emphasis>
</taxonomicName>
and 
<emphasis id="B949EAF6F3153E1CFD08FBD9A03ADF0D" box="[682,835,1124,1148]" italics="true" pageId="4" pageNumber="595">
<taxonomicName id="4C3D4D67F3153E1CFD08FBD9A046DF0D" authorityName="Bruce" authorityYear="1990" box="[682,831,1124,1148]" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">Altopontonia</taxonomicName>
,
</emphasis>
though differ in having a distinct hepatic tooth like 
<taxonomicName id="4C3D4D67F3153E1CFF35FB35A206DFD1" authorityName="Bruce" authorityYear="1986" box="[151,383,1160,1184]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3153E1CFF35FB35A206DFD1" box="[151,383,1160,1184]" italics="true" pageId="4" pageNumber="595">Periclimenes milleri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D4D67F3153E1CFE1CFB34A1F6DFD1" authorityName="Berggren &amp; Svane" authorityYear="1989" box="[446,655,1160,1184]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="ingressicolumbi">
<emphasis id="B949EAF6F3153E1CFE1CFB34A1F6DFD1" box="[446,655,1160,1184]" italics="true" pageId="4" pageNumber="595">P. ingressicolumbi</emphasis>
</taxonomicName>
have. With 
<taxonomicName id="4C3D4D67F3153E1CFC86FB35A0CBDFEE" authorityName="Bruce" authorityYear="1986" box="[804,946,1160,1183]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFC86FB35A0CBDFEE" box="[804,946,1160,1183]" italics="true" pageId="4" pageNumber="595">Diapontonia</emphasis>
</taxonomicName>
they share the denticulation of the flexor margin of the unguis of the ambulatory dactyli and the irregular dentition of the dactylar accessory tooth. They differ in having the flexor margin of the dactyli entire while it is denticulate in 
<taxonomicName id="4C3D4D67F3153E1CFB8EFB6DA7C0DF96" authorityName="Bruce" authorityYear="1986" box="[1068,1209,1232,1255]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFB8EFB6DA7C0DF96" box="[1068,1209,1232,1255]" italics="true" pageId="4" pageNumber="595">Diapontonia</emphasis>
</taxonomicName>
. With 
<taxonomicName id="4C3D4D67F3153E1CFAA8FB6DA6E5DF99" authorityName="Bruce" authorityYear="1990" box="[1290,1436,1232,1256]" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFAA8FB6DA6E5DF99" box="[1290,1436,1232,1256]" italics="true" pageId="4" pageNumber="595">Altopontonia</emphasis>
</taxonomicName>
they share the entire flexor margin of the dactyli of the ambulatory pereiopods. However. 
<taxonomicName id="4C3D4D67F3153E1CFB03FB49A64ADE7D" authorityName="Bruce" authorityYear="1990" box="[1185,1331,1268,1292]" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFB03FB49A64ADE7D" box="[1185,1331,1268,1292]" italics="true" pageId="4" pageNumber="595">Altopontonia</emphasis>
</taxonomicName>
does not show the denticulate accessory tooth nor a denticulate flexor margin of the unguis.
</paragraph>
<paragraph id="8B8236E4F3153E1CFF65FA81A01CDD79" blockId="4.[151,1440,151,1544]" pageId="4" pageNumber="595">
Mitochondrial DNA COI barcodes were obtained for the species of 
<taxonomicName id="4C3D4D67F3153E1CFB8BFA80A7CEDE25" authorityName="Bruce" authorityYear="1986" box="[1065,1207,1341,1364]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFB8BFA80A7CEDE25" box="[1065,1207,1341,1364]" italics="true" pageId="4" pageNumber="595">Diapontonia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D4D67F3153E1CFAA1FA81A6ECDE25" authorityName="Bruce" authorityYear="1990" box="[1283,1429,1340,1364]" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFAA1FA81A6ECDE25" box="[1283,1429,1340,1364]" italics="true" pageId="4" pageNumber="595">Altopontonia</emphasis>
</taxonomicName>
. Unfortunately, DNA extraction of 
<typeStatus id="54868846F3153E1CFD8BFADDA1F0DE09" box="[553,649,1376,1400]" pageId="4" pageNumber="595" type="paratype">paratype</typeStatus>
specimens of 
<taxonomicName id="4C3D4D67F3153E1CFC95FADDA766DE09" authorityName="Bruce" authorityYear="1986" box="[823,1055,1376,1400]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B949EAF6F3153E1CFC95FADDA766DE09" box="[823,1055,1376,1400]" italics="true" pageId="4" pageNumber="595">Periclimenes milleri</emphasis>
</taxonomicName>
(RMNH.CRUS.D.36342) and 
<taxonomicName id="4C3D4D67F3153E1CFA25FADCA230DEED" authorityName="Berggren &amp; Svane" authorityYear="1989" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="ingressicolumbi">
<emphasis id="B949EAF6F3153E1CFA25FADCA230DEED" italics="true" pageId="4" pageNumber="595">P. ingressicolumbi</emphasis>
</taxonomicName>
(RMNH.CRUS.D.37738) failed. The genetic distance between 
<taxonomicName id="4C3D4D67F3153E1CFB87FA38A7CBDEED" authorityName="Bruce" authorityYear="1986" box="[1061,1202,1413,1436]" class="Malacostraca" family="Palaemonidae" genus="Diapontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFB87FA38A7CBDEED" box="[1061,1202,1413,1436]" italics="true" pageId="4" pageNumber="595">Diapontonia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D4D67F3153E1CFB4FFA39A606DEED" authorityName="Bruce" authorityYear="1990" box="[1261,1407,1412,1436]" class="Malacostraca" family="Palaemonidae" genus="Altopontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAF6F3153E1CFB4FFA39A606DEED" box="[1261,1407,1412,1436]" italics="true" pageId="4" pageNumber="595">Altopontonia</emphasis>
</taxonomicName>
is 0.25, which is substantial. COI barcodes of the four 
<taxonomicName id="4C3D4D67F3153E1CFCA6FA15A747DEB1" box="[772,1086,1447,1472]" class="Malacostraca" family="Palaemonidae" genus="Echinopericlimenes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B949EAF6F3153E1CFCA6FA15A09BDEB1" box="[772,994,1448,1472]" italics="true" pageId="4" pageNumber="595">Echinopericlimenes</emphasis>
species
</taxonomicName>
were used by 
<bibRefCitation id="EFAC4B15F3153E1CFB52FA1AA398DE94" author="Marin, I. &amp; Chan, T. - Y." pageId="4" pageNumber="595" pagination="301 - 324" refId="ref3284" refString="Marin, I. &amp; Chan, T. - Y. (2014) Deep water echinoid-associated pontoniine shrimp &quot; Periclimenes hertwigi Balss, 1913 &quot; species group (Crustacea: Decapoda: Caridea: Palaemonidae): species review, description of a new genus and species from Philippines. Zootaxa, 3835 (3), 301 - 324. http: // dx. doi. org / 10.11646 / zootaxa. 3835.3.1" type="journal article" year="2014">Marin &amp; Chan (2014)</bibRefCitation>
to show the molecular distinction between these species. Unfortunately these COI barcodes are not yet available on GenBank for comparison with the present material.
</paragraph>
</subSubSection>
</treatment>
</document>