309 lines
38 KiB
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309 lines
38 KiB
XML
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<mods:title id="11EF2A6D7732A1B3ED01A02F1ECB4053">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:date id="87CD7755C47820511F99ABDDDA990870">2022</mods:date>
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<emphasis id="B949EAD5FFD9FFF3FFBAFA45696CF9C9" bold="true" box="[151,286,1518,1545]" italics="true" pageId="91" pageNumber="92">‘Clavatula’</emphasis>
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<taxonomicName id="4C3D4D44FFD9FFF3FE08FA456A5CF9C8" ID-CoL="86YTS" authority="Bellardi, 1877" authorityName="Bellardi" authorityYear="1877" box="[293,558,1518,1545]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="91" pageNumber="92" phylum="Mollusca" rank="species" species="inedita">
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<emphasis id="B949EAD5FFD9FFF3FE08FA45690AF9C9" bold="true" box="[293,376,1518,1545]" italics="true" pageId="91" pageNumber="92">inedita</emphasis>
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<bibRefCitation id="EFAC4B36FFD9FFF3FEADFA456A5CF9C8" author="Bellardi, L" box="[384,558,1518,1544]" pageId="91" pageNumber="92" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
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</emphasis>
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<paragraph id="8B8236C7FFD9FFF3FFBAF9B9693FF9ED" blockId="91.[151,558,1518,1581]" box="[151,333,1554,1587]" pageId="91" pageNumber="92">
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<figureCitation id="13062A42FFD9FFF3FFBAF9B9688DF9EC" box="[151,255,1554,1580]" captionStart="FIGURE 29" captionStartId="96.[151,250,1397,1421]" captionTargetBox="[151,1436,236,1373]" captionTargetId="figure-45@96.[151,1436,236,1373]" captionTargetPageId="96" captionText="FIGURE 29 A. ‘Clavatula’ inedita Bellardi, 1877, Letkés (Hungary), private collection of Zoltán Kovács and Zoltán Vicián. B. ‘Clavatula’ rusticula (Mayer-Eymar, 1886), lectotype, S 1615, Grund (Austria). C. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). D1–D2. ‘Clavatula’ orientoromana Báldi, 1960, paratype, M60.146, Szokolya (Hungary). E1–E2. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), paralectotypes, NHMW 1849/0005/0110b, Buituri (Romania).F1–F3. ‘Clavatula’ justinae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0110a, Buituri (Romania). G. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0005/0110c, Buituri (Romania)." figureDoi="http://doi.org/10.5281/zenodo.6399893" httpUri="https://zenodo.org/record/6399893/files/figure.png" pageId="91" pageNumber="92">Figs 29A</figureCitation>
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<figureCitation id="13062A42FFD9FFF3FFD2F98E695BF9EC" box="[255,297,1554,1587]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="91" pageNumber="92">
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<subScript id="17B93482FFD9FFF3FFD2F98E6975F9F3" attach="left" box="[255,263,1573,1587]" fontSize="6" pageId="91" pageNumber="92">1</subScript>
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–A
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</figureCitation>
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<figureCitation id="13062A42FFD9FFF3FE04F98E6943F9F3" box="[297,305,1573,1587]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="91" pageNumber="92">
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<subScript id="17B93482FFD9FFF3FE04F98E6943F9F3" attach="left" box="[297,305,1573,1587]" fontSize="6" pageId="91" pageNumber="92">2</subScript>
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,
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<figureCitation id="13062A42FFD9FFF3FE12F9B9693FF9ED" box="[319,333,1554,1581]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="91" pageNumber="92">5</figureCitation>
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</paragraph>
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<paragraph id="8B8236C7FFD9FFF3FFBAF9FC6A8AF9AF" blockId="91.[151,1391,1623,1739]" box="[151,760,1623,1647]" pageId="91" pageNumber="92">
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<emphasis id="B949EAD5FFD9FFF3FFBAF9FC68D6F9AE" box="[151,164,1623,1646]" italics="true" pageId="91" pageNumber="92">*</emphasis>
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<taxonomicName id="4C3D4D44FFD9FFF3FF87F9FC69FDF9AF" ID-CoL="86YTS" authority="Bell." authorityName="Bellardi" authorityYear="1877" box="[170,399,1623,1647]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="91" pageNumber="92" phylum="Mollusca" rank="species" species="inedita">
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<emphasis id="B949EAD5FFD9FFF3FF87F9FC6928F9AF" box="[170,346,1623,1647]" italics="true" pageId="91" pageNumber="92">Clavatula inedita</emphasis>
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Bell.
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</taxonomicName>
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—
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<treatmentCitation id="0A9C10D6FFD9FFF3FE84F9FC6A19F9AF" author="Bellardi, L" box="[425,619,1623,1647]" page="203" pageId="91" pageNumber="92" year="1877">
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<bibRefCitation id="EFAC4B36FFD9FFF3FE84F9FC6A19F9AF" author="Bellardi, L" box="[425,619,1623,1647]" pageId="91" pageNumber="92" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi 1877: 203</bibRefCitation>
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, pl. 6, fig. 28.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8236C7FFD9FFF3FFBAF9DE6B99F94D" blockId="91.[151,1391,1623,1739]" box="[151,1003,1653,1677]" pageId="91" pageNumber="92">
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<treatmentCitationGroup id="AB2D11E9FFD9FFF3FFBAF9DE6B99F94D" box="[151,1003,1653,1677]" pageId="91" pageNumber="92">
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<taxonomicName id="4C3D4D44FFD9FFF3FFBAF9DE69AEF94D" ID-CoL="86YTS" authority="Bellardi, 1877" authorityName="Bellardi" authorityYear="1877" box="[151,476,1653,1677]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="91" pageNumber="92" phylum="Mollusca" rank="species" species="inedita">
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<emphasis id="B949EAD5FFD9FFF3FFBAF9DE6935F94D" box="[151,327,1653,1677]" italics="true" pageId="91" pageNumber="92">Clavatula inedita</emphasis>
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<bibRefCitation id="EFAC4B36FFD9FFF3FE63F9DE69AEF94D" author="Bellardi, L" box="[334,476,1653,1677]" pageId="91" pageNumber="92" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
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</taxonomicName>
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—
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<treatmentCitation id="0A9C10D6FFD9FFF3FEDBF9DE6B44F94D" author="Ferrero Mortara, E. & Montefameglio, L. & Pavia, G. & Tampieri, R." box="[502,822,1653,1677]" page="80" pageId="91" pageNumber="92" year="1981">
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<bibRefCitation id="EFAC4B36FFD9FFF3FEDBF9DE6B44F94D" author="Ferrero Mortara, E. & Montefameglio, L. & Pavia, G. & Tampieri, R." box="[502,822,1653,1677]" pageId="91" pageNumber="92" pagination="1 - 327" refId="ref131938" refString="Ferrero Mortara, E., Montefameglio, L., Pavia, G. & Tampieri, R. (1981) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. Parte I. Museo Regionale di Scienze Naturali, Catalogo, 6, 1 - 327. [https: // www. biodiversitylibrary. org / item / 269998 # page / 5 / mode / 1 up]" type="journal article" year="1981">
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Ferrero Mortara
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<emphasis id="B949EAD5FFD9FFF3FD8CF9DD6AA7F94D" box="[673,725,1653,1677]" italics="true" pageId="91" pageNumber="92">et al.</emphasis>
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1981: 80
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</bibRefCitation>
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</treatmentCitation>
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, pl. 15, figs 8a–b.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8236C7FFD9FFF3FFBAF93E6BBEF90A" blockId="91.[151,1391,1623,1739]" pageId="91" pageNumber="92">
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<taxonomicName id="4C3D4D44FFD9FFF3FFBAF93E69ADF96C" ID-CoL="7SNP8" authority="(Bellardi, 1877)" baseAuthorityName="Bellardi" baseAuthorityYear="1877" box="[151,479,1684,1708]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="91" pageNumber="92" phylum="Mollusca" rank="species" species="inedita">
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<emphasis id="B949EAD5FFD9FFF3FFBAF93E6945F96C" box="[151,311,1684,1708]" italics="true" pageId="91" pageNumber="92">Perrona inedita</emphasis>
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(
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||
<bibRefCitation id="EFAC4B36FFD9FFF3FE68F93F69AAF96C" author="Bellardi, L" box="[325,472,1684,1708]" pageId="91" pageNumber="92" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
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)
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</taxonomicName>
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<emphasis id="B949EAD5FFD9FFF3FECDF93F698BF96C" bold="true" box="[480,505,1684,1708]" pageId="91" pageNumber="92">—</emphasis>
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<bibRefCitation id="EFAC4B36FFD9FFF3FED4F93F6A9EF96C" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[505,748,1684,1708]" pageId="91" pageNumber="92" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
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||
Landau
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||
<emphasis id="B949EAD5FFD9FFF3FD66F93E6A0CF96C" box="[587,638,1684,1708]" italics="true" pageId="91" pageNumber="92">et al.</emphasis>
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||
2013: 291
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||
</bibRefCitation>
|
||
, pl. 49, figs 10, 11, pl 50, fig. 14, pl. 74, fig. 2, pl. pl. 2, fig. 12.
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||
<taxonomicName id="4C3D4D44FFD9FFF3FFBAF91869ADF90B" ID-CoL="7SNP8" authority="(Bellardi, 1877)" baseAuthorityName="Bellardi" baseAuthorityYear="1877" box="[151,479,1715,1739]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="91" pageNumber="92" phylum="Mollusca" rank="species" species="inedita">
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<emphasis id="B949EAD5FFD9FFF3FFBAF9186945F90B" box="[151,311,1715,1739]" italics="true" pageId="91" pageNumber="92">Perrona inedita</emphasis>
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(
|
||
<bibRefCitation id="EFAC4B36FFD9FFF3FE68F91869AAF90B" author="Bellardi, L" box="[325,472,1715,1739]" pageId="91" pageNumber="92" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B949EAD5FFD9FFF3FECDF918698BF90B" bold="true" box="[480,505,1715,1739]" pageId="91" pageNumber="92">—</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFD9FFF3FED4F9186B67F90A" author="Kovacs, Z. & Vician, Z." box="[505,789,1714,1739]" pageId="91" pageNumber="92" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián 2021: 145</bibRefCitation>
|
||
, pl. 3, figs 15–16.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFD9FFF4FFBAF9526A32FE35" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92" type="materials_examined">
|
||
<paragraph id="8B8236C7FFD9FFF3FFBAF9526C02F8F7" blockId="91.[151,1437,1785,2027]" pageId="91" pageNumber="92">
|
||
<emphasis id="B949EAD5FFD9FFF3FFBAF952693AF8D3" bold="true" box="[151,328,1785,1811]" pageId="91" pageNumber="92">Type material.</emphasis>
|
||
<materialsCitation id="3B553C9AFFD9FFF3FE7FF9526C1EF8F7" country="Italy" county="Museo Regionale di Scienze Naturali Torino" location="Pino Torinese" municipality="Burdigalian" pageId="91" pageNumber="92" specimenCount="1" typeStatus="syntype">
|
||
BS.011.07.073,
|
||
<typeStatus id="54868865FFD9FFF3FD27F9526A16F8D3" box="[522,612,1785,1811]" pageId="91" pageNumber="92" type="syntype">syntype</typeStatus>
|
||
,
|
||
<location id="8EE2601CFFD9FFF3FD5FF9526B7DF8D3" LSID="urn:lsid:plazi:treatment:039487D1FFD9FFF4FFBAFA456DE0FBBD:8EE2601CFFD9FFF3FD5FF9526B7DF8D3" box="[626,783,1785,1811]" country="Italy" county="Museo Regionale di Scienze Naturali Torino" municipality="Burdigalian" name="Pino Torinese" pageId="91" pageNumber="92">Pino Torinese</location>
|
||
(
|
||
<location id="8EE2601CFFD9FFF3FC0CF9526BB1F8D3" LSID="urn:lsid:plazi:treatment:039487D1FFD9FFF4FFBAFA456DE0FBBD:8EE2601CFFD9FFF3FC0CF9526BB1F8D3" box="[801,963,1785,1811]" country="Italy" county="Museo Regionale di Scienze Naturali Torino" municipality="Burdigalian" name="Colli Torinesi" pageId="91" pageNumber="92">Colli Torinesi</location>
|
||
,
|
||
<collectingCountry id="F32A7657FFD9FFF3FCFCF9526C7AF8D4" box="[977,1032,1785,1812]" name="Italy" pageId="91" pageNumber="92">Italy</collectingCountry>
|
||
),
|
||
<collectingMunicipality id="6BE6ACBDFFD9FFF3FB32F9526CD4F8D3" box="[1055,1190,1785,1811]" pageId="91" pageNumber="92">Burdigalian</collectingMunicipality>
|
||
, stored in the
|
||
<collectingCounty id="62E34E4BFFD9FFF3FA7DF9526A45F8F7" pageId="91" pageNumber="92">Museo Regionale di Scienze Naturali Torino</collectingCounty>
|
||
(
|
||
<collectingCountry id="F32A7657FFD9FFF3FD6BF8B66A0CF8F8" box="[582,638,1821,1848]" name="Italy" pageId="91" pageNumber="92">Italy</collectingCountry>
|
||
), illustrated in
|
||
<bibRefCitation id="EFAC4B36FFD9FFF3FC0BF8B66BB9F8F7" author="Bellardi, L" box="[806,971,1821,1847]" pageId="91" pageNumber="92" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi (1877</bibRefCitation>
|
||
, pl. 6, fig. 28)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFD9FFF3FFEAF8EA6BD8F8BF" blockId="91.[151,1437,1785,2027]" pageId="91" pageNumber="92">
|
||
<emphasis id="B949EAD5FFD9FFF3FFEAF8EA6A64F89B" bold="true" box="[199,534,1857,1883]" pageId="91" pageNumber="92">Additional studied material.</emphasis>
|
||
<materialsCitation id="3B553C9AFFD9FFF3FD31F8EA6BD1F845" country="Hungary" county="Zoltan Vician" location="Letkes" municipality="Zoltan Kovacs" pageId="91" pageNumber="92" specimenCount="1">
|
||
<specimenCount id="9D3BFD4EFFD9FFF3FD31F8EA6A19F89B" box="[540,619,1857,1883]" pageId="91" pageNumber="92" type="generic">1 spec.</specimenCount>
|
||
,
|
||
<location id="8EE2601CFFD9FFF3FD5BF8EA6AB3F89B" LSID="urn:lsid:plazi:treatment:039487D1FFD9FFF4FFBAFA456DE0FBBD:8EE2601CFFD9FFF3FD5BF8EA6AB3F89B" box="[630,705,1857,1883]" country="Hungary" county="Zoltan Vician" municipality="Zoltan Kovacs" name="Letkes" pageId="91" pageNumber="92">Letkés</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFD9FFF3FDE3F8EA6B44F89B" box="[718,822,1857,1883]" name="Hungary" pageId="91" pageNumber="92">Hungary</collectingCountry>
|
||
), private collection of
|
||
<collectingMunicipality id="6BE6ACBDFFD9FFF3FB00F8EA6CA2F89B" box="[1069,1232,1857,1883]" pageId="91" pageNumber="92">Zoltán Kovács</collectingMunicipality>
|
||
and
|
||
<collectingCounty id="62E34E4BFFD9FFF3FA28F8EA6DEEF89B" box="[1285,1436,1857,1883]" pageId="91" pageNumber="92">Zoltán Vicián</collectingCounty>
|
||
(illustrated in
|
||
<bibRefCitation id="EFAC4B36FFD9FFF3FE19F8CE6A44F8BF" author="Kovacs, Z. & Vician, Z." box="[308,566,1893,1919]" pageId="91" pageNumber="92" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián 2021</bibRefCitation>
|
||
, pl. 3, figs 15–16), figs 29A
|
||
<subScript id="17B93482FFD9FFF3FC5CF8DC6B0BF845" attach="left" box="[881,889,1911,1925]" fontSize="6" pageId="91" pageNumber="92">1</subScript>
|
||
–A
|
||
<subScript id="17B93482FFD9FFF3FCB6F8DC6BD1F845" attach="left" box="[923,931,1911,1925]" fontSize="6" pageId="91" pageNumber="92">2</subScript>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFD9FFF4FFEAF8226A32FE35" blockId="91.[151,1437,1785,2027]" lastBlockId="92.[151,1437,151,1150]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
|
||
<emphasis id="B949EAD5FFD9FFF3FFEAF82269C2F863" bold="true" box="[199,432,1929,1955]" pageId="91" pageNumber="92">Revised description</emphasis>
|
||
(based on Paratethyan material). Medium sized, broad fusiform with gradate conical spire; apical angle ~36°. Protoconch not preserved. Teleoconch of at least ten whorls (earliest whorls not preserved). Early teleoconch whorls with bipartite sculpture of weak, smooth subsutural cord and weakly beaded suprasutural cord, separated by concave mid-portion. Subsutural collar on later whorls weak, indistinct with faint spiral threads on spire whorls. Mid-portion markedly concave. Shoulder weakly angulated, protruding, distinctly above suture. Suture narrow, linear. Last whorl 64% of total height, subsutural collar rounded, very weakly developed. Subsutural ramp broad, concave. Shoulder elevated, roundly angled, bearing faint, opisthocline axial ribs marking whorl periphery, convex below. Base not delimited, strongly constricted. Siphonal fasciole not delimited, weakly rounded, slightly twisted. Surface smooth except for weak shoulder ribs and one or two subobsolete cords over base. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately deep and wide, symmetrically U-shaped with apex mid-ramp. Base convex, rapidly contracting into long, moderately narrow siphonal canal, slightly deflected to the left. Columella weakly convex, smooth. Columellar and parietal callus weakly thickened, forming broad rim.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFDEFFF4FFEAFE556942FC09" pageId="92" pageNumber="93" type="discussion">
|
||
<paragraph id="8B8236C7FFDEFFF4FFEAFE556D58FC41" blockId="92.[151,1437,151,1150]" pageId="92" pageNumber="93">
|
||
<emphasis id="B949EAD5FFDEFFF4FFEAFE55693EFDD8" bold="true" box="[199,332,510,536]" pageId="92" pageNumber="93">Discussion.</emphasis>
|
||
<emphasis id="B949EAD5FFDEFFF4FE78FE5469A5FDD9" box="[341,471,511,537]" italics="true" pageId="92" pageNumber="93">‘Clavatula’</emphasis>
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FECDFE546AABFDD9" authority="Bellardi, 1877" authorityName="Bellardi" authorityYear="1877" box="[480,729,511,537]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="inedita">
|
||
<emphasis id="B949EAD5FFDEFFF4FECDFE546A5CFDD9" box="[480,558,511,537]" italics="true" pageId="92" pageNumber="93">inedita</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FD1BFE546AABFDD9" author="Bellardi, L" box="[566,729,511,537]" pageId="92" pageNumber="93" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
is based on a shell with high, slender conical spire with very weak, smooth subsutural collar, a concave whorl profile and a strongly protruding shoulder bearing faint, opisthocline beads. Its base is strongly constricted passing rapidly into a long and narrow siphonal canal. The surface is smooth, except for weak opisthocline ribs at the shoulder and even weaker spiral cords on the base. Serravallian specimens from
|
||
<collectingCountry id="F32A7657FFDEFFF4FFF5FD24695BFD69" box="[216,297,655,681]" name="Turkey" pageId="92" pageNumber="93">Turkey</collectingCountry>
|
||
, recorded by
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FEE7FD246ACEFD69" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[458,700,655,681]" pageId="92" pageNumber="93" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
|
||
Landau
|
||
<emphasis id="B949EAD5FFDEFFF4FD04FD246A2CFD69" box="[553,606,655,681]" italics="true" pageId="92" pageNumber="93">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
, correspond to that Italian morphotype. However, placement in
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FFBAFD186881FD0C" box="[151,243,691,716]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFDEFFF4FFBAFD186881FD0C" box="[151,243,691,716]" italics="true" pageId="92" pageNumber="93">Perrona</emphasis>
|
||
</taxonomicName>
|
||
is incorrect, as this species lacks comma-shaped axial riblets on early teleoconch whorls, as is typical for
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FFBAFD7C6938FD31" box="[151,330,727,753]" pageId="92" pageNumber="93">
|
||
<emphasis id="B949EAD5FFDEFFF4FFBAFD7C6881FD30" box="[151,243,727,752]" italics="true" pageId="92" pageNumber="93">Perrona</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FEACFD7C6A20FD31" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[385,594,727,753]" pageId="92" pageNumber="93" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
|
||
Landau
|
||
<emphasis id="B949EAD5FFDEFFF4FEF6FD7C6A63FD31" box="[475,529,727,753]" italics="true" pageId="92" pageNumber="93">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
, pl. 74, fig. 2). The bipartite early teleoconch sculpture excludes a placement in
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FF9BFD50692CFCD5" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[182,350,763,789]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFDEFFF4FF9BFD50692CFCD5" box="[182,350,763,789]" italics="true" pageId="92" pageNumber="93">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
and in the otherwise slightly reminiscent ‘
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FC65FD506BC4FCD5" box="[840,950,763,789]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFDEFFF4FC65FD506BC4FCD5" box="[840,950,763,789]" italics="true" pageId="92" pageNumber="93">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FCEAFD506C4BFCD4" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[967,1081,763,788]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="interrupta">
|
||
<emphasis id="B949EAD5FFDEFFF4FCEAFD506C4BFCD4" box="[967,1081,763,788]" italics="true" pageId="92" pageNumber="93">interrupta</emphasis>
|
||
</taxonomicName>
|
||
group. Therefore, we treat this species as ‘
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FE35FCB469F4FCF9" box="[280,390,799,825]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFDEFFF4FE35FCB469F4FCF9" box="[280,390,799,825]" italics="true" pageId="92" pageNumber="93">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’. The specimen from Letkés (
|
||
<collectingCountry id="F32A7657FFDEFFF4FDF7FCB46B30FCF9" box="[730,834,799,825]" name="Hungary" pageId="92" pageNumber="93">Hungary</collectingCountry>
|
||
), described herein and by
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FB47FCB46D0FFCF9" author="Kovacs, Z. & Vician, Z." box="[1130,1405,799,825]" pageId="92" pageNumber="93" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián (2021)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FFBAFCE8693BFC9D" authorityName="Bellardi" authorityYear="1877" box="[151,329,835,861]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="inedita">
|
||
<emphasis id="B949EAD5FFDEFFF4FFBAFCE8693BFC9D" box="[151,329,835,861]" italics="true" pageId="92" pageNumber="93">Perrona inedita</emphasis>
|
||
</taxonomicName>
|
||
, agrees generally with ‘
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FD75FCE86AB4FC9D" box="[600,710,835,861]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFDEFFF4FD75FCE86AB4FC9D" box="[600,710,835,861]" italics="true" pageId="92" pageNumber="93">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FFDEFFF4FDF8FCE86B51FC9D" authorityName="Bellardi" authorityYear="1877" box="[725,803,835,861]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="inedita">
|
||
<emphasis id="B949EAD5FFDEFFF4FDF8FCE86B51FC9D" box="[725,803,835,861]" italics="true" pageId="92" pageNumber="93">inedita</emphasis>
|
||
</taxonomicName>
|
||
, but has a wider apical angle, a broader last whorl and a shorter siphonal canal. Further material is required to decide on the status of the Paratethyan occurrence.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFDEFFF4FFEAFC216942FC09" blockId="92.[151,1437,151,1150]" pageId="92" pageNumber="93">
|
||
<emphasis id="B949EAD5FFDEFFF4FFEAFC2169DAFC64" bold="true" box="[199,424,906,932]" pageId="92" pageNumber="93">Paleoenvironment.</emphasis>
|
||
The occurrence at Letkés in
|
||
<collectingCountry id="F32A7657FFDEFFF4FC25FC206B18FC65" box="[776,874,907,933]" name="Hungary" pageId="92" pageNumber="93">Hungary</collectingCountry>
|
||
suggests inner neritic environments (
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FA08FC206956FC09" author="Kovacs, Z. & Vician, Z." pageId="92" pageNumber="93" pagination="53 - 100" refId="ref136963" refString="Kovacs, Z. & Vician, Z. (2013) Badenian (middle Miocene) Conoidean (Neogastropoda) fauna from Letkes (N Hungary). Fragmenta Palaeontologica Hungarica, 30, 53 - 100." type="journal article" year="2013">Kovács & Vicián 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFDEFFF4FFEAFC796DE0FBBD" pageId="92" pageNumber="93" type="distribution">
|
||
<paragraph id="8B8236C7FFDEFFF4FFEAFC79693EFBD1" blockId="92.[151,1437,151,1150]" pageId="92" pageNumber="93">
|
||
<emphasis id="B949EAD5FFDEFFF4FFEAFC796A11FC2D" bold="true" box="[199,611,978,1005]" pageId="92" pageNumber="93">Distribution in Central Paratethys.</emphasis>
|
||
Badenian (middle Miocene):
|
||
<emphasis id="B949EAD5FFDEFFF4FC87FC786C18FC2C" box="[938,1130,979,1004]" italics="true" pageId="92" pageNumber="93">Pannonian Basin</emphasis>
|
||
: Letkés (
|
||
<collectingCountry id="F32A7657FFDEFFF4FBE0FC786D40FC2D" box="[1229,1330,979,1005]" name="Hungary" pageId="92" pageNumber="93">Hungary</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FA6AFC78694DFBD1" author="Kovacs, Z. & Vician, Z." pageId="92" pageNumber="93" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFDEFFF4FFEAFBB16892FB99" blockId="92.[151,1437,151,1150]" pageId="92" pageNumber="93">
|
||
<emphasis id="B949EAD5FFDEFFF4FFEAFBB16A7BFBF4" bold="true" box="[199,521,1050,1076]" pageId="92" pageNumber="93">Proto-Mediterranean Sea:</emphasis>
|
||
Burdigalian (early Miocene):
|
||
<emphasis id="B949EAD5FFDEFFF4FC51FBB06C6CFBF4" box="[892,1054,1051,1077]" italics="true" pageId="92" pageNumber="93">Colli Torinesi</emphasis>
|
||
: Pino Torinese (
|
||
<collectingCountry id="F32A7657FFDEFFF4FBC1FBB06D50FBF6" box="[1260,1314,1051,1078]" name="Italy" pageId="92" pageNumber="93">Italy</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FA6DFBB068A1FB99" author="Bellardi, L" pageId="92" pageNumber="93" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi 1877</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFDEFFF4FFEAFBC86DE0FBBD" blockId="92.[151,1437,151,1150]" box="[199,1426,1122,1150]" pageId="92" pageNumber="93">
|
||
Serravallian (middle Miocene):
|
||
<emphasis id="B949EAD5FFDEFFF4FD03FBC86AACFBBC" box="[558,734,1123,1148]" italics="true" pageId="92" pageNumber="93">
|
||
<collectingRegion id="49F9F825FFDEFFF4FD03FBC86AEAFBBC" box="[558,664,1123,1148]" country="Turkey" name="Karaman" pageId="92" pageNumber="93">Karaman</collectingRegion>
|
||
Basin
|
||
</emphasis>
|
||
: Akboðaz, Akpýnar, Tilkikaya (
|
||
<collectingCountry id="F32A7657FFDEFFF4FB69FBC86CE5FBBE" box="[1092,1175,1123,1150]" name="Turkey" pageId="92" pageNumber="93">Turkey</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FFDEFFF4FB83FBC86DF4FBBD" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[1198,1414,1123,1149]" pageId="92" pageNumber="93" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
|
||
Landau
|
||
<emphasis id="B949EAD5FFDEFFF4FA27FBC86D31FBBD" box="[1290,1347,1123,1149]" italics="true" pageId="92" pageNumber="93">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |