283 lines
37 KiB
XML
283 lines
37 KiB
XML
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<mods:title id="36CB33500753C614B705E968E429C0AC">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<taxonomicName id="4C3D4D44FFC7FFEDFFBAFA296A98FA5C" ID-CoL="8YN4D" authority="(Kovacs & Vician, 2021)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Kovacs & Vician" baseAuthorityYear="2021" box="[151,746,1410,1437]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="69" pageNumber="70" phylum="Mollusca" rank="species" species="nemethi" status="comb. nov.">
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<emphasis id="B949EAD5FFC7FFEDFFBAFA2969C6FA5D" bold="true" box="[151,436,1410,1437]" italics="true" pageId="69" pageNumber="70">Megaclavatula nemethi</emphasis>
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(
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<bibRefCitation id="EFAC4B36FFC7FFEDFEEEFA296A90FA5C" author="Kovacs, Z. & Vician, Z." box="[451,738,1410,1437]" pageId="69" pageNumber="70" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián, 2021</bibRefCitation>
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)
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<taxonomicNameLabel id="A27A57AEFFC7FFEDFDDCFA286B05FA5D" box="[753,887,1411,1437]" pageId="69" pageNumber="70" rank="species">nov. comb.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C327654CFFC7FFEDFFBAFA0C69FAFA01" box="[151,392,1446,1479]" pageId="69" pageNumber="70" type="description">
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<paragraph id="8B8236C7FFC7FFEDFFBAFA0C69FAFA01" blockId="69.[151,887,1410,1479]" box="[151,392,1446,1479]" pageId="69" pageNumber="70">
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<figureCitation id="13062A42FFC7FFEDFFBAFA0C688FFA01" box="[151,253,1447,1473]" captionStart="FIGURE 22" captionStartId="71.[151,250,1791,1815]" captionTargetBox="[164,1423,198,1784]" captionTargetId="figure-18@71.[164,1423,198,1785]" captionTargetPageId="71" captionText="FIGURE 22 A1–A3. Megaclavatula laevigata (Eichwald, 1830), NHMW 1846/0037/0308, Baden (Austria). B1–B3. Megaclavatula subsculpta (Schaffer, 1912), lectotype, KM Inv. No. F/1125, Maria Dreieichen (Austria). C1–C2. Megaclavatula nemethi (Kovács & Vicián, 2021), holotype, HNHM, PAL 2021.21.1, Letkés (Hungary). D. Megaclavatula nemethi (Kovács & Vicián, 2021), paratype, NHMW 2021/0002/0003, Letkés (Hungary). E1–E2. Megaclavatula? szontaghyi (Strausz, 1926), holotype, M.116, Bán, Baranya County (Hungary). F. Megaclavatula neudorfensis (Schaffer, 1898), holotype, NHMW 2021/0137/0001, Devínska Nová Ves (Slovakia). G1–G3. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). H1–H3. Megaclavatula pilleri nov. sp., paratype, NHMW 1949/0005/0051b, Baden-Sooss (Austria)." figureDoi="http://doi.org/10.5281/zenodo.6399881" httpUri="https://zenodo.org/record/6399881/files/figure.png" pageId="69" pageNumber="70">Figs 22C</figureCitation>
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<figureCitation id="13062A42FFC7FFEDFFD0FA126954FA01" box="[253,294,1447,1479]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="69" pageNumber="70">
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<subScript id="17B93482FFC7FFEDFFD0FA126977FA07" attach="left" box="[253,261,1465,1479]" fontSize="6" pageId="69" pageNumber="70">1</subScript>
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–C
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</figureCitation>
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<figureCitation id="13062A42FFC7FFEDFE0BFA12695CFA07" box="[294,302,1465,1479]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="69" pageNumber="70">
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<subScript id="17B93482FFC7FFEDFE0BFA12695CFA07" attach="left" box="[294,302,1465,1479]" fontSize="6" pageId="69" pageNumber="70">2</subScript>
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, D,
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<figureCitation id="13062A42FFC7FFEDFE73FA0D691CFA01" box="[350,366,1446,1473]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="69" pageNumber="70">5</figureCitation>
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,
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<figureCitation id="13062A42FFC7FFEDFE57FA0C69FAFA01" box="[378,392,1447,1473]" captionStart="FIGURE 6" captionStartId="16.[151,250,1938,1962]" captionTargetBox="[151,1436,345,1915]" captionTargetId="figure-106@16.[151,1436,345,1915]" captionTargetPageId="16" captionText="FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia)." figureDoi="http://doi.org/10.5281/zenodo.6399841" httpUri="https://zenodo.org/record/6399841/files/figure.png" pageId="69" pageNumber="70">6</figureCitation>
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</paragraph>
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<subSubSection id="C327654CFFC7FFEDFFBAFA406C64F981" pageId="69" pageNumber="70" type="reference_group">
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<paragraph id="8B8236C7FFC7FFEDFFBAFA406960F9E2" blockId="69.[151,1436,1515,1601]" pageId="69" pageNumber="70">
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<taxonomicName id="4C3D4D44FFC7FFEDFFBAFA4069B1F9C4" ID-CoL="1a20f969-690f-419e-bd9f-57a21c145356" authority="Mayer" authorityName="Mayer" box="[151,451,1515,1540]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="69" pageNumber="70" phylum="Mollusca" rank="species" species="taurinensis">
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<emphasis id="B949EAD5FFC7FFEDFFBAFA406905F9C3" box="[151,375,1515,1539]" italics="true" pageId="69" pageNumber="70">Clavatula taurinensis</emphasis>
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Mayer
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</taxonomicName>
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(in Bellardi)—
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<treatmentCitation id="0A9C10D6FFC7FFEDFD45FA406B58F9C3" author="Strausz, L." box="[616,810,1515,1539]" page="403" pageId="69" pageNumber="70" year="1966">
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<bibRefCitation id="EFAC4B36FFC7FFEDFD45FA406B58F9C3" author="Strausz, L." box="[616,810,1515,1539]" pageId="69" pageNumber="70" refId="ref145008" refString="Strausz, L. (1966) Die Miozan-Mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp." type="book" year="1966">Strausz 1966: 403</bibRefCitation>
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</treatmentCitation>
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||
, pl. 14, figs 20–23 [
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<emphasis id="B949EAD5FFC7FFEDFB21FA476C40F9C3" box="[1036,1074,1516,1539]" italics="true" pageId="69" pageNumber="70">non</emphasis>
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<taxonomicName id="4C3D4D44FFC7FFEDFB11FA476975F9E2" ID-CoL="8ZFR4" authority="(Bellardi, 1877)" baseAuthorityName="Bellardi" baseAuthorityYear="1877" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="69" pageNumber="70" phylum="Mollusca" rank="species" species="taurinensis">
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<emphasis id="B949EAD5FFC7FFEDFB11FA476D40F9C3" box="[1084,1330,1516,1539]" italics="true" pageId="69" pageNumber="70">Neoperrona taurinensis</emphasis>
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(
|
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<bibRefCitation id="EFAC4B36FFC7FFEDFA6EFA40688DF9E2" author="Bellardi, L" pageId="69" pageNumber="70" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
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)
|
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</taxonomicName>
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].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8236C7FFC7FFEDFFBAF9826C64F981" blockId="69.[151,1436,1515,1601]" box="[151,1046,1576,1601]" pageId="69" pageNumber="70">
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<treatmentCitationGroup id="AB2D11E9FFC7FFEDFFBAF9826C64F981" box="[151,1046,1576,1601]" pageId="69" pageNumber="70">
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<emphasis id="B949EAD5FFC7FFEDFFBAF9826926F980" box="[151,340,1576,1600]" italics="true" pageId="69" pageNumber="70">
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*
|
||
<taxonomicName id="4C3D4D44FFC7FFEDFF87F9826926F980" ID-CoL="8KTHV" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[170,340,1576,1600]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="69" pageNumber="70" phylum="Mollusca" rank="species" species="nemethi" status="sp. nov.">Perrona nemethi</taxonomicName>
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</emphasis>
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||
<taxonomicNameLabel id="A27A57AEFFC7FFEDFE76F98269E3F980" box="[347,401,1577,1600]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
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<emphasis id="B949EAD5FFC7FFEDFEBCF98269D8F981" bold="true" box="[401,426,1577,1601]" pageId="69" pageNumber="70">—</emphasis>
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<treatmentCitation id="0A9C10D6FFC7FFEDFE87F9826AB4F981" author="Kovacs, Z. & Vician, Z." box="[426,710,1576,1601]" page="147" pageId="69" pageNumber="70" year="2021">
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<bibRefCitation id="EFAC4B36FFC7FFEDFE87F9826AB4F981" author="Kovacs, Z. & Vician, Z." box="[426,710,1576,1601]" pageId="69" pageNumber="70" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián 2021: 147</bibRefCitation>
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</treatmentCitation>
|
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, pl. 3, figs 30–32, pl. 4, figs 1–3.
|
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C327654CFFC7FFEEFFBAF9C4690FFE6D" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="materials_examined">
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<paragraph id="8B8236C7FFC7FFEDFFBAF9C468C7F8D9" blockId="69.[151,1437,1646,2033]" pageId="69" pageNumber="70">
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<emphasis id="B949EAD5FFC7FFEDFFBAF9C4693AF948" bold="true" box="[151,328,1646,1673]" pageId="69" pageNumber="70">Type material.</emphasis>
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<materialsCitation id="3B553C9AFFC7FFEDFE7FF9C46B4DF973" collectionCode="HNHM, MD" country="Hungary" location="Letkes" pageId="69" pageNumber="70" specimenCode="PAL 2021.21" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54868865FFC7FFEDFE7FF9C469CDF949" box="[338,447,1647,1673]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
|
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:
|
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<collectionCode id="ED2CAE02FFC7FFEDFEE1F9C46A56F949" box="[460,548,1647,1673]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="69" pageNumber="70">HNHM</collectionCode>
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||
,
|
||
<specimenCode id="DB9B9EBCFFC7FFEDFD1EF9C46AA7F949" box="[563,725,1647,1673]" collectionCode="PAL" country="Italy" lsid="urn:lsid:biocol.org:col:15292" name="Herbarium Mediterraneum Panormitanum" pageId="69" pageNumber="70" type="Herbarium">PAL 2021.21</specimenCode>
|
||
.1, SL: 36.0 mm,
|
||
<collectionCode id="ED2CAE02FFC7FFEDFCB7F9C46BB9F949" box="[922,971,1647,1673]" country="Germany" name="Museum Donaueschingen" pageId="69" pageNumber="70">MD</collectionCode>
|
||
:
|
||
<quantity id="4CC59B22FFC7FFEDFCF7F9C46C31F949" box="[986,1091,1647,1673]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="69" pageNumber="70" unit="mm" value="12.4">12.4 mm</quantity>
|
||
,
|
||
<location id="8EE2601CFFC7FFEDFB7FF9C46CEFF949" LSID="urn:lsid:plazi:treatment:039487D1FFC7FFEEFFBAFA29693EFBD1:8EE2601CFFC7FFEDFB7FF9C46CEFF949" box="[1106,1181,1647,1673]" country="Hungary" name="Letkes" pageId="69" pageNumber="70">Letkés</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFC7FFEDFB83F9C46D64F949" box="[1198,1302,1647,1673]" name="Hungary" pageId="69" pageNumber="70">Hungary</collectingCountry>
|
||
), illustrated in
|
||
<bibRefCitation id="EFAC4B36FFC7FFEDFF9BF93869BAF96D" author="Kovacs, Z. & Vician, Z." box="[182,456,1683,1709]" pageId="69" pageNumber="70" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián (2021</bibRefCitation>
|
||
, pl. 3, figs 30–31), figs 22C
|
||
<subScript id="17B93482FFC7FFEDFC23F90E6B64F973" attach="left" box="[782,790,1701,1715]" fontSize="6" pageId="69" pageNumber="70">1</subScript>
|
||
–C
|
||
<subScript id="17B93482FFC7FFEDFC1AF90E6B4DF973" attach="left" box="[823,831,1701,1715]" fontSize="6" pageId="69" pageNumber="70">2</subScript>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B553C9AFFC7FFEDFC62F9386DEAF96D" box="[847,1432,1682,1709]" collectionCode="NHMW" country="Hungary" pageId="69" pageNumber="70" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54868865FFC7FFEDFC62F9386BB2F96D" box="[847,960,1683,1709]" pageId="69" pageNumber="70" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectionCode id="ED2CAE02FFC7FFEDFCE0F9386C5BF96D" box="[973,1065,1683,1709]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="69" pageNumber="70" type="Museum">NHMW</collectionCode>
|
||
, 2021/0002/0003, SL: 25.0 mm
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B553C9AFFC7FFEDFFBAF91C6D5DF911" box="[151,1327,1719,1745]" collectionCode="MD" country="Hungary" location="Letkes" municipality="Ac" pageId="69" pageNumber="70" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2CAE02FFC7FFEDFFBAF91C68BAF911" box="[151,200,1719,1745]" country="Germany" name="Museum Donaueschingen" pageId="69" pageNumber="70">MD</collectionCode>
|
||
: 10.0 mm,
|
||
<location id="8EE2601CFFC7FFEDFE64F91C69E6F911" LSID="urn:lsid:plazi:treatment:039487D1FFC7FFEEFFBAFA29693EFBD1:8EE2601CFFC7FFEDFE64F91C69E6F911" box="[329,404,1719,1745]" country="Hungary" municipality="Ac" name="Letkes" pageId="69" pageNumber="70">Letkés</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFC7FFEDFE89F91C6A7EF911" box="[420,524,1719,1745]" name="Hungary" pageId="69" pageNumber="70">Hungary</collectingCountry>
|
||
), illustrated in
|
||
<bibRefCitation id="EFAC4B36FFC7FFEDFD9AF91C6BB5F911" author="Kovacs, Z. & Vician, Z." box="[695,967,1719,1745]" pageId="69" pageNumber="70" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián (2021</bibRefCitation>
|
||
, pl. 4, figs 1–2), figs 22D, 4
|
||
<collectingMunicipality id="6BE6ACBDFFC7FFEDFA21F91C6D5DF911" box="[1292,1327,1719,1745]" pageId="69" pageNumber="70">Ac</collectingMunicipality>
|
||
</materialsCitation>
|
||
;
|
||
<collectionCode id="ED2CAE02FFC7FFEDFA11F91C6DEAF911" box="[1340,1432,1719,1745]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="69" pageNumber="70" type="Museum">NHMW</collectionCode>
|
||
, 2021/0002/0004, SL: 41.0 mm,
|
||
<materialsCitation id="3B553C9AFFC7FFEDFD2CF97068C3F8D9" collectionCode="MD" country="Hungary" location="Letkes" pageId="69" pageNumber="70" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2CAE02FFC7FFEDFD2CF9706A40F935" box="[513,562,1755,1781]" country="Germany" name="Museum Donaueschingen" pageId="69" pageNumber="70">MD</collectionCode>
|
||
:
|
||
<quantity id="4CC59B22FFC7FFEDFD10F9716AD6F935" box="[573,676,1754,1781]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="69" pageNumber="70" unit="mm" value="15.4">15.4 mm</quantity>
|
||
,
|
||
<location id="8EE2601CFFC7FFEDFD9DF9706A89F935" LSID="urn:lsid:plazi:treatment:039487D1FFC7FFEEFFBAFA29693EFBD1:8EE2601CFFC7FFEDFD9DF9706A89F935" box="[688,763,1755,1781]" country="Hungary" name="Letkes" pageId="69" pageNumber="70">Letkés</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFC7FFEDFC26F9706B01F935" box="[779,883,1755,1781]" name="Hungary" pageId="69" pageNumber="70">Hungary</collectingCountry>
|
||
), illustrated in
|
||
<bibRefCitation id="EFAC4B36FFC7FFEDFB36F9706D5BF935" author="Kovacs, Z. & Vician, Z." box="[1051,1321,1755,1781]" pageId="69" pageNumber="70" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián (2021</bibRefCitation>
|
||
, pl. 4, fig. 3)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFC7FFEEFFEAF889690FFE6D" blockId="69.[151,1437,1646,2033]" lastBlockId="70.[151,1437,150,1041]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
|
||
<emphasis id="B949EAD5FFC7FFEDFFEAF88969C9F8FC" bold="true" box="[199,443,1826,1852]" pageId="69" pageNumber="70">Revised description.</emphasis>
|
||
Shell medium-sized, moderately broad fusiform with conical spire, apical angle ~35°. Protoconch not preserved. Teleoconch of at least ten whorls. Early teleoconch whorls straight-sided with tripartite sculpture, subsutural and mid-cord finely beaded, suprasutural cord with larger beads made bifid by intercalation of a spiral groove. On sixth whorl subsutural cord becomes very weak collar, smooth to subobsoletely beaded; midrow of beads fades leaving smooth, very weakly concave mid-portion. Suprasutural beads become opisthocline, axially elongated, subdivided by further narrow spiral cords. Entire surface covered by even finer subobsolete threads. Suture narrowly impressed, linear. Last whorl 65% of total height. Subsutural cord weak, subsutural ramp of mid-width, weakly concave. Shoulder broad, delimited by row of weak tubercles, somewhat inflated at midwhorl. Base weakly delimited, strongly constricted. Siphonal fasciole poorly developed. Sculpture of spiral threads on entire whorl and weak, subobsolete cords below shoulder. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately wide, moderately deep, asymmetrically U-shaped, with apex at mid-ramp. Siphonal canal moderately long, moderately wide, straight, shallowly notched at tip. Columella strongly excavated in upper third, weakly twisted at fasciole, smooth. Columellar and parietal callus thin, adherent, forming indistinct callus rim.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFC4FFEEFFEAFE1D6AC9FC09" pageId="70" pageNumber="71" type="discussion">
|
||
<paragraph id="8B8236C7FFC4FFEEFFEAFE1D6B4FFD69" blockId="70.[151,1437,150,1041]" pageId="70" pageNumber="71">
|
||
<emphasis id="B949EAD5FFC4FFEEFFEAFE1D693EFE10" bold="true" box="[199,332,438,464]" pageId="70" pageNumber="71">Discussion.</emphasis>
|
||
Within
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFE85FE1C6A22FE11" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[424,592,439,465]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFC4FFEEFE85FE1C6A22FE11" box="[424,592,439,465]" italics="true" pageId="70" pageNumber="71">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
this species has the most reduced sculpture. Its tripartite sculpture of early teleoconch whorls excludes a placement in
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFD5DFE706ABEFE34" box="[624,716,475,500]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFC4FFEEFD5DFE706ABEFE34" box="[624,716,475,500]" italics="true" pageId="70" pageNumber="71">Perrona</emphasis>
|
||
</taxonomicName>
|
||
as suggested by
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFCADFE706CFCFE35" author="Kovacs, Z. & Vician, Z." box="[896,1166,475,501]" pageId="70" pageNumber="71" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián (2021)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFBBAFE706DEFFE35" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Kovacs & Vician" baseAuthorityYear="2021" box="[1175,1437,475,501]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="nemethi">
|
||
<emphasis id="B949EAD5FFC4FFEEFBBAFE706DEFFE35" box="[1175,1437,475,501]" italics="true" pageId="70" pageNumber="71">Megaclavatula nemethi</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFE23FE5469B9FDD9" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[270,459,511,537]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="dulaii" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFC4FFEEFE23FE5469B9FDD9" box="[270,459,511,537]" italics="true" pageId="70" pageNumber="71">Tomellana dulaii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFC4FFEEFEFEFE546A5BFDD9" box="[467,553,511,537]" pageId="70" pageNumber="71" rank="species">nov. sp.</taxonomicNameLabel>
|
||
in general shape but has a slightly higher spire and a deeper and narrower anal sinus. Moreover,
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFE76FD8869CAFDFD" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[347,440,547,573]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="dulaii">
|
||
<emphasis id="B949EAD5FFC4FFEEFE76FD8869CAFDFD" box="[347,440,547,573]" italics="true" pageId="70" pageNumber="71">T. dulaii</emphasis>
|
||
</taxonomicName>
|
||
lacks the beads at the abapical suture and lacks a tripartite sculpture of early teleoconch whorls.
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFFDEFDEC6AAFFDA1" authority="(Lamarck, 1822)" authorityName="Lozouet et al." authorityYear="2001" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[243,733,583,609]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="semimarginata">
|
||
<emphasis id="B949EAD5FFC4FFEEFFDEFDEC6A65FDA0" box="[243,535,583,609]" italics="true" pageId="70" pageNumber="71">Tomellana semimarginata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFD0AFDEC6AA6FDA1" author="Lamarck, J. - B. M. de" box="[551,724,583,609]" pageId="70" pageNumber="71" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is also slightly reminiscent of
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFB11FDEC6CCEFDA1" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Kovacs & Vician" baseAuthorityYear="2021" box="[1084,1212,583,609]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="nemethi">
|
||
<emphasis id="B949EAD5FFC4FFEEFB11FDEC6CCEFDA1" box="[1084,1212,583,609]" italics="true" pageId="70" pageNumber="71">M. nemethi</emphasis>
|
||
</taxonomicName>
|
||
, but lacks a beaded spiral cord at the abapical suture, is much larger, has a much higher spire and axial riblets on the early teleoconch whorls. For further discussions see
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFD09FD246B4BFD69" author="Kovacs, Z. & Vician, Z." box="[548,825,655,681]" pageId="70" pageNumber="71" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián (2021)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFC4FFEEFFEAFD186BD0FC41" blockId="70.[151,1437,150,1041]" pageId="70" pageNumber="71">
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFFEAFD1869BCFD0D" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Kovacs & Vician" baseAuthorityYear="2021" box="[199,462,691,717]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="nemethi">
|
||
<emphasis id="B949EAD5FFC4FFEEFFEAFD1869BCFD0D" box="[199,462,691,717]" italics="true" pageId="70" pageNumber="71">Megaclavatula nemethi</emphasis>
|
||
</taxonomicName>
|
||
was already recorded from Letkés by
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFC5BFD186C69FD0E" author="Strausz, L." box="[886,1051,691,718]" pageId="70" pageNumber="71" refId="ref145008" refString="Strausz, L. (1966) Die Miozan-Mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp." type="book" year="1966">Strausz (1966)</bibRefCitation>
|
||
, who confused the specimens with
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFFBAFD7C6A12FD31" authority="(Bellardi, 1877)" baseAuthorityName="Bellardi" baseAuthorityYear="1877" box="[151,608,727,753]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="taurinensis">
|
||
<emphasis id="B949EAD5FFC4FFEEFFBAFD7C69D0FD30" box="[151,418,727,752]" italics="true" pageId="70" pageNumber="71">Neoperrona taurinensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFE9FFD7C6A2BFD31" author="Bellardi, L" box="[434,601,727,753]" pageId="70" pageNumber="71" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, described from the Langhian of
|
||
<collectingCountry id="F32A7657FFC4FFEEFCF7FD7C6C7DFD31" box="[986,1039,727,753]" name="Italy" pageId="70" pageNumber="71">Italy</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFB36FD7C6D57FD30" baseAuthorityName="Bellardi" baseAuthorityYear="1877" box="[1051,1317,727,752]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="taurinensis">
|
||
<emphasis id="B949EAD5FFC4FFEEFB36FD7C6D57FD30" box="[1051,1317,727,752]" italics="true" pageId="70" pageNumber="71">Neoperrona taurinensis</emphasis>
|
||
</taxonomicName>
|
||
is slightly reminiscent of
|
||
<taxonomicName id="4C3D4D44FFC4FFEEFE6BFD5069B5FCD5" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Kovacs & Vician" baseAuthorityYear="2021" box="[326,455,763,789]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="nemethi">
|
||
<emphasis id="B949EAD5FFC4FFEEFE6BFD5069B5FCD5" box="[326,455,763,789]" italics="true" pageId="70" pageNumber="71">M. nemethi</emphasis>
|
||
</taxonomicName>
|
||
in its beads along the abapical suture of early teleoconch whorls, but is much more slender and has a much higher spire (see
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFD44FCB46B4DFCF9" author="Bellardi, L" box="[617,831,799,825]" pageId="70" pageNumber="71" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi 1877: 187</bibRefCitation>
|
||
, pl. 6, fig. 10). The species compared above illustrate the importance of the early teleoconch whorl sculpture in generic placement. All the above species have similar late teleoconch whorl features, but differ markedly in their neanic whorls.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFC4FFEEFFEAFC216AC9FC09" blockId="70.[151,1437,150,1041]" pageId="70" pageNumber="71">
|
||
<emphasis id="B949EAD5FFC4FFEEFFEAFC2169DAFC64" bold="true" box="[199,424,906,932]" pageId="70" pageNumber="71">Paleoenvironment.</emphasis>
|
||
The occurrence at Letkés (
|
||
<collectingCountry id="F32A7657FFC4FFEEFDF1FC206B33FC65" box="[732,833,907,933]" name="Hungary" pageId="70" pageNumber="71">Hungary</collectingCountry>
|
||
) suggests a preference for inner neritic environments in the vicinity of corals (
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFE87FC046ADDFC09" author="Kovacs, Z. & Vician, Z." box="[426,687,943,969]" pageId="70" pageNumber="71" pagination="53 - 100" refId="ref136963" refString="Kovacs, Z. & Vician, Z. (2013) Badenian (middle Miocene) Conoidean (Neogastropoda) fauna from Letkes (N Hungary). Fragmenta Palaeontologica Hungarica, 30, 53 - 100." type="journal article" year="2013">Kovács & Vicián 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFC4FFEEFFEAFC79693EFBD1" pageId="70" pageNumber="71" type="distribution">
|
||
<paragraph id="8B8236C7FFC4FFEEFFEAFC79693EFBD1" blockId="70.[151,1437,150,1041]" pageId="70" pageNumber="71">
|
||
<emphasis id="B949EAD5FFC4FFEEFFEAFC796A16FC2D" bold="true" box="[199,612,978,1005]" pageId="70" pageNumber="71">Distribution in Central Paratethys.</emphasis>
|
||
Badenian (middle Miocene): Pannonian Basin: Letkés (
|
||
<collectingCountry id="F32A7657FFC4FFEEFBE0FC786D40FC2D" box="[1229,1330,979,1005]" name="Hungary" pageId="70" pageNumber="71">Hungary</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FFC4FFEEFA6AFC78694DFBD1" author="Kovacs, Z. & Vician, Z." pageId="70" pageNumber="71" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |