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<mods:title id="8A3987B691177682D303FE83B4ECA88E">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="9A2030A484E99C305539D1445E648D71">Harzhauser, Mathias</mods:namePart>
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<subSubSection id="C327654CFF91FFBBFFBAFE706AB1FE34" box="[151,707,474,501]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8B8236C7FF91FFBBFFBAFE706AB1FE34" blockId="19.[151,707,474,501]" box="[151,707,474,501]" pageId="19" pageNumber="20">
<emphasis id="B949EAD5FF91FFBBFFBAFE706AB1FE34" bold="true" box="[151,707,474,501]" pageId="19" pageNumber="20">
Key to
<taxonomicName id="4C3D4D44FF91FFBBFFD6FE716902FE35" ID-CoL="7NZRZ" authorityName="Lamarck" authorityYear="1801" box="[251,368,474,501]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF91FFBBFFD6FE716902FE35" bold="true" box="[251,368,474,501]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
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groups recognized herein:
</emphasis>
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<subSubSection id="C327654CFF91FFBBFFBAFD8B6DEEFA48" pageId="19" pageNumber="20" type="key">
<paragraph id="8B8236C7FF91FFBBFFBAFD8B6DEEFDAF" blockId="19.[151,1437,544,1416]" pageId="19" pageNumber="20">1. Early whorls bipartite...................................................................................2 Early whorls tripartite.................................................................................. 5 Early whorls with&gt;3 spiral cords......................................................................... 8</paragraph>
<paragraph id="8B8236C7FF91FFBBFFBAFDD36DEEFD6A" blockId="19.[151,1437,544,1416]" pageId="19" pageNumber="20">2. Shell medium-large size, siphonal canal long to moderately long................................................ 3 Shell medium-small, siphonal canal short.................................................................. 4</paragraph>
<paragraph id="8B8236C7FF91FFBBFFBAFD196DEEFC55" blockId="19.[151,1437,544,1416]" pageId="19" pageNumber="20">
3. Profile broad biconic, subsutural cord weak, smooth, suprasutural cord prominent with large beads overrun by weak spiral cords........................................................................................
<taxonomicName id="4C3D4D44FF91FFBBFA13FD7B6951FCC3" ID-CoL="7NZRZ" authority="Profile" authorityName="Profile" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF91FFBBFA13FD7B6DEFFD25" box="[1342,1437,720,741]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
Profile
</taxonomicName>
slender fusiform, subsutural and suprasutural cords of equal strength with indistinct, bifid beads, mid-whorl moderately concave interspace bearing prominent, comma-shaped axial riblets................................
<emphasis id="B949EAD5FF91FFBBFBC1FCA16D18FCDF" box="[1260,1386,778,799]" italics="true" pageId="19" pageNumber="20">bimarginata-</emphasis>
clade Profile with broad conical spire, strongly constricted at base, long siphonal canal, narrow adsutural cords separated by a wide, concave mid-portion.................................................................
<taxonomicName id="4C3D4D44FF91FFBBFB96FCEE6D68FC9A" ID-CoL="7NZRZ" box="[1211,1306,837,858]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF91FFBBFB96FCEE6D68FC9A" box="[1211,1306,837,858]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FF91FFBBFA05FCEE6D2DFC9A" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[1320,1375,837,858]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="juliae">
<emphasis id="B949EAD5FF91FFBBFA05FCEE6D2DFC9A" box="[1320,1375,837,858]" italics="true" pageId="19" pageNumber="20">juliae</emphasis>
</taxonomicName>
group Profile slender fusiform, subsutural and suprasutural cords of equal strength, separated by a broad, smooth, concave midportion.........................................................................
<taxonomicName id="4C3D4D44FF91FFBBFB8CFC2B6D72FC55" ID-CoL="7NZRZ" box="[1185,1280,896,917]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF91FFBBFB8CFC2B6D72FC55" box="[1185,1280,896,917]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
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<taxonomicName id="4C3D4D44FF91FFBBFA21FC2B6D2DFC55" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[1292,1375,896,917]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="camillae">
<emphasis id="B949EAD5FF91FFBBFA21FC2B6D2DFC55" box="[1292,1375,896,917]" italics="true" pageId="19" pageNumber="20">camillae</emphasis>
</taxonomicName>
group
</paragraph>
<paragraph id="8B8236C7FF91FFBBFFBAFC366DEEFC2E" blockId="19.[151,1437,544,1416]" pageId="19" pageNumber="20">
4. Prominent, smooth subsutural spiral cord, coarsely tuberculate suprasutural cord, mid-whorl concave with comma-shaped riblets, later whorls coarsely tubercular........................................................
<taxonomicName id="4C3D4D44FF91FFBBFBD3FC116D4DFC0F" ID-CoL="8NW8G" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1278,1343,954,975]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949EAD5FF91FFBBFBD3FC116D4DFC0F" box="[1278,1343,954,975]" italics="true" pageId="19" pageNumber="20">Olegia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFF91FFBBFA68FC106DEFFC10" box="[1349,1437,955,976]" pageId="19" pageNumber="20" rank="genus">nov. gen.</taxonomicNameLabel>
Profile slender fusiform with high spire, short last whorl...........................................
<emphasis id="B949EAD5FF91FFBBFA23FC736D10FC2D" box="[1294,1378,984,1005]" italics="true" pageId="19" pageNumber="20">caerulea</emphasis>
-clade
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<paragraph id="8B8236C7FF91FFBBFFBAFC5E6DEEFBA2" blockId="19.[151,1437,544,1416]" pageId="19" pageNumber="20">
5. Shell profile bucciniform, with short siphonal canal, entire shell covered in beaded cords or subquadrate tubercles....................................................................................... ..
<taxonomicName id="4C3D4D44FF91FFBBFB8DFBB96D4DFBE7" ID-CoL="8YKXH" authority="Shell" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1184,1343,1042,1063]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B949EAD5FF91FFBBFB8DFBB96D4DFBE7" box="[1184,1343,1042,1063]" italics="true" pageId="19" pageNumber="20">Granulatocincta</emphasis>
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<taxonomicNameLabel id="A27A57AEFF91FFBBFA68FBB86DEFFBE8" box="[1349,1437,1043,1064]" pageId="19" pageNumber="20" rank="genus">nov. gen.</taxonomicNameLabel>
Shell profile fusiform with medium-long siphonal canal....................................................... 6 Shell profile fusiform with tall spire about 50-55% total height, short last whorl.................................... 7
</paragraph>
<paragraph id="8B8236C7FF91FFBBFFBAFBC16DEFFAF0" blockId="19.[151,1437,544,1416]" pageId="19" pageNumber="20">
6. Slender to stout biconic profile, early beginning with weakly opisthocline axial ribs, grading into three spiral rows of beads, suprasutural spiral row most prominent..........................................................
<emphasis id="B949EAD5FF91FFBBFA35FB236D1BFB5D" box="[1304,1385,1160,1181]" italics="true" pageId="19" pageNumber="20">mystica-</emphasis>
clade Moderately slender fusiform profile, prominently beaded adsutural cords, weaker beaded mid-whorl cord, subsutural collar coronate with pointed tubercles or spines, base with weak sculpture, siphonal canal moderately long to long..............................................................................................
<taxonomicName id="4C3D4D44FF91FFBBFB82FB4B6D4DFB35" ID-CoL="8YKXM" authority="Opisthocline" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1199,1343,1248,1269]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B949EAD5FF91FFBBFB82FB4B6D4DFB35" box="[1199,1343,1248,1269]" italics="true" pageId="19" pageNumber="20">Megaclavatula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFF91FFBBFA68FB4B6DEFFB35" box="[1349,1437,1248,1269]" pageId="19" pageNumber="20" rank="genus">nov. gen.</taxonomicNameLabel>
Opisthocline axial ribs on the first teleoconch whorl, weak subsutural spiral cord on second teleoconch whorl, beaded suprasutural spiral cord evolves from the abapical tips of the axial ribs on subsequent whorls..............
<emphasis id="B949EAD5FF91FFBBFA22FAB16D18FAEF" box="[1295,1386,1306,1327]" italics="true" pageId="19" pageNumber="20">diadema-</emphasis>
clade
</paragraph>
<paragraph id="8B8236C7FF91FFBBFFBAFA936DEFFA8D" blockId="19.[151,1437,544,1416]" box="[151,1437,1336,1358]" pageId="19" pageNumber="20">
7. Tripartite earliest whorl sculpture, weakly beaded subsutural, strongly beaded suprasutural cord.......
<taxonomicName id="4C3D4D44FF91FFBBFBE1FA936D4DFA8D" ID-CoL="8ZH4Q" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1228,1343,1336,1357]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B949EAD5FF91FFBBFBE1FA936D4DFA8D" box="[1228,1343,1336,1357]" italics="true" pageId="19" pageNumber="20">Neoperrona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFF91FFBBFA68FA936DEFFA8D" box="[1349,1437,1336,1357]" pageId="19" pageNumber="20" rank="genus">nov. gen.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B8236C7FF91FFBBFFBAFAFE6DEEFA48" blockId="19.[151,1437,544,1416]" pageId="19" pageNumber="20">
8. Close-set spiral cords on flat-sided early teleoconch whorls comparatively slender, smooth and high spired shells, and the weak subsutural collar, which lacks spines..............................................................
<emphasis id="B949EAD5FF91FFBBFA03FAD86D10FA48" box="[1326,1378,1395,1416]" italics="true" pageId="19" pageNumber="20">taxea</emphasis>
-clade
</paragraph>
</subSubSection>
</treatment>
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