treatments-xml/data/03/94/87/039487C7E52CFFBF599AA37BFE6CFADA.xml
2024-06-21 12:22:17 +02:00

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<document id="109D292DED25DA84AFAB5BD6F2EA2631" ID-DOI="10.15468/k7klsp" ID-ISSN="1407-8619" ID-Zenodo-Dep="10667535" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708023866638" checkinUser="felipe" docAuthor="Alekseev, Vitalii I." docDate="2016" docId="039487C7E52CFFBF599AA37BFE6CFADA" docLanguage="en" docName="BalticJColeopterol.16.1.21-26.pdf" docOrigin="Baltic Journal of Coleopterology 16 (1)" docSource="http://dx.doi.org/10.15468/k7klsp" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.1:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="1" docTitle="Clypastraea primainterpares Alekseev 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="24" masterDocId="FFADFFBFE52DFFBC5979A634FFAEFFCD" masterDocTitle="New records and species of Crepidodera Chevrolat (Coleoptera: Chrysomelidae) in Eocene European amber, with a brief review of described fossil beetles from Bitterfeld amber" masterLastPageNumber="26" masterPageNumber="21" pageNumber="22" updateTime="1708024172291" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="E733D0F6B1912D08828017B10315931F">New records and species of Crepidodera Chevrolat (Coleoptera: Chrysomelidae) in Eocene European amber, with a brief review of described fossil beetles from Bitterfeld amber</mods:title>
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<mods:namePart id="597A642845B59DDEAD8961BB3BA971CD">Alekseev, Vitalii I.</mods:namePart>
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<mods:title id="BC7C7B0BEE1B92377F09CCD155762727">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="A413258CCBF2451AE1EBA1F84B731FF8">2016</mods:date>
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<mods:identifier id="2E1FE06CC39CB207F04AF29F133ED34D" type="DOI">10.15468/k7klsp</mods:identifier>
<mods:identifier id="3BF12D133F2A3E927F106B6BDE07835D" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="251E45AD597F7CC8A7D7237524D66EC6" type="Zenodo-Dep">10667535</mods:identifier>
</mods:mods>
<treatment id="039487C7E52CFFBF599AA37BFE6CFADA" LSID="urn:lsid:plazi:treatment:039487C7E52CFFBF599AA37BFE6CFADA" httpUri="http://treatment.plazi.org/id/039487C7E52CFFBF599AA37BFE6CFADA" lastPageId="3" lastPageNumber="24" pageId="1" pageNumber="22">
<subSubSection id="C327655AE52CFFBD599AA37BFEC8FA4A" pageId="1" pageNumber="22" type="nomenclature">
<paragraph id="8B8236D1E52CFFBD599AA37BFDB2FAAA" blockId="1.[227,540,1359,1415]" box="[227,540,1359,1383]" pageId="1" pageNumber="22">
<heading id="D0CA81BDE52CFFBD599AA37BFDB2FAAA" bold="true" box="[227,540,1359,1383]" fontSize="10" level="2" pageId="1" pageNumber="22" reason="4">
<emphasis id="B949EAC3E52CFFBD599AA37BFDB2FAAA" bold="true" box="[227,540,1359,1383]" pageId="1" pageNumber="22">
<taxonomicName id="4C3D4D52E52CFFBD599AA37BFE14FAAA" authority="Alekseev, 2016" authorityName="Alekseev" authorityYear="2016" box="[227,442,1359,1383]" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="primainterpares" status="sp. nov.">
<emphasis id="B949EAC3E52CFFBD599AA37BFE14FAAA" bold="true" box="[227,442,1359,1383]" italics="true" pageId="1" pageNumber="22">C. primainterpares</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B8E52CFFBD58BAA37BFDB2FAAA" box="[451,540,1359,1383]" pageId="1" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236D1E52CFFBD599AA35BFEC8FA4A" blockId="1.[227,540,1359,1415]" box="[227,358,1391,1415]" pageId="1" pageNumber="22">(Figs. 1 - 4)</paragraph>
</subSubSection>
<subSubSection id="C327655AE52CFFBD599AA39BFB0BFEBA" pageId="1" pageNumber="22" type="materials_examined">
<paragraph id="8B8236D1E52CFFBD599AA39BFE56F9AA" blockId="1.[227,737,1455,1736]" pageId="1" pageNumber="22">
<emphasis id="B949EAC3E52CFFBD599AA39BFE7EFA0A" bold="true" box="[227,464,1455,1479]" pageId="1" pageNumber="22">Material examined</emphasis>
.
<materialsCitation id="3B553C8CE52CFFBD589DA39BFE56F9AA" collectionCode="CCHH" pageId="1" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="54868873E52CFFBD589DA39BFDE3FA0A" box="[484,589,1455,1479]" pageId="1" pageNumber="22" type="holotype">Holotype</typeStatus>
No. 1615-3 [
<collectionCode id="ED2CAE14E52CFFBD5990A3FBFE95FA2A" box="[233,315,1487,1511]" pageId="1" pageNumber="22">CCHH</collectionCode>
], adult, possibly female (evenly curved apical sternite). Inclusion in a small yellow amber piece, embedded in a block of GTSpolyester resin measuring 15 x 7 x
<quantity id="4CC59B34E52CFFBD5BEAA01BFD72F98A" box="[659,732,1583,1607]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="22" unit="mm" value="4.0">4 mm</quantity>
. Syninclusions are absent.
</materialsCitation>
</paragraph>
<paragraph id="8B8236D1E52CFFBD599AA05BFB84FF3A" blockId="1.[227,737,1455,1736]" lastBlockId="1.[774,1285,191,247]" pageId="1" pageNumber="22">
The amber piece was obtained from a commercial source and will be deposited at the Senckenber g Deutsch es Entomologisch es Institut (Müncheberg,
<collectingCountry id="F32A7641E52CFFBD5A84A68BFBCAFF1A" box="[1021,1124,191,215]" name="Germany" pageId="1" pageNumber="22">Germany</collectingCountry>
) as part of the institute amber collection.
</paragraph>
<paragraph id="8B8236D1E52CFFBD5A7FA72BFB23FEFA" blockId="1.[774,1284,287,375]" box="[774,1165,287,311]" pageId="1" pageNumber="22">
<emphasis id="B949EAC3E52CFFBD5A7FA72BFC3CFEFA" bold="true" box="[774,914,287,311]" pageId="1" pageNumber="22">
<typeStatus id="54868873E52CFFBD5A7FA72BFC91FEFA" box="[774,831,287,311]" pageId="1" pageNumber="22">Type</typeStatus>
strata.
</emphasis>
Baltic Amber, Eocene.
</paragraph>
<paragraph id="8B8236D1E52CFFBD5A7FA70BFB0BFEBA" blockId="1.[774,1284,287,375]" pageId="1" pageNumber="22">
<materialsCitation id="3B553C8CE52CFFBD5A7FA70BFB0CFEBA" country="Russia" location="Yantarny" municipality="Sambian" pageId="1" pageNumber="22" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B949EAC3E52CFFBD5A7FA70BFC08FE9A" bold="true" box="[774,934,319,343]" pageId="1" pageNumber="22">
<typeStatus id="54868873E52CFFBD5A7FA70BFCEEFE9A" box="[774,832,319,343]" pageId="1" pageNumber="22">Type</typeStatus>
locality.
</emphasis>
<location id="8EE2600AE52CFFBD5AC9A774FBBBFE95" LSID="urn:lsid:plazi:treatment:039487C7E52CFFBF599AA37BFE6CFADA:8EE2600AE52CFFBD5AC9A774FBBBFE95" box="[944,1045,320,344]" country="Russia" municipality="Sambian" name="Yantarny" pageId="1" pageNumber="22" stateProvince="Kaliningrad">Yantarny</location>
settlement,
<collectingMunicipality id="6BE6ACABE52CFFBD5DDDA70BFAAAFE9A" box="[1188,1284,319,343]" pageId="1" pageNumber="22">Sambian</collectingMunicipality>
peninsula,
<collectingRegion id="49F9F833E52CFFBD5A02A76BFBE3FEBA" box="[891,1101,351,375]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="22">Kaliningrad region</collectingRegion>
,
<collectingCountry id="F32A7641E52CFFBD5D2EA76BFB0CFEBA" box="[1111,1186,351,375]" name="Russia" pageId="1" pageNumber="22">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C327655AE52CFFBD5A7FA7ABFC09FB3A" pageId="1" pageNumber="22" type="diagnosis">
<paragraph id="8B8236D1E52CFFBD5A7FA7ABFBBCFE7A" blockId="1.[774,1285,415,855]" box="[774,1042,415,439]" pageId="1" pageNumber="22">
<emphasis id="B949EAC3E52CFFBD5A7FA7ABFBBCFE7A" bold="true" box="[774,1042,415,439]" pageId="1" pageNumber="22">Differential diagnosis.</emphasis>
</paragraph>
<paragraph id="8B8236D1E52CFFBD5A7FA78BFC6CFC9A" blockId="1.[774,1285,415,855]" pageId="1" pageNumber="22">
The specimen under study corresponds to the diagnosis of the subfamily
<taxonomicName id="4C3D4D52E52CFFBD5D59A7EBFB1FFE3A" authorityName="LeConte" authorityYear="1852" box="[1056,1201,479,503]" class="Insecta" family="Corylophidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Corylophinae">Corylophinae</taxonomicName>
and the tribe
<taxonomicName id="4C3D4D52E52CFFBD5A44A7CBFB39FDDA" authority="(Slipinski et al. 2009)" baseAuthorityName="Slipinski" baseAuthorityYear="2009" box="[829,1175,511,535]" class="Insecta" family="Corylophidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Parmulini">
Parmulini (
<bibRefCitation id="EFAC4B20E52CFFBD5ACCA7CBFB21FDDA" author="Slipinski A. &amp; Tomaszewska W. &amp; Lawrence J. F." box="[949,1167,511,535]" pageId="1" pageNumber="22" pagination="409 - 433" refId="ref2923" refString="Slipinski A., Tomaszewska W., Lawrence J. F. 2009. Phylogeny and classification of Corylophidae (Coleoptera: Cucujoidea) with descr iption s of n ew gen er a an d lar vae. Systematic Entomology. 34: 409 - 433." type="journal article" year="2009">Ślipiński et al. 2009</bibRefCitation>
)
</taxonomicName>
. However, mandibles, labial palps, details of head, internal characters of abdomen and venation of hind wings are invisible in this specimen. The characters of the genus (
<bibRefCitation id="EFAC4B20E52CFFBD5D63A44BFB7FFD5A" author="Bowestead S." box="[1050,1233,639,663]" pageId="1" pageNumber="22" pagination="1 - 203" refId="ref2478" refString="Bowestead S. 1999. A revision of the Corylophidae (Coleoptera) of the West Palaearctic Region. Instrumenta Biodiversitatis. 3: 1 - 203." type="journal article" year="1999">Bowestead 1999</bibRefCitation>
) are well discernible: anterior margin of prosternum sharply emarginated by antennal slots, leaving a truncate median plate; antennae 11-segmented, with well-formed club. These characters and the large body fit with the genus
<taxonomicName id="4C3D4D52E52CFFBD5D0FA52BFC10FC9A" authority="Haldeman, 1842" authorityName="Haldeman" authorityYear="1842" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAC3E52CFFBD5D0FA52BFAABFCFA" box="[1142,1285,799,823]" italics="true" pageId="1" pageNumber="22">Clypastraea</emphasis>
Haldeman, 1842
</taxonomicName>
.
</paragraph>
<paragraph id="8B8236D1E52CFFBD5A7FA54BFC09FB3A" blockId="1.[774,1286,895,1271]" pageId="1" pageNumber="22">
The narrow sutural interval indicates the probable placement of the newly described species into the
<taxonomicName id="4C3D4D52E52CFFBD5A8CA58BFBC0FC1A" baseAuthorityName="Reitter" baseAuthorityYear="1877" box="[1013,1134,959,983]" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B949EAC3E52CFFBD5A8CA58BFBC0FC1A" box="[1013,1134,959,983]" italics="true" pageId="1" pageNumber="22">orientalis</emphasis>
</taxonomicName>
group.
<taxonomicName id="4C3D4D52E52CFFBD5D93A5F4FC64FC3A" authority="Alekseev, 2016" authorityName="Alekseev" authorityYear="2016" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="primainterpares" status="sp. nov.">
<emphasis id="B949EAC3E52CFFBD5D93A5F4FC64FC3A" italics="true" pageId="1" pageNumber="22">C. primainterpares</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B8E52CFFBD5AA1A5D4FB95FC3A" box="[984,1083,992,1015]" pageId="1" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C3D4D52E52CFFBD5D93A5D4FB91FBDA" authority="(Kraatz, 1869)" baseAuthorityName="Kraatz" baseAuthorityYear="1869" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="maderae">
<emphasis id="B949EAC3E52CFFBD5D93A5D4FCC1FBDA" italics="true" pageId="1" pageNumber="22">C. maderae</emphasis>
(Kraatz, 1869)
</taxonomicName>
in the smooth interspaces between punctures and from
<taxonomicName id="4C3D4D52E52CFFBD5D92A214FBA7FB9A" authority="Bowestead, 1999" authorityName="Bowestead" authorityYear="1999" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="palmi">
<emphasis id="B949EAC3E52CFFBD5D92A214FCEAFB9A" italics="true" pageId="1" pageNumber="22">C. palmi</emphasis>
<bibRefCitation id="EFAC4B20E52CFFBD5A34A20BFBA7FB9A" author="Bowestead S." box="[845,1033,1087,1111]" pageId="1" pageNumber="22" pagination="1 - 203" refId="ref2478" refString="Bowestead S. 1999. A revision of the Corylophidae (Coleoptera) of the West Palaearctic Region. Instrumenta Biodiversitatis. 3: 1 - 203." type="journal article" year="1999">
Bowestead,
<quantity id="4CC59B34E52CFFBD5AAAA20BFB86FB9A" box="[979,1064,1087,1111]" metricMagnitude="1" metricUnit="m" metricValue="5.07746" pageId="1" pageNumber="22" unit="in" value="1999.0">1999 in</quantity>
</bibRefCitation>
</taxonomicName>
the fully developed hind wings and the comparatively long elytra. The Eocene beetle shows a distinctly larger body length than the recent south-European
<taxonomicName id="4C3D4D52E52CFFBD5D93A294FBB1FB1A" authority="(Reitter, 1877)" baseAuthorityName="Reitter" baseAuthorityYear="1877" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B949EAC3E52CFFBD5D93A294FCC1FB1A" italics="true" pageId="1" pageNumber="22">C. orientalis</emphasis>
(Reitter, 1877)
</taxonomicName>
, which is only
<quantity id="4CC59B34E52CFFBD5DB6A28BFCC6FB3A" metricMagnitude="-3" metricUnit="m" metricValue="1.28" metricValueMax="1.46" metricValueMin="1.1" pageId="1" pageNumber="22" unit="mm" value="1.28" valueMax="1.46" valueMin="1.1">1.1 - 1.46 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C327655AE52CFFBF5A7FA32BFDE6FBDA" lastPageId="3" lastPageNumber="24" pageId="1" pageNumber="22" type="description">
<paragraph id="8B8236D1E52CFFBD5A7FA32BFC35FAFA" blockId="1.[774,1284,1311,1495]" box="[774,923,1311,1335]" pageId="1" pageNumber="22">
<emphasis id="B949EAC3E52CFFBD5A7FA32BFC3AFAFA" bold="true" box="[774,916,1311,1335]" pageId="1" pageNumber="22">Description</emphasis>
.
</paragraph>
<paragraph id="8B8236D1E52CFFBD5A7FA30BFB5EFA1A" blockId="1.[774,1284,1311,1495]" pageId="1" pageNumber="22">
<emphasis id="B949EAC3E52CFFBD5A7FA30BFCF2FA9A" box="[774,860,1343,1367]" italics="true" pageId="1" pageNumber="22">Habitus</emphasis>
. Large (body length
<quantity id="4CC59B34E52CFFBD5D40A30BFB35FA9A" box="[1081,1179,1343,1367]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="1" pageNumber="22" unit="mm" value="1.67">1.67 mm</quantity>
, maximal body width
<quantity id="4CC59B34E52CFFBD5AFAA36BFC4AFABA" box="[899,996,1375,1399]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="22" unit="mm" value="0.91">0.91 mm</quantity>
, maximal body height
<quantity id="4CC59B34E52CFFBD5DACA36BFC98FA5A" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="1" pageNumber="22" unit="mm" value="0.57">0.57 mm</quantity>
), elongate, oval, moderately depressed, shortly pubescent. Color: entirely brown, slightly darker at center of the pronotal disc.
</paragraph>
<paragraph id="8B8236D1E52CFFBD5A7FA3CBFAABF91A" blockId="1.[774,1286,1535,1752]" pageId="1" pageNumber="22">
<emphasis id="B949EAC3E52CFFBD5A7FA3CBFCEFF9DA" box="[774,833,1535,1559]" italics="true" pageId="1" pageNumber="22">Head</emphasis>
. Concealed, completely covered by the pronotum. Eyes large, oval. Mentum subquadrate. Submentum very short and broad. Antennae doubly geniculate in repose, 11- segmented, with a 3-segmented, weakly asymmetrical club. Pedicel and scape subequal in length. First and second segments of the club
</paragraph>
<paragraph id="8B8236D1E52FFFBE5D3DA77EFB4FFD8F" blockId="2.[1092,1354,330,578]" pageId="2" pageNumber="23">
Figures 1-3.
<taxonomicName id="4C3D4D52E52FFFBE5D3DA75EFB59FE6E" authority="Alekseev, 2016" authorityName="Alekseev" authorityYear="2016" box="[1092,1271,362,419]" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="primainterpares" status="sp. nov.">
<emphasis id="B949EAC3E52FFFBE5D3DA75EFB59FE6E" box="[1092,1271,362,419]" italics="true" pageId="2" pageNumber="23">Clypastraea primainterpares</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B8E52FFFBE5C7AA7BFFBDDFE0F" pageId="2" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
<typeStatus id="54868873E52FFFBE5D03A79EFB4EFE0F" box="[1146,1248,426,450]" pageId="2" pageNumber="23" type="holotype">Holotype</typeStatus>
; No. 1615-3 [
<collectionCode id="ED2CAE14E52FFFBE5DDBA7FEFB5AFE2F" box="[1186,1268,458,482]" pageId="2" pageNumber="23">CCHH</collectionCode>
]. Habitus: 1 - Dorsal view; 2 - Ventral view; 3 - Lateral view.
</paragraph>
<caption id="DF426659E52FFFBE5856A20FFE79FB7E" ID-DOI="http://doi.org/10.5281/zenodo.10667537" ID-Zenodo-Dep="10667537" httpUri="https://zenodo.org/record/10667537/files/figure.png" pageId="2" pageNumber="23" startId="2.[303,374,1083,1107]" targetBox="[322,541,674,1015]" targetPageId="2" targetType="figure">
<paragraph id="8B8236D1E52FFFBE5856A20FFB96FBBE" blockId="2.[303,1265,1083,1203]" pageId="2" pageNumber="23">
Figure 4.
<taxonomicName id="4C3D4D52E52FFFBE58D9A20FFD8BFB9E" authorityName="Haldeman" authorityYear="1842" box="[416,549,1083,1107]" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAC3E52FFFBE58D9A20FFD8BFB9E" box="[416,549,1083,1107]" italics="true" pageId="2" pageNumber="23">Clypastraea</emphasis>
</taxonomicName>
Figures 5-6.
<taxonomicName id="4C3D4D52E52FFFBE5B82A20FFC3CFB9E" authorityName="LeConte" authorityYear="1852" box="[763,914,1083,1107]" class="Insecta" family="Corylophidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Corylophinae">Corylophinae</taxonomicName>
. Specimen No 1303-2 [CCHH].
<taxonomicName id="4C3D4D52E52FFFBE5856A268FE71FBBE" authority="Alekseev, 2016" authorityName="Alekseev" authorityYear="2016" box="[303,479,1116,1139]" class="Insecta" family="Corylophidae" genus="Clypastraea" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="primainterpares" status="sp. nov.">
<emphasis id="B949EAC3E52FFFBE5856A268FE71FBBE" box="[303,479,1116,1139]" italics="true" pageId="2" pageNumber="23">primainterpares</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B8E52FFFBE5891A268FDEFFBBE" box="[488,577,1116,1139]" pageId="2" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
Habitus: 1 - Ventral view; 2 - Dorsal view.
</paragraph>
<paragraph id="8B8236D1E52FFFBE5856A24FFE43FB5E" blockId="2.[303,1265,1083,1203]" box="[303,493,1147,1171]" pageId="2" pageNumber="23">Habitus ventrally,</paragraph>
<paragraph id="8B8236D1E52FFFBE5856A2AFFE79FB7E" blockId="2.[303,1265,1083,1203]" box="[303,471,1179,1203]" pageId="2" pageNumber="23">reconstruction.</paragraph>
</caption>
<paragraph id="8B8236D1E52FFFBE5856A2D7FDFAFA96" blockId="2.[303,813,1251,1371]" pageId="2" pageNumber="23">subtriangular; apical segment of the club oval, broad and rounded apically. Ratio of antennomere lengths: 5.0: 5.0: 2.0: 1.0: 1.0: 1.0: 2.0: 1.0: 4.0: 4.0: 5.0.</paragraph>
<paragraph id="8B8236D1E52FFFBE5856A3B7FBE2F9CD" blockId="2.[303,814,1411,1755]" lastBlockId="2.[851,1362,1251,1536]" pageId="2" pageNumber="23">
<emphasis id="B949EAC3E52FFFBE5856A3B7FED3FA56" box="[303,381,1411,1435]" italics="true" pageId="2" pageNumber="23">Thorax</emphasis>
. Pronotum finely bordered; transverse (median length
<quantity id="4CC59B34E52FFFBE58AFA397FD96FA76" box="[470,568,1443,1467]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="23" unit="mm" value="0.48">0.48 mm</quantity>
; basal width
<quantity id="4CC59B34E52FFFBE5BC6A397FC8EFA76" box="[703,800,1443,1467]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="23" unit="mm" value="0.85">0.85 mm</quantity>
), with broadly arcuate anterior margin and slightly bisinuate posterior margin. Anterior angles absent, posterior angles straight. Pronotal punctuation dense, distinct (especially basally), round, separated by distance almost equal to that of a puncture diameter. Prosternal process flat, narrow between coxae, expanding posteriorly into a triangle meeting postcoxal projections. Anterior margin of the prosternum emarginated by antennal slots, leaving a truncate median plate. Mesepisternum and mesepimeron not separated by a suture. Sinuate line at base of the mesosternum not visible due to position of legs. Metepisternum narrowly triangular, slightly shorter than epipleuron, reaching the metacoxal cavities. Postcoxal lines absent. Scutellum triangular, rounded apically, transverse (1.4 times as wide as long).
</paragraph>
<paragraph id="8B8236D1E52FFFBE5A2AA01DFAFEF92D" blockId="2.[851,1362,1576,1760]" pageId="2" pageNumber="23">
<emphasis id="B949EAC3E52FFFBE5A2AA01DFC32F98D" box="[851,924,1577,1600]" italics="true" pageId="2" pageNumber="23">Wings.</emphasis>
Elytra broadest in anterior third; almost as wide as pronotal base anteriorly; elongate (sutural length
<quantity id="4CC59B34E52FFFBE5A84A05CFBCAF94D" box="[1021,1124,1640,1664]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="2" pageNumber="23" unit="mm" value="1.14">1.14 mm</quantity>
, maximal width
<quantity id="4CC59B34E52FFFBE5C59A05CFC2CF96D" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="23" unit="mm" value="0.91">0.91 mm</quantity>
), separately rounded apically; irregularly punctured and recumbently pubescent. Puncture of elytra longitudinally oval, fine and
</paragraph>
<paragraph id="8B8236D1E52EFFBF5BC4A6A5FC83FF69" blockId="3.[701,813,145,164]" box="[701,813,145,164]" pageId="3" pageNumber="24">
<emphasis id="B949EAC3E52EFFBF5BC4A6A5FC83FF69" box="[701,813,145,164]" italics="true" pageId="3" pageNumber="24">Alekseev V.I.</emphasis>
</paragraph>
<paragraph id="8B8236D1E52EFFBF599AA68BFD0CFE3A" blockId="3.[227,737,191,503]" pageId="3" pageNumber="24">comparatively sparse, separated by distance 1.5- 2 times as diameter of punctures, with a long pubescence overlapping the next row of punctures, interspaces smooth and shining. Sutural stria long (two-thirds of the elytral length apically, not discernible in the basal third). Sutural interval narrow, with one row of punctures. Epipleura present, widest at humeral area; reaching the first ventrite. Metathoracical wings fully developed, partially exposed.</paragraph>
<paragraph id="8B8236D1E52EFFBF599AA414FEDDFCBA" blockId="3.[227,738,543,887]" pageId="3" pageNumber="24">
<emphasis id="B949EAC3E52EFFBF599AA414FEB8FDFA" box="[227,278,544,567]" italics="true" pageId="3" pageNumber="24">Legs</emphasis>
. Pro- and mesocoxae rounded; procoxae narrowly separated by the prosternal process; distance between mesocoxae twice broader than mesocoxal diameter; metacoxae str on gly transverse, separated by distance 4 x that of the metacoxal diameter. Femora broad, flattened. Tibiae slender. Tarsi 4-segmented; last metatarsomere as long as tarsomeres 1 - 3 combined. Claws simple, long (three times sh or ter th an metatar somer e 4), cur ved, symmetrical.
</paragraph>
<paragraph id="8B8236D1E52EFFBF599AA5ABFDE6FBDA" blockId="3.[227,737,927,1047]" pageId="3" pageNumber="24">
<emphasis id="B949EAC3E52EFFBF599AA5ABFEE7FC7A" box="[227,329,927,951]" italics="true" pageId="3" pageNumber="24">Abdomen</emphasis>
. With six visible ventrites. Ratio of ventrite lengths: 22.0-5.0-4.0-5.0-7.0-7.0. Basal ventrite without femoral lines. Last ventrite widely rounded apically.
</paragraph>
</subSubSection>
<subSubSection id="C327655AE52EFFBF599AA20BFE6CFADA" pageId="3" pageNumber="24" type="etymology">
<paragraph id="8B8236D1E52EFFBF599AA20BFE6CFADA" blockId="3.[227,737,1087,1303]" pageId="3" pageNumber="24">
<emphasis id="B949EAC3E52EFFBF599AA20BFE67FB9A" bold="true" box="[227,457,1087,1111]" pageId="3" pageNumber="24">Derivatio nominis.</emphasis>
The specific epithet is derived from “primus inter pares” (feminine: prima inter pares), a Latin phrase meaning “first among equals”. The name is used as noun in apposition and refers to the habitual similarity to the recent congeners and Tertiary origin of the fossil specimen.
</paragraph>
</subSubSection>
</treatment>
</document>