389 lines
49 KiB
XML
389 lines
49 KiB
XML
<document id="3B5B6723328B8B9D988CE9C0B2E2E214" ID-CLB-Dataset="20492" ID-DOI="10.5169/seals-787049" ID-GBIF-Dataset="ce0756e2-5184-4b74-8472-ab69fed7951c" ID-ISSN="2624-9170" ID-Zenodo-Dep="6285683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935326602" checkinUser="carolina" docAuthor="Huber, Charles & Schmidt, Joachim" docDate="2017" docId="03948799FFE9FFF6FF41FC88FC68FBFE" docLanguage="en" docName="ContribNatHist.36.1-85.pdf.imf" docOrigin="Contributions to Natural History 36" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Nebria (Epinebriola) numburica Huber & Schmidt, 2017, sp. nov." docType="treatment" docVersion="12" lastPageNumber="54" masterDocId="FFADFFE1FFDBFFC0FF9CFFE9FFB0FFA7" masterDocTitle="Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="51" updateTime="1699470525451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="A73BD77A0EB68925B8543655117452B7">Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)</mods:title>
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<mods:namePart id="06F450D52B6DBF43C2739A964C0C7535">Huber, Charles</mods:namePart>
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<mods:namePart id="6A1CB614E2DE1CBD26E8DFF3D636ACF1">Schmidt, Joachim</mods:namePart>
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<mods:title id="5F3A768CC6AE8E36A2F3AD58C5E3F1AD">Contributions to Natural History</mods:title>
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<mods:date id="E30EA5FDB1394D703FD5A3C763DE6D9B">2017</mods:date>
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<emphasis id="B949EA9DFFE9FFF2FF41FC88FE99FCDD" bold="true" box="[221,297,865,890]" italics="true" pageId="50" pageNumber="51">Nebria</emphasis>
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(
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)
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<emphasis id="B949EA9DFFE9FFF2FE57FC88FDF3FCDD" bold="true" box="[459,579,865,890]" italics="true" pageId="50" pageNumber="51">numburica</emphasis>
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(
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)
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:
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,
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,
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, leg.
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(
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, same data as holotype (
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<collectingCountry id="F32A761FFFE9FFF2FF41FBF5FE94FB93" box="[221,292,1052,1076]" name="Nepal" pageId="50" pageNumber="51">Nepal</collectingCountry>
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<location id="8EE26054FFE9FFF2FE4AFBF5FDDEFB93" LSID="urn:lsid:plazi:treatment:03948799FFE9FFF6FF41FC88FC68FBFE:8EE26054FFE9FFF2FE4AFBF5FDDEFB93" box="[470,622,1052,1076]" country="Nepal" latitude="27.676666" longLatPrecision="20" longitude="86.59056" name="S Dudh Kund" pageId="50" pageNumber="51">S Dudh Kund</location>
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, leg.
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<collectingCountry id="F32A761FFFE9FFF2FC9DFBA8FCF8FBFE" box="[769,840,1089,1113]" name="Nepal" pageId="50" pageNumber="51">Nepal</collectingCountry>
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,
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,
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,
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|
||
<geoCoordinate id="EE095048FFE9FFF2FDE7FB8FFCBAFBD9" box="[635,778,1126,1150]" degrees="86" direction="east" minutes="35" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="26" value="86.59056">86°35'26" E</geoCoordinate>
|
||
,
|
||
<date id="FF83104FFFE9FFF2FC8AFB8FFC2AFBD9" box="[790,922,1126,1150]" pageId="50" pageNumber="51" value="2013-05-25">
|
||
<collectingDate id="EFC7E9A7FFE9FFF2FC8AFB8FFC2AFBD9" box="[790,922,1126,1150]" pageId="50" pageNumber="51" value="2013-05-25">25. V. 2013</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26C85359FFE9FFF2FC41FB8FFBE2FBD9" box="[989,1106,1126,1150]" pageId="50" pageNumber="51">J. Schmidt</collectorName>
|
||
(
|
||
<collectionCode id="ED2CAE4AFFE9FFF2FF78FB65FEF1FB03" box="[228,321,1164,1188]" pageId="50" pageNumber="51">cSCHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE9FFF5FE9FFB58FC80FDD3" lastPageId="53" lastPageNumber="54" pageId="50" pageNumber="51" type="description">
|
||
<paragraph id="8B82368FFFE9FFF2FE9FFB58FCD0FAB3" blockId="50.[221,1107,937,1598]" pageId="50" pageNumber="51">
|
||
Body length
|
||
<quantity id="4CC59B6AFFE9FFF2FE05FB58FDA3FB6E" box="[409,531,1201,1225]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="50" pageNumber="51" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
|
||
. Colour black, shiny, appendages of head brown. Antennae brown, rarely the antennal scape or the entire antenna blackish darkened. Legs black, knees brownish lightened, tarsi brown.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFE9FFF3FE9FFAC8FD31FEC9" blockId="50.[221,1107,937,1598]" lastBlockId="51.[184,1069,152,1636]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
|
||
Head with a shallow transverse impression behind the prominent eyes. Labrum with anterior margin straight, bearing six setae. Apical margin of clypeus straight. Clypeus laterally unisetose. Head supraorbitally unisetose, rarely asymmetrically bisetose or bisetose (in a single case in the
|
||
<typeStatus id="5486882DFFE9FFF2FC58FA78FC47FA0E" box="[964,1015,1425,1449]" pageId="50" pageNumber="51">type</typeStatus>
|
||
series). Frontal furrows shallow. Vertex finely transversally wrinkled but impunctate. Antennae long and slender extending to the middle of the elytra. Antennal scape elongate, as long as the eye's diameter, basally narrowed, cylindrical apically, with one dorsal seta. 2
|
||
<superScript id="7C489BC7FFE9FFF2FDD6F9CAFDD1F991" attach="left" box="[586,609,1571,1590]" fontSize="8" pageId="50" pageNumber="51">nd</superScript>
|
||
Antennomere with one seta ventroapically. 4
|
||
<superScript id="7C489BC7FFE8FFF3FF5AFF71FF69FF0C" attach="left" box="[198,217,152,171]" fontSize="8" pageId="51" pageNumber="52">th</superScript>
|
||
antennomere apically with a collar of long setae, without additional short setae. Penultimate labial palpomere bi- or trisetose, rarely asymmetrically with two and three setae. Ligula spiny, with two lateral setae. Maxillary stipes laterally straight, with 5–7 setae. Mentum with bifid medial tooth. Submentum with a row of 10–13 setae. Not or inconspicuously punctate on vertex. Microsculpture of the head isodiametric.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFE8FFF3FF41FE95FEB0FBD9" blockId="51.[184,1069,152,1636]" pageId="51" pageNumber="52">
|
||
Pronotum transverse, subcordate, moderately convex, narrowed basally, lateral margin straight in basal half, rarely with a faint concave sinuation in front of the basal angles (
|
||
<figureCitation id="13062A0AFFE8FFF3FE43FE2FFD8FFE79" box="[479,575,454,478]" captionStart="Fig" captionStartId="61.[184,213,1317,1335]" captionTargetBox="[185,1068,153,1276]" captionTargetId="figure-106@61.[551,733,151,716]" captionTargetPageId="61" captionText="Fig. 31: Left part of the pronotum of Epinebriola species. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) retingensis sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov.; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285816" httpUri="https://zenodo.org/record/6285816/files/figure.png" pageId="51" pageNumber="52">Fig. 31F</figureCitation>
|
||
). Lateral groove very narrow, conspicuous, broadened basally. Lateral margin basally blade-like. Posterior angles acutely projected posteriorly. Anterior angles wide, rounded, weakly protruding. Basal margin bisinuate, with a deep incision towards the basal fovea. Pronotal disc weakly convex. Basal fovea deepened, apical and posterior transverse impressions well-defined; median longitudinal impression weak. Basal fovea and transverse impressions sparsely punctate, lateral groove only in the basal, broadened part punctate. Punctation of the apical transverse impression extended to the apical margin. Posterior part of the median longitudinal impression with faint wrinkles. Apical margination of the pronotum restricted to lateral one-third, merges with a distinct ridge at the apex of the anterior angle; the ridge continues along the lateral groove to the hindmost lateral seta. Basal margination of the pronotum absent. Basolateral seta present. 3–5 (rarely two) midlateral setae in the apical half of the pronotum. Midlateral setae conspicuously inserted on the egde of the lateral margin, the large insertion grooves disturb the outline of the pronotum. Microsculpture of the pronotum isodiametric, impunctate on disc. Proepisternum sparsely and faintly punctate; prosternal process elongate, rounded apically, unmargined.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFE8FFF3FF41FB65FE34F9C3" blockId="51.[184,1069,152,1636]" pageId="51" pageNumber="52">
|
||
Elytral silhouette moderately convex, ovoid-elongate, narrowed basally, widest in the apical half of the elytra, no subapical sinuation. Elytral apex sharp, rounded. Basal margination straight, smoothly merged, or joined at an obtuse angle, with the lateral margination. Humeral carina faintly developed. Subapical carina absent. Striae on disc finely punctuate, striae only very weakly deepened, nearly inexistent. Striae and punctation obliterate towards the basal margin and towards the apex; striae 6–8 visible as a faint trace of punctation. Intervals flat; 3
|
||
<superScript id="7C489BC7FFE8FFF3FE71FA67FDB0FA06" attach="left" box="[493,512,1422,1441]" fontSize="8" pageId="51" pageNumber="52">rd</superScript>
|
||
interval with 5–8 long setae adjoining stria 3. 5
|
||
<superScript id="7C489BC7FFE8FFF3FB85FA67FB9CFA06" attach="none" box="[1049,1068,1422,1441]" fontSize="8" pageId="51" pageNumber="52">th</superScript>
|
||
interval often (59%) with 1–3 setae. 7
|
||
<superScript id="7C489BC7FFE8FFF3FDF0FA5AFDCFFA61" attach="left" box="[620,639,1459,1478]" fontSize="8" pageId="51" pageNumber="52">th</superScript>
|
||
interval rarely (14%) with 1–4 setae. Scutellar seta present. Microsculpture transverse-oblong. Mesepisterna smooth with a few strong punctulae. Metepisterna 1.6 times as long as wide; smooth. Hind coxa with 2–4 basal and one apical setae. Hindwings present as a short strap-like vestige.
|
||
</paragraph>
|
||
<caption id="DF426607FFEFFFF4FF41FF49FECAFF5B" ID-DOI="http://doi.org/10.5281/zenodo.6285794" ID-Zenodo-Dep="6285794" httpUri="https://zenodo.org/record/6285794/files/figure.png" pageId="52" pageNumber="53" startId="52.[221,249,160,178]" targetBox="[463,1106,153,1171]" targetPageId="52">
|
||
<paragraph id="8B82368FFFEFFFF4FF41FF49FECAFF5B" blockId="52.[221,429,160,252]" pageId="52" pageNumber="53">
|
||
Fig. 26:
|
||
<taxonomicName id="4C3D4D0CFFEFFFF4FEBDFF49FEFEFF6D" authority="(Epinebriola)" authorityName="LATREILLE" authorityYear="1802" baseAuthorityName="Epinebriola" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
Nebria
|
||
<emphasis id="B949EA9DFFEFFFF4FF41FF51FEFEFF6D" bold="true" box="[221,334,184,202]" pageId="52" pageNumber="53">
|
||
(
|
||
<emphasis id="B949EA9DFFEFFFF4FF78FF51FEF7FF6D" bold="true" box="[228,327,184,202]" italics="true" pageId="52" pageNumber="53">Epinebriola</emphasis>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EA9DFFEFFFF4FECEFF51FECAFF5B" bold="true" pageId="52" pageNumber="53">
|
||
<emphasis id="B949EA9DFFEFFFF4FECEFF51FE1DFF6D" bold="true" box="[338,429,184,202]" italics="true" pageId="52" pageNumber="53">numburica</emphasis>
|
||
sp. nov.
|
||
<typeStatus id="5486882DFFEFFFF4FEB8FF39FEC4FF45" box="[292,372,208,226]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
|
||
. Scale bar =
|
||
<quantity id="4CC59B6AFFEFFFF4FED9FF03FEC6FF5B" box="[325,374,234,252]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="52" pageNumber="53" unit="mm" value="5.0">5 mm</quantity>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B82368FFFEFFFF4FE9FFB3FFD15FA9E" blockId="52.[221,1107,1238,1636]" pageId="52" pageNumber="53">
|
||
Sternum 3 with a row of 4–7 medial setae. Sterna 4–6 each with 3–6 posterior paramedial setae; anal sternum with 1–3 paramedial setae in male,
|
||
<quantity id="4CC59B6AFFEFFFF4FC24FB15FC4CFAB3" box="[952,1020,1276,1300]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="52" pageNumber="53" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
|
||
female. All sterna with faint impressions laterally.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFEFFFF4FE9FFAAFFD85F9BE" blockId="52.[221,1107,1238,1636]" pageId="52" pageNumber="53">Legs long and slender. All tarsi dorsally glabrous, rarely with a few short setae. Protarsus of male with tarsomeres 1–3 distinctly broadened, with pads of adhesive setae ventrally. Apex of hind tarsomere 4 ventrally markedly projected. Meso- and metatarsomeres 2–4 ventroapically with long setae. All tarsomeres 5 ventrally with two rows of 3–4 short setae. Metatarsomere 5 longer than the metatarsomeres 3+4.</paragraph>
|
||
<paragraph id="8B82368FFFEFFFF5FE9FF9CFFDEDFF59" blockId="52.[221,1107,1238,1636]" lastBlockId="53.[183,1069,156,1598]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
|
||
Male genitalia: Edeagus (
|
||
<figureCitation id="13062A0AFFEFFFF4FDB3F9CFFD26F999" box="[559,662,1574,1598]" captionStart="Fig" captionStartId="38.[221,250,1317,1335]" captionTargetBox="[221,1106,153,1296]" captionTargetId="figure-70@38.[221,704,627,867]" captionTargetPageId="38" captionText="Fig. 20: Male edeagi of new Nebria species of the subgenus Epinebriola K. DANIEL, 1904. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) numburica sp. nov., with parameres; F: N. (E.) retingensis sp. nov.; G: N. (E.) incarinata sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov., apex in dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285764" httpUri="https://zenodo.org/record/6285764/files/figure.png" pageId="52" pageNumber="53">Fig. 20E</figureCitation>
|
||
): Basal part of the median lobe wide, with triangular basolateral lobes. Mid-shaft strongly curved at base on the inner side, and shallowly curved on the outer side forming a flat angle; moderately curved at apex. Apex long, slender, acute, faintly deflected to the left. Midshaft of the endophallus asetose.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFEEFFF5FF41FEE5FF42FE79" blockId="53.[183,1069,156,1598]" pageId="53" pageNumber="54">
|
||
Female genitalia: Gonocoxa (
|
||
<figureCitation id="13062A0AFFEEFFF5FDAEFEE5FD27FE83" box="[562,663,268,292]" captionStart="Fig" captionStartId="20.[221,250,1430,1448]" captionTargetBox="[228,1106,153,1365]" captionTargetId="figure-16@20.[574,757,751,1260]" captionTargetPageId="20" captionText="Fig. 10: Right gonocoxae of Epinebriola species, in ventral view. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) numburica sp. nov. in lateral view with the distale gonocoxite 2 dorsally grooved (arrow); H: N. (E.) christinae HUBER & SCHMIDT, 2007; I: N. (E.) pseudorestias sp. nov.; J: N. (E.) retingensis sp. nov.; Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285726" httpUri="https://zenodo.org/record/6285726/files/figure.png" pageId="53" pageNumber="54">Fig. 10F</figureCitation>
|
||
): Gonocoxite 2 triangular, half the length of gonocoxite 1, broad at base. Apex broadly rounded, slightly arcuate dorsally, dorsally slightly grooved (
|
||
<figureCitation id="13062A0AFFEEFFF5FDE1FEBFFD55FEC9" box="[637,741,342,366]" captionStart="Fig" captionStartId="20.[221,250,1430,1448]" captionTargetBox="[228,1106,153,1365]" captionTargetId="figure-16@20.[574,757,751,1260]" captionTargetPageId="20" captionText="Fig. 10: Right gonocoxae of Epinebriola species, in ventral view. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) numburica sp. nov. in lateral view with the distale gonocoxite 2 dorsally grooved (arrow); H: N. (E.) christinae HUBER & SCHMIDT, 2007; I: N. (E.) pseudorestias sp. nov.; J: N. (E.) retingensis sp. nov.; Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285726" httpUri="https://zenodo.org/record/6285726/files/figure.png" pageId="53" pageNumber="54">Fig. 10G</figureCitation>
|
||
). Ventral preapical insertion furrow elongate-oval, with two nematiforme setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFEEFFF5FF41FE05FC80FDD3" blockId="53.[183,1069,156,1598]" pageId="53" pageNumber="54">Body ratios: hea.w/fro.w = 1.46± 0.034 (1.39–1.53); prm.w/hea.w = 1.24± 0.023 (1.19–1.30); prm.w/prp.w = 1.53±0.047 (1.41–1.57); pra.w/prp.w = 1.16± 0.030 (1.09–1.20); ely.w/prm.w = 1.59±0.044 (1.51–1.69); prm.w/prm.l = 1.31± 0.038 (1.26–1.38); ely.l/ely.w = 1.62±0.027 (1.58–1.67).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFEEFFF5FF41FD68FEC5FD19" pageId="53" pageNumber="54" type="etymology">
|
||
<paragraph id="8B82368FFFEEFFF5FF41FD68FEC5FD19" blockId="53.[183,1069,156,1598]" pageId="53" pageNumber="54">
|
||
Etymology: The specific epithet refers to the Numbur Mountain North of the
|
||
<typeStatus id="5486882DFFEEFFF5FF7EFD4FFEA5FD19" box="[226,277,678,702]" pageId="53" pageNumber="54">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFEEFFF5FF41FD25FE69FA9E" pageId="53" pageNumber="54" type="diagnosis">
|
||
<paragraph id="8B82368FFFEEFFF5FF41FD25FC75FBD9" blockId="53.[183,1069,156,1598]" pageId="53" pageNumber="54">
|
||
Diagnosis: Due to the polysetose lateral margin of the pronotum and the polysetose 3
|
||
<superScript id="7C489BC7FFEEFFF5FEDBFD07FEEAFCA6" attach="left" box="[327,346,750,769]" fontSize="8" pageId="53" pageNumber="54">rd</superScript>
|
||
abdominal sternum
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FDA3FD18FD65FCAE" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[575,725,753,777]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="numburica" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFEEFFF5FDA3FD18FD65FCAE" box="[575,725,753,777]" italics="true" pageId="53" pageNumber="54">N. numburica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFEEFFF5FD47FD18FC82FCAE" box="[731,818,753,777]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FC42FD18FF6CFC89" authorityName="HUBER & SCHMIDT" authorityYear="2007" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FC42FD18FF6CFC89" italics="true" pageId="53" pageNumber="54">molendai</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<emphasis id="B949EA9DFFEEFFF5FE05FCFFFCC4FC89" box="[409,884,790,814]" italics="true" pageId="53" pageNumber="54">
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FE05FCFFFD98FC89" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[409,552,790,814]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">N. molendai</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FDAAFCFFFD7BFC89" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[566,715,790,814]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christinae">N. christinae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FD44FCFFFCC4FC89" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[728,884,790,814]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="numburica">N. numburica</taxonomicName>
|
||
</emphasis>
|
||
). The
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FC5DFCFFFB9CFC89" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[961,1068,790,814]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FC5DFCFFFB9CFC89" box="[961,1068,790,814]" italics="true" pageId="53" pageNumber="54">molendai</emphasis>
|
||
</taxonomicName>
|
||
species group is also characterized by the similar shape of gonocoxite 2, which have a broadly rounded apex in contrast to the similar gonocoxite 2 of
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FC56FC88FF4CFC39" authorityName="C.Huber & J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFEEFFF5FC56FC88FF4CFC39" italics="true" pageId="53" pageNumber="54">N. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFEEFFF5FE99FC6FFEEFFC39" box="[261,351,902,926]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with an acuminate apex. These three species of the
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FC5EFC6FFB9CFC39" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[962,1068,902,926]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FC5EFC6FFB9CFC39" box="[962,1068,902,926]" italics="true" pageId="53" pageNumber="54">molendai</emphasis>
|
||
</taxonomicName>
|
||
group inhabit the neighboring massifs of the Rolwaling Himal and the Solu Khumbu Massif.
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FEEAFC38FDBEFC4E" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[374,526,977,1001]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="numburica" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFEEFFF5FEEAFC38FDBEFC4E" box="[374,526,977,1001]" italics="true" pageId="53" pageNumber="54">N. numburica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFEEFFF5FD8AFC38FDDFFC4E" box="[534,623,977,1001]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FC9FFC38FC23FC4E" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[771,915,977,1001]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christinae">
|
||
<emphasis id="B949EA9DFFEEFFF5FC9FFC38FC23FC4E" box="[771,915,977,1001]" italics="true" pageId="53" pageNumber="54">N. christinae</emphasis>
|
||
</taxonomicName>
|
||
by the transverse pronotum, which is very slender in
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FD0EFC1FFC92FBA9" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[658,802,1014,1038]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christinae">
|
||
<emphasis id="B949EA9DFFEEFFF5FD0EFC1FFC92FBA9" box="[658,802,1014,1038]" italics="true" pageId="53" pageNumber="54">N. christinae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FCAEFC1FFC7AFBA9" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[818,970,1014,1038]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="numburica" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFEEFFF5FCAEFC1FFC7AFBA9" box="[818,970,1014,1038]" italics="true" pageId="53" pageNumber="54">N. numburica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFEEFFF5FC4FFC1FFB9DFBA9" box="[979,1069,1014,1038]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FEDFFBF5FE7CFB93" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[323,460,1052,1076]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FEDFFBF5FE7CFB93" box="[323,460,1052,1076]" italics="true" pageId="53" pageNumber="54">N. molendai</emphasis>
|
||
</taxonomicName>
|
||
by the sparsely punctate pronotum, which is coarsely and densely punctate in
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FE53FBA8FDE8FBFE" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[463,600,1089,1113]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FE53FBA8FDE8FBFE" box="[463,600,1089,1113]" italics="true" pageId="53" pageNumber="54">N. molendai</emphasis>
|
||
</taxonomicName>
|
||
, and by the long and slender apex of the median lobe, which is shorter in both
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FDC6FB8FFD52FBD9" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[602,738,1126,1150]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FDC6FB8FFD52FBD9" box="[602,738,1126,1150]" italics="true" pageId="53" pageNumber="54">N. molendai</emphasis>
|
||
</taxonomicName>
|
||
and in
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FCAEFB8FFC0FFBD9" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[818,959,1126,1150]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christinae">
|
||
<emphasis id="B949EA9DFFEEFFF5FCAEFB8FFC0FFBD9" box="[818,959,1126,1150]" italics="true" pageId="53" pageNumber="54">N. christinae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFEEFFF5FF41FB65FE69FA9E" blockId="53.[183,1069,156,1598]" pageId="53" pageNumber="54">
|
||
The
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FE8CFB65FEF2FB03" authorityName="ANDREWES" authorityYear="1936" box="[272,322,1164,1188]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rasa">
|
||
<emphasis id="B949EA9DFFEEFFF5FE8CFB65FEF2FB03" box="[272,322,1164,1188]" italics="true" pageId="53" pageNumber="54">rasa</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<emphasis id="B949EA9DFFEEFFF5FD9DFB65FC11FB03" box="[513,929,1164,1188]" italics="true" pageId="53" pageNumber="54">
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FD9DFB65FDE7FB03" authorityName="ANDREWES" authorityYear="1936" box="[513,599,1164,1188]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rasa">N. rasa</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FDF9FB65FCBCFB03" authorityName="SHILENKOV" authorityYear="1998" box="[613,780,1164,1188]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="schawalleri">N. schawalleri</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FC86FB65FC11FB03" authorityName="HUBER & SCHMIDT" authorityYear="2012" box="[794,929,1164,1188]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="martensi">N. martensi</taxonomicName>
|
||
</emphasis>
|
||
) of Eastern
|
||
<collectingCountry id="F32A761FFFEEFFF5FF24FB58FEB1FB6E" box="[184,257,1201,1225]" name="Nepal" pageId="53" pageNumber="54">Nepal</collectingCountry>
|
||
, which is characterized by the polysetose lateral margin of the pronotum as in the
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FEE6FB3FFE55FB49" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[378,485,1238,1262]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FEE6FB3FFE55FB49" box="[378,485,1238,1262]" italics="true" pageId="53" pageNumber="54">molendai</emphasis>
|
||
</taxonomicName>
|
||
species group, differs from the
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FCFEFB3FFC7DFB49" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[866,973,1238,1262]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FCFEFB3FFC7DFB49" box="[866,973,1238,1262]" italics="true" pageId="53" pageNumber="54">molendai</emphasis>
|
||
</taxonomicName>
|
||
species group by the absence of setae on the 3
|
||
<superScript id="7C489BC7FFEEFFF5FD1CFB11FD23FAAC" attach="left" box="[640,659,1272,1291]" fontSize="8" pageId="53" pageNumber="54">rd</superScript>
|
||
abdominal sternum (present in the
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FF24FAC8FE93FA9E" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[184,291,1313,1337]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FF24FAC8FE93FA9E" box="[184,291,1313,1337]" italics="true" pageId="53" pageNumber="54">molendai</emphasis>
|
||
</taxonomicName>
|
||
species group).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFEEFFF5FF41FAAFFDA2FA0E" pageId="53" pageNumber="54" type="distribution">
|
||
<paragraph id="8B82368FFFEEFFF5FF41FAAFFDA2FA0E" blockId="53.[183,1069,156,1598]" pageId="53" pageNumber="54">
|
||
Distribution (
|
||
<figureCitation id="13062A0AFFEEFFF5FEEEFAAFFE0FFAF9" box="[370,447,1350,1374]" captionStart="Fig" captionStartId="76.[221,250,160,178]" captionTargetId="figure-8@75.[184,1068,152,1645]" captionTargetPageId="75" captionText="Fig. 41: Locations with occurrences of Nebria (Epinebriola) species from the Central and Eastern Himalaya and from South Tibet." figureDoi="http://doi.org/10.5281/zenodo.6285892" httpUri="https://zenodo.org/record/6285892/files/figure.png" pageId="53" pageNumber="54">Fig. 41</figureCitation>
|
||
): Known only from the
|
||
<typeStatus id="5486882DFFEEFFF5FD57FAAFFD4EFAF9" box="[715,766,1350,1374]" pageId="53" pageNumber="54">type</typeStatus>
|
||
locality in the headwaters of the Dudhkund Khola on southern slope of the Shorong Himal, Solu Khumbu district, eastern Central
|
||
<collectingCountry id="F32A761FFFEEFFF5FE54FA78FDBEFA0E" box="[456,526,1425,1449]" name="Nepal" pageId="53" pageNumber="54">Nepal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFEEFFF5FF41FA5FFC2FFA53" pageId="53" pageNumber="54" type="biology_ecology">
|
||
<paragraph id="8B82368FFFEEFFF5FF41FA5FFC2FFA53" blockId="53.[183,1069,156,1598]" pageId="53" pageNumber="54">
|
||
Habitat (
|
||
<figureCitation id="13062A0AFFEEFFF5FEA6FA5FFE35FA69" box="[314,389,1462,1486]" captionStart="Fig" captionStartId="54.[221,250,832,850]" captionTargetBox="[221,1105,153,808]" captionTargetId="figure-185@54.[221,1105,153,812]" captionTargetPageId="54" captionText="Fig. 27: Habitat of Nebria (Epinebriola) numburica sp. nov.: Headwater of the Dudhkund Khola, 4100 m a.s.l., subalpine zone. Specimens were collected alongside the stream. Photo: Joachim Schmidt." figureDoi="http://doi.org/10.5281/zenodo.6285800" httpUri="https://zenodo.org/record/6285800/files/figure.png" pageId="53" pageNumber="54">Fig. 27</figureCitation>
|
||
): The specimens were collected alongside melt water streams and at the edge of a rivulet in the lower alpine and subalpine zones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFEEFFF6FF41F9E8FC68FBFE" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="discussion">
|
||
<paragraph id="8B82368FFFEEFFF6FF41F9E8FC68FBFE" blockId="53.[183,1069,156,1598]" lastBlockId="54.[221,1106,977,1113]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
|
||
Monophyly. The monophyly of the
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FDEDF9E8FD6CF9BE" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[625,732,1537,1561]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEEFFF5FDEDF9E8FD6CF9BE" box="[625,732,1537,1561]" italics="true" pageId="53" pageNumber="54">molendai</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<emphasis id="B949EA9DFFEEFFF5FC04F9E8FEF8F999" italics="true" pageId="53" pageNumber="54">
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FC04F9E8FB98F9BE" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[920,1064,1537,1561]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="molendai">N. molendai</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FF24F9CFFEF8F999" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[184,328,1574,1598]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christinae">N. christinae</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3D4D0CFFEEFFF5FE1FF9CFFDACF999" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[387,540,1574,1598]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="numburica" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFEEFFF5FE1FF9CFFDACF999" box="[387,540,1574,1598]" italics="true" pageId="53" pageNumber="54">N. numburica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFEEFFF5FDB9F9CFFDCEF999" box="[549,638,1574,1598]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) appears clearly and is suggested by four characters: the polysetose lateral margin of the pronotum, the absence of setae on the 3
|
||
<superScript id="7C489BC7FFEDFFF6FE07FC1AFE1EFBA1" attach="left" box="[411,430,1011,1030]" fontSize="8" pageId="54" pageNumber="55">rd</superScript>
|
||
abdominal sternum, and the resemblances in the shapes of the median lobes and of the gonocoxites 2. Moreover, the separated distribution of the
|
||
<taxonomicName id="4C3D4D0CFFEDFFF6FEB8FBA8FE3EFBFE" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[292,398,1089,1113]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFEDFFF6FEB8FBA8FE3EFBFE" box="[292,398,1089,1113]" italics="true" pageId="54" pageNumber="55">molendai</emphasis>
|
||
</taxonomicName>
|
||
species group is also consistent with its monophyly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |