434 lines
46 KiB
XML
434 lines
46 KiB
XML
<document ID-DOI="10.11646/zootaxa.5231.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7571485" ID-ZooBank="9C66A388-C221-4D17-B2AA-F9EB7F1EC940" checkinTime="1674711822273" checkinUser="plazi" docAuthor="Kraus, Fred" docDate="2023" docId="0381A239FFF9FFDF7E8DFF75FECFFCC2" docLanguage="en" docName="zootaxa.5231.1.1.pdf" docOrigin="Zootaxa 5231 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gerrhopilus flavinotatus Kraus 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="FFB8DA41FFF4FFD17E1AFFE2FFBAFFD7" masterDocTitle="A revision of Gerrhopilus inornatus (Squamata: Gerrhopilidae) reveals a multi-species complex" masterLastPageNumber="23" masterPageNumber="1" pageNumber="14" updateTime="1674753466069" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of Gerrhopilus inornatus (Squamata: Gerrhopilidae) reveals a multi-species complex</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kraus, Fred</mods:namePart>
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<mods:affiliation>Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, U. S. A</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5231</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:start>1</mods:start>
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<mods:end>23</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5231.1.1</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5231.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7571485</mods:identifier>
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<mods:identifier type="ZooBank">9C66A388-C221-4D17-B2AA-F9EB7F1EC940</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0381A239FFF9FFDF7E8DFF75FECFFCC2" httpUri="http://treatment.plazi.org/id/0381A239FFF9FFDF7E8DFF75FECFFCC2" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<subSubSection box="[151,550,151,178]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph blockId="13.[151,550,151,214]" box="[151,550,151,178]" pageId="13" pageNumber="14">
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<heading bold="true" box="[151,550,151,178]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis bold="true" box="[151,550,151,178]" pageId="13" pageNumber="14">
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<taxonomicName authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[151,448,151,178]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="flavinotatus" status="sp. nov.">
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<emphasis bold="true" box="[151,448,151,178]" italics="true" pageId="13" pageNumber="14">Gerrhopilus flavinotatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[456,550,151,177]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,280,187,214]" pageId="13" pageNumber="14" type="description">
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<paragraph blockId="13.[151,550,151,214]" box="[151,280,187,214]" pageId="13" pageNumber="14">
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<figureCitation box="[151,280,187,214]" captionStart="FIGURE 5" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[188,1398,197,1908]" captionTargetId="figure-27@16.[174,1413,181,1920]" captionTargetPageId="16" captionText="FIGURE 5. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 22 midbody scale rows and venters marked with yellow. A, B holotype of Gerrhopilus wallachi (MCZ 140724); C, D holotype of Gerrhopilus flavinotatus (MCZ 175100); E, F holotype of Gerrhopilus lorealis (BPBM 2272)." figureDoi="http://doi.org/10.5281/zenodo.7571501" httpUri="https://zenodo.org/record/7571501/files/figure.png" pageId="13" pageNumber="14">Figs. 5C, D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="reference_group">
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<paragraph blockId="13.[151,662,255,311]" box="[151,548,255,280]" pageId="13" pageNumber="14">
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<treatmentCitationGroup box="[151,548,255,280]" pageId="13" pageNumber="14">
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<treatmentCitation author="Wallach, V." box="[151,544,255,280]" page="113" pageId="13" pageNumber="14" year="1996">
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<taxonomicName authority="Wallach 1996: 113" authorityName="Wallach" authorityPageNumber="113" authorityYear="1996" box="[151,544,255,280]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
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<emphasis box="[151,346,256,280]" italics="true" pageId="13" pageNumber="14">Typhlops inornatus</emphasis>
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<bibRefCitation author="Wallach, V." box="[352,544,255,280]" pageId="13" pageNumber="14" pagination="107 - 118" refId="ref19774" refString="Wallach, V. (1996) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996">Wallach 1996: 113</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="13.[151,662,255,311]" box="[151,662,286,311]" pageId="13" pageNumber="14">
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<treatmentCitationGroup box="[151,662,286,311]" pageId="13" pageNumber="14">
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<treatmentCitation author="Pyron, R. A. & Wallach, V." box="[151,658,286,311]" page="79" pageId="13" pageNumber="14" year="2014">
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<taxonomicName authority="Pyron & Wallach 2014: 79" authorityName="Pyron & Wallach" authorityPageNumber="79" authorityYear="2014" box="[151,658,286,311]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
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<emphasis box="[151,378,286,310]" italics="true" pageId="13" pageNumber="14">Gerrhopilus inornatus</emphasis>
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<bibRefCitation author="Pyron, R. A. & Wallach, V." box="[384,658,286,311]" pageId="13" pageNumber="14" pagination="1 - 81" refId="ref19325" refString="Pyron, R. A. & Wallach, V. (2014) Systematics of the blind snakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa, 3829 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 3829.1.1" type="journal article" year="2014">Pyron & Wallach 2014: 79</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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</treatmentCitationGroup>
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||
</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="materials_examined">
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<paragraph blockId="13.[151,1437,356,2039]" pageId="13" pageNumber="14">
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<materialsCitation collectingDate="1969-03-29" collectionCode="MCZ" country="Papua New Guinea" county="Western Province" elevation="1830" latitude="-5.88" location="Fred Parker" longLatPrecision="783" longitude="141.23" municipality="Menemsore" pageId="13" pageNumber="14" specimenCode="MCZ 175100" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[151,267,357,383]" pageId="13" pageNumber="14">
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<typeStatus box="[151,260,357,383]" pageId="13" pageNumber="14">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode box="[276,429,356,383]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14" type="Museum">MCZ 175100</specimenCode>
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(field tag Y-31495), collected on
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<date box="[817,992,356,383]" pageId="13" pageNumber="14" value="1969-03-29">
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<collectingDate box="[817,992,356,383]" pageId="13" pageNumber="14" value="1969-03-29">29 March 1969</collectingDate>
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</date>
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by
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<location LSID="urn:lsid:plazi:treatment:0381A239FFF9FFDF7E8DFF75FECFFCC2:8EF745F4FFF9FFDC7A17FE86FB28FEA9" box="[1037,1170,356,382]" country="Papua New Guinea" county="Western Province" latitude="-5.88" longLatPrecision="783" longitude="141.23" municipality="Menemsore" name="Fred Parker" pageId="13" pageNumber="14">Fred Parker</location>
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at
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<collectingMunicipality box="[1208,1343,357,383]" pageId="13" pageNumber="14">Menemsore</collectingMunicipality>
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,
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<geoCoordinate box="[1356,1431,356,383]" degrees="5.88" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="555" value="-5.88">5.88 S</geoCoordinate>
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,
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<geoCoordinate box="[151,254,393,419]" degrees="141.23" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="555" value="141.23">141.23 E</geoCoordinate>
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,
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<quantity box="[266,351,393,419]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="13" pageNumber="14" unit="m" value="1830.0">1830 m</quantity>
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a.s.l.
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</elevation>
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,
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<collectingCounty box="[424,624,392,419]" pageId="13" pageNumber="14">Western Province</collectingCounty>
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,
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<collectingCountry box="[635,856,392,419]" name="Papua New Guinea" pageId="13" pageNumber="14">Papua New Guinea</collectingCountry>
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.
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||
</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="diagnosis">
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||
<paragraph blockId="13.[151,1437,356,2039]" pageId="13" pageNumber="14">
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||
<emphasis bold="true" box="[199,320,428,454]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
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A
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<taxonomicName box="[353,575,428,454]" pageId="13" pageNumber="14">
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||
<emphasis box="[353,488,428,454]" italics="true" pageId="13" pageNumber="14">Gerrhopilus</emphasis>
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species
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</taxonomicName>
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||
characterized by the unique combination of having a rounded snout in lateral view; LSR = 22 at midbody; TSR = 327; loreal absent; supralabial imbrication pattern T-V; subocular scale one; presubocular scale absent; a sharp, protruding spine on the tail tip that is white; 3 glands in the prefrontal,
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<quantity box="[1326,1396,500,526]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="13" pageNumber="14" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">6–8 in</quantity>
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||
the supraocular, 0 in the ocular and frontal,
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<quantity box="[606,706,536,563]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" metricValueMax="7.112" metricValueMin="6.096" pageId="13" pageNumber="14" unit="in" value="26.0" valueMax="28.0" valueMin="24.0">24–28 in</quantity>
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the preocular, and
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<quantity box="[924,997,536,563]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="10.16" metricValueMin="0.0" pageId="13" pageNumber="14" unit="in" value="2.0" valueMax="4.0" valueMin="0.0">0–4 in</quantity>
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||
the subocular; L/W = 26.6; TL/SVL = 0.037; and a dark-brown venter with 2–4 rows of yellow scales mid-ventrally on the posterior half of the body with brown scales scattered in this yellow field. Its size is small for this species group (
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||
<tableCitation box="[1059,1142,608,635]" captionStart="TABLE 1" captionStartId="17.[151,244,552,578]" captionTargetPageId="17" captionText="TABLE 1. Mensural and count data for members of the Gerrhopilus inornatus Group. Forward slashes separate counts on the right and left sides, respectively." pageId="13" pageNumber="14">Table 1</tableCitation>
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||
).
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||
</paragraph>
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||
<paragraph blockId="13.[151,1437,356,2039]" pageId="13" pageNumber="14">
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||
<emphasis bold="true" box="[199,584,644,671]" pageId="13" pageNumber="14">Comparisons with other species.</emphasis>
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||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[592,866,644,670]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="flavinotatus">
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||
<emphasis box="[592,866,644,670]" italics="true" pageId="13" pageNumber="14">Gerrhopilus flavinotatus</emphasis>
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||
</taxonomicName>
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is distinguished from
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<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[1119,1279,644,671]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="fredparkeri">
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<emphasis box="[1119,1279,644,671]" italics="true" pageId="13" pageNumber="14">G. fredparkeri</emphasis>
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</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[1294,1395,644,671]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hades">
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<emphasis box="[1294,1395,644,671]" italics="true" pageId="13" pageNumber="14">G. hades</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
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<emphasis italics="true" pageId="13" pageNumber="14">G. inornatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Kraus" authorityYear="2023" box="[271,428,680,707]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="slapcinskyi">
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<emphasis box="[271,428,680,707]" italics="true" pageId="13" pageNumber="14">G. slapcinskyi</emphasis>
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</taxonomicName>
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, and
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<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[489,619,680,707]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="suturalis">
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<emphasis box="[489,619,680,707]" italics="true" pageId="13" pageNumber="14">G. suturalis</emphasis>
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</taxonomicName>
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in having 22 longitudinal scale rows at midbody (vs
|
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<quantity box="[1207,1264,680,706]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="13" pageNumber="14" unit="in" value="16.0">16 in</quantity>
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||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[1270,1429,680,707]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="fredparkeri">
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||
<emphasis box="[1270,1429,680,707]" italics="true" pageId="13" pageNumber="14">G. fredparkeri</emphasis>
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||
</taxonomicName>
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||
,
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||
<quantity box="[151,211,716,743]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="13" pageNumber="14" unit="in" value="18.0">18 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[221,323,716,743]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hades">
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||
<emphasis box="[221,323,716,743]" italics="true" pageId="13" pageNumber="14">G. hades</emphasis>
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||
</taxonomicName>
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||
and
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||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[383,518,716,743]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="suturalis">
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||
<emphasis box="[383,518,716,743]" italics="true" pageId="13" pageNumber="14">G. suturalis</emphasis>
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||
</taxonomicName>
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||
, and
|
||
<quantity box="[585,645,716,743]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="14" unit="in" value="20.0">20 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[655,798,717,743]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
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||
<emphasis box="[655,798,717,743]" italics="true" pageId="13" pageNumber="14">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[859,1019,716,743]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[859,1019,716,743]" italics="true" pageId="13" pageNumber="14">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
). It is further distinguished from
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="fredparkeri">
|
||
<emphasis italics="true" pageId="13" pageNumber="14">G. fredparkeri</emphasis>
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||
</taxonomicName>
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||
in having fewer transverse scale rows (327 vs
|
||
<quantity box="[790,860,752,779]" metricMagnitude="1" metricUnit="m" metricValue="1.36906" pageId="13" pageNumber="14" unit="in" value="539.0">539 in</quantity>
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[865,1024,752,779]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="fredparkeri">
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||
<emphasis box="[865,1024,752,779]" italics="true" pageId="13" pageNumber="14">G. fredparkeri</emphasis>
|
||
</taxonomicName>
|
||
), as it is from
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||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1182,1357,753,779]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="papuanorum">
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||
<emphasis box="[1182,1357,753,779]" italics="true" pageId="13" pageNumber="14">G. papuanorum</emphasis>
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||
</taxonomicName>
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||
and
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||
<taxonomicName authorityName="Kraus" authorityYear="2023" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
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<emphasis italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
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||
</taxonomicName>
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||
(407–414 rows in
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<taxonomicName authorityName="Kraus" authorityYear="2023" box="[452,627,789,815]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[452,627,789,815]" italics="true" pageId="13" pageNumber="14">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
and 430 rows in
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[813,941,788,815]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
|
||
<emphasis box="[813,941,788,815]" italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
|
||
</taxonomicName>
|
||
) and apparently from
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[1185,1324,789,815]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[1185,1324,789,815]" italics="true" pageId="13" pageNumber="14">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
(374–375 rows in
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[242,383,825,851]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[242,383,825,851]" italics="true" pageId="13" pageNumber="14">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
). It is further distinguished from
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[758,898,825,851]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[758,898,825,851]" italics="true" pageId="13" pageNumber="14">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
in being more robust (L/W = 26.6 vs
|
||
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.06934" metricValueMax="1.2141199999999999" metricValueMin="0.9245599999999999" pageId="13" pageNumber="14" unit="in" value="42.099999999999994" valueMax="47.8" valueMin="36.4">36.4–47.8 in</quantity>
|
||
<taxonomicName authorityName="Pyron & Wallach" authorityYear="2014" box="[180,321,861,887]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[180,321,861,887]" italics="true" pageId="13" pageNumber="14">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[406,564,860,887]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[406,564,860,887]" italics="true" pageId="13" pageNumber="14">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
in lacking glands in the ocular (vs
|
||
<quantity box="[958,1029,860,887]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.286" metricValueMin="1.016" pageId="13" pageNumber="14" unit="in" value="6.5" valueMax="9.0" valueMin="4.0">4–9 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1035,1193,860,887]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[1035,1193,860,887]" italics="true" pageId="13" pageNumber="14">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
), having more glands in the supraocular (6–8 vs
|
||
<quantity box="[448,519,896,923]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="7.62" metricValueMin="0.0" pageId="13" pageNumber="14" unit="in" value="1.5" valueMax="3.0" valueMin="0.0">0–3 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[526,683,896,923]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[526,683,896,923]" italics="true" pageId="13" pageNumber="14">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
), fewer glands in the preocular (24–28 vs
|
||
<quantity box="[1157,1256,896,923]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="9.906" metricValueMin="8.89" pageId="13" pageNumber="14" unit="in" value="37.0" valueMax="39.0" valueMin="35.0">35–39 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1263,1420,896,923]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[1263,1420,896,923]" italics="true" pageId="13" pageNumber="14">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
), and a white tail spine (vs black in
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[560,721,932,959]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[560,721,932,959]" italics="true" pageId="13" pageNumber="14">G. slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[813,993,933,959]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[813,993,933,959]" italics="true" pageId="13" pageNumber="14">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
in being more robust (L/W = 26.6 vs
|
||
<quantity box="[151,290,968,995]" metricMagnitude="0" metricUnit="m" metricValue="1.3296899999999998" metricValueMax="1.3589000000000002" metricValueMin="1.3004799999999999" pageId="13" pageNumber="14" unit="in" value="52.35" valueMax="53.5" valueMin="51.2">51.2–53.5 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[296,471,969,995]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[296,471,969,995]" italics="true" pageId="13" pageNumber="14">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
) and having a longer tail (TL/SVL = 0.037 vs
|
||
<quantity box="[979,1146,968,994]" metricMagnitude="-4" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.604" metricValueMin="5.334" pageId="13" pageNumber="14" unit="in" value="0.0235" valueMax="0.026" valueMin="0.021">
|
||
<geoCoordinate box="[979,1040,968,994]" degrees="0.021" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="0.021">0.021</geoCoordinate>
|
||
<geoCoordinate box="[1040,1118,968,994]" degrees="0.026" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="55" value="-0.026">–0.026</geoCoordinate>
|
||
in
|
||
</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1152,1327,969,995]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[1152,1327,969,995]" italics="true" pageId="13" pageNumber="14">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis italics="true" pageId="13" pageNumber="14">G. hades</emphasis>
|
||
</taxonomicName>
|
||
in having 22 longitudinal scale rows behind the head (vs
|
||
<quantity box="[856,912,1004,1031]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="13" pageNumber="14" unit="in" value="18.0">18 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[919,1018,1004,1031]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[919,1018,1004,1031]" italics="true" pageId="13" pageNumber="14">G. hades</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[1102,1233,1004,1031]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[1102,1233,1004,1031]" italics="true" pageId="13" pageNumber="14">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
in having a single postocular on each side of the head (vs two in
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[673,804,1040,1067]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[673,804,1040,1067]" italics="true" pageId="13" pageNumber="14">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
) and lacking a presubocular (vs present in
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[1288,1419,1040,1067]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[1288,1419,1040,1067]" italics="true" pageId="13" pageNumber="14">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[256,384,1076,1103]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
|
||
<emphasis box="[256,384,1076,1103]" italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
|
||
</taxonomicName>
|
||
in having a sharp, protruding tail spine (vs short, blunt, and downturned in
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1205,1332,1076,1103]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
|
||
<emphasis box="[1205,1332,1076,1103]" italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
|
||
</taxonomicName>
|
||
), lacking glands in the ocular (vs
|
||
<quantity box="[421,505,1112,1139]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="13" pageNumber="14" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9–11 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[512,642,1112,1139]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
|
||
<emphasis box="[512,642,1112,1139]" italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
|
||
</taxonomicName>
|
||
), and having fewer glands in the prefrontal (3 vs
|
||
<quantity box="[1198,1242,1112,1139]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="13" pageNumber="14" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1249,1379,1112,1139]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
|
||
<emphasis box="[1249,1379,1112,1139]" italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
|
||
</taxonomicName>
|
||
) and preocular (24–28 vs
|
||
<quantity box="[380,478,1148,1175]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" metricValueMax="8.89" metricValueMin="8.382" pageId="13" pageNumber="14" unit="in" value="34.0" valueMax="35.0" valueMin="33.0">33–35 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[485,614,1148,1175]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
|
||
<emphasis box="[485,614,1148,1175]" italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[638,911,1148,1174]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="flavinotatus">
|
||
<emphasis box="[638,911,1148,1174]" italics="true" pageId="13" pageNumber="14">Gerrhopilus flavinotatus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1159,1319,1148,1175]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="polyadenus">
|
||
<emphasis box="[1159,1319,1148,1175]" italics="true" pageId="13" pageNumber="14">G. polyadenus</emphasis>
|
||
</taxonomicName>
|
||
in lacking glands in the ocular (vs
|
||
<quantity box="[434,535,1184,1211]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="5.842" metricValueMin="2.794" pageId="13" pageNumber="14" unit="in" value="17.0" valueMax="23.0" valueMin="11.0">11–23 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[545,708,1184,1211]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="polyadenus">
|
||
<emphasis box="[545,708,1184,1211]" italics="true" pageId="13" pageNumber="14">G. polyadenus</emphasis>
|
||
</taxonomicName>
|
||
) and having fewer glands in the subocular (0–4 vs
|
||
<quantity box="[1311,1399,1184,1211]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="4.064" metricValueMin="1.27" pageId="13" pageNumber="14" unit="in" value="10.5" valueMax="16.0" valueMin="5.0">5–16 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="polyadenus">
|
||
<emphasis italics="true" pageId="13" pageNumber="14">G. polyadenus</emphasis>
|
||
</taxonomicName>
|
||
) and preocular (24–28 vs
|
||
<quantity box="[563,660,1220,1247]" metricMagnitude="0" metricUnit="m" metricValue="1.1684" metricValueMax="1.27" metricValueMin="1.0668" pageId="13" pageNumber="14" unit="in" value="46.0" valueMax="50.0" valueMin="42.0">42–50 in</quantity>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[665,824,1220,1247]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="polyadenus">
|
||
<emphasis box="[665,824,1220,1247]" italics="true" pageId="13" pageNumber="14">G. polyadenus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[846,1118,1220,1246]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="flavinotatus">
|
||
<emphasis box="[846,1118,1220,1246]" italics="true" pageId="13" pageNumber="14">Gerrhopilus flavinotatus</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from all other Melanesian
|
||
<taxonomicName authorityName="Fitzinger" authorityYear="1843" box="[384,519,1256,1282]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[384,519,1256,1282]" italics="true" pageId="13" pageNumber="14">Gerrhopilus</emphasis>
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[604,733,1256,1283]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="wallachi">
|
||
<emphasis box="[604,733,1256,1283]" italics="true" pageId="13" pageNumber="14">G. wallachi</emphasis>
|
||
</taxonomicName>
|
||
in having a venter that is brown with yellow mid-ventral scales throughout the posterior third of the body (vs venter uniformly brown in all the other species).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
|
||
<paragraph blockId="13.[151,1437,356,2039]" lastBlockId="14.[151,1437,151,789]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<emphasis bold="true" box="[199,529,1328,1355]" pageId="13" pageNumber="14">
|
||
Description of the
|
||
<typeStatus box="[420,523,1329,1355]" pageId="13" pageNumber="14">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Immature female. L =
|
||
<quantity box="[789,905,1328,1355]" metricMagnitude="-1" metricUnit="m" metricValue="1.545" pageId="13" pageNumber="14" unit="mm" value="154.5">154.5 mm</quantity>
|
||
, SVL =
|
||
<quantity box="[998,1093,1329,1355]" metricMagnitude="-1" metricUnit="m" metricValue="1.49" pageId="13" pageNumber="14" unit="mm" value="149.0">149 mm</quantity>
|
||
, TL =
|
||
<quantity box="[1168,1256,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="13" pageNumber="14" unit="mm" value="5.5">5.5 mm</quantity>
|
||
, HW =
|
||
<quantity box="[1344,1432,1329,1355]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="14" unit="mm" value="4.1">4.1 mm</quantity>
|
||
, SN =
|
||
<quantity box="[216,303,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="14" unit="mm" value="1.6">1.6 mm</quantity>
|
||
, SW =
|
||
<quantity box="[385,472,1365,1391]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="14" unit="mm" value="3.3">3.3 mm</quantity>
|
||
, PSN =
|
||
<quantity box="[564,651,1365,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="14" unit="mm" value="1.1">1.1 mm</quantity>
|
||
, RW =
|
||
<quantity box="[734,822,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="14" unit="mm" value="1.6">1.6 mm</quantity>
|
||
, EW =
|
||
<quantity box="[905,993,1365,1391]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="mm" value="0.3">0.3 mm</quantity>
|
||
, W =
|
||
<quantity box="[1058,1146,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="14" unit="mm" value="5.8">5.8 mm</quantity>
|
||
, VW =
|
||
<quantity box="[1231,1319,1365,1391]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="13" pageNumber="14" unit="mm" value="4.3">4.3 mm</quantity>
|
||
, TW =
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="14" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, L/W = 26.6, TL/SVL = 0.037. Head slightly wider than neck. Snout rounded in dorsal and lateral views. Snout anterior to lower jaw horizontal, parallel to body axis. Rostral rather narrow (RW/HW = 0.39), oval dorsally, lateral margins convex, posterior border extending approximately two-thirds way between naris and eye, posterior margin convex; ventrally surface papillose, with sides and posterior margin straight. Nasals separated dorsally by prefrontal (
|
||
<figureCitation box="[159,246,1544,1571]" captionStart="FIGURE 5" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[188,1398,197,1908]" captionTargetId="figure-27@16.[174,1413,181,1920]" captionTargetPageId="16" captionText="FIGURE 5. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 22 midbody scale rows and venters marked with yellow. A, B holotype of Gerrhopilus wallachi (MCZ 140724); C, D holotype of Gerrhopilus flavinotatus (MCZ 175100); E, F holotype of Gerrhopilus lorealis (BPBM 2272)." figureDoi="http://doi.org/10.5281/zenodo.7571501" httpUri="https://zenodo.org/record/7571501/files/figure.png" pageId="13" pageNumber="14">Fig. 5C</figureCitation>
|
||
); superior nasal large, with slightly sinuous posterior margin, concave dorsally, convex ventrally (
|
||
<figureCitation box="[1334,1423,1544,1571]" captionStart="FIGURE 5" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[188,1398,197,1908]" captionTargetId="figure-27@16.[174,1413,181,1920]" captionTargetPageId="16" captionText="FIGURE 5. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 22 midbody scale rows and venters marked with yellow. A, B holotype of Gerrhopilus wallachi (MCZ 140724); C, D holotype of Gerrhopilus flavinotatus (MCZ 175100); E, F holotype of Gerrhopilus lorealis (BPBM 2272)." figureDoi="http://doi.org/10.5281/zenodo.7571501" httpUri="https://zenodo.org/record/7571501/files/figure.png" pageId="13" pageNumber="14">Fig. 5D</figureCitation>
|
||
). External naris semicircular, close to rostral, anterior half covered by inferior nasal; superior nasal suture extending anterodorsally from naris to rostral; inferior nasal suture complete, contacting second supralabial posterior to latter’s contact with first supralabial. Prefrontal subequal to supraoculars. Supraoculars wider than long, longest at medial third, tapering toward eye, larger than parietals and interparietal, which are subequal in size. Frontal divided into two scales, left overlapping right, subequal in size to parietals. Preocular large, triangular; somewhat larger than ocular but smaller than superior nasal. Ocular large, smaller than preocular, extending dorsally well above preocular and to center of suproculars, extending ventrally to ~2/3 depth of preocular, bordered posteroventrally by subocular of approximately half its size. Eye fairly obscure, with pale pupil, situated at widest point of ocular and approximately midway along its height, anterior half covered by preocular plate in lateral view. Four postoculars bordering ocular and subocular between parietal and fourth supralabial. Four supralabials, third the largest, all except first with long axis oblique to long axis of body, first approximately square. Supralabial imbrication pattern T-V, posterior border of second supralabial overlaps anteroventral margin of preocular, that of third supralabial overlaps anteroventral margin of subocular and extends posterior to rear margin of preocular. Mental hexagonal, wider than long, projecting slightly beyond curve of lower jaw and fitting into notch on upper lip when mouth is closed. Infralabials two on each side, second much longer.
|
||
</paragraph>
|
||
<paragraph blockId="14.[151,1437,151,789]" pageId="14" pageNumber="15">
|
||
Longitudinal scale rows uniformly 22 behind head, at midbody, and anterior to vent; transverse scale rows 327,
|
||
<specimenCount box="[208,368,259,285]" pageId="14" pageNumber="15" type="intercalary">six intercalary</specimenCount>
|
||
scales along vertebral row; subcaudals 12; dorsocaudals 12; apical region with sharp, laterally compressed, corneous spine that extends well past last scales.
|
||
</paragraph>
|
||
<paragraph blockId="14.[151,1437,151,789]" pageId="14" pageNumber="15">Rostral, nasals, and preoculars densely covered in pale glands; oculars with 0 glands, preoculars with 24 (R) and 28 (L), supraoculars with 6 (R) and 5 (L), prefrontal with 3; frontal with 0, and suboculars with 0 (R) and 4 (L).</paragraph>
|
||
<paragraph blockId="14.[151,1437,151,789]" pageId="14" pageNumber="15">In preservative, 53 years after preservation, dorsum and sides uniformly brown, each scale darker anteriorly. Venter brown, with a few pale straw-yellow scales scattered mid-ventrally posterior to midbody, with posterior third of venter comprised of 2–4 mid-ventral rows of pale straw-yellow scales with brown scales scattered among them, the yellow scales becoming more continuous and predominant toward the tail. Subcaudals and tail spine pale straw yellow. Chin, mid-ventral throat, infralabials, first three supralabials, and anterior half of rostral pale straw yellow. Iris black; pupil pale gray.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph blockId="14.[151,1437,151,789]" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[199,334,619,645]" pageId="14" pageNumber="15">Etymology.</emphasis>
|
||
The specific epithet is a masculine compound adjective from the Latin
|
||
<emphasis box="[1127,1194,619,645]" italics="true" pageId="14" pageNumber="15">flavus</emphasis>
|
||
, meaning
|
||
<emphasis box="[1309,1382,619,645]" italics="true" pageId="14" pageNumber="15">yellow</emphasis>
|
||
, and
|
||
<emphasis box="[151,233,655,680]" italics="true" pageId="14" pageNumber="15">notatus</emphasis>
|
||
, meaning
|
||
<emphasis box="[351,409,655,681]" italics="true" pageId="14" pageNumber="15">mark</emphasis>
|
||
, in reference to the distinctive ventral color pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,845,690,717]" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph blockId="14.[151,1437,151,789]" box="[199,845,690,717]" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[199,353,690,716]" pageId="14" pageNumber="15">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus box="[608,656,691,717]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality (
|
||
<figureCitation box="[763,832,691,717]" captionStart="FIGURE 2" captionStartId="5.[152,255,1069,1093]" captionTargetBox="[181,1408,456,1045]" captionTargetId="figure-148@5.[179,1408,453,1045]" captionTargetPageId="5" captionText="FIGURE 2. Map of southern Papua New Guinea showing collecting localities for Gerrhopilus inornatus (star), G. flavinotatus (diamond), G. lorealis (hexagon), G. papuanorum (square), G. polyadenus (open circles), G. slapcinskyi (filled circle), and G. wallachi (triangle)." figureDoi="http://doi.org/10.5281/zenodo.7571491" httpUri="https://zenodo.org/record/7571491/files/figure.png" pageId="14" pageNumber="15">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph blockId="14.[151,1437,151,789]" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[199,315,727,753]" pageId="14" pageNumber="15">Remarks.</emphasis>
|
||
The sole specimen was examined by
|
||
<bibRefCitation author="Wallach, V." box="[746,920,727,753]" pageId="14" pageNumber="15" pagination="107 - 118" refId="ref19774" refString="Wallach, V. (1996) Two new blind snakes of the Typhlops ater species group from Papua New Guinea (Serpentes: Typhlopidae). Russian Journal of Herpetology, 3, 107 - 118." type="journal article" year="1996">Wallach (1996)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Pyron, R. A. & Wallach, V." box="[977,1256,727,753]" pageId="14" pageNumber="15" pagination="1 - 81" refId="ref19325" refString="Pyron, R. A. & Wallach, V. (2014) Systematics of the blind snakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa, 3829 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 3829.1.1" type="journal article" year="2014">Pyron & Wallach (2014)</bibRefCitation>
|
||
who listed it as
|
||
<taxonomicName authorityName="Wallach" authorityYear="1996" box="[151,366,763,789]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[151,366,763,789]" italics="true" pageId="14" pageNumber="15">Typhlops inornatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |