364 lines
36 KiB
XML
364 lines
36 KiB
XML
<document ID-DOI="10.11646/zootaxa.5231.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7571485" ID-ZooBank="9C66A388-C221-4D17-B2AA-F9EB7F1EC940" checkinTime="1674711822273" checkinUser="plazi" docAuthor="Kraus, Fred" docDate="2023" docId="0381A239FFF1FFD67E8DFB3DFF73FDCE" docLanguage="en" docName="zootaxa.5231.1.1.pdf" docOrigin="Zootaxa 5231 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gerrhopilus slapcinskyi Kraus 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="FFB8DA41FFF4FFD17E1AFFE2FFBAFFD7" masterDocTitle="A revision of Gerrhopilus inornatus (Squamata: Gerrhopilidae) reveals a multi-species complex" masterLastPageNumber="23" masterPageNumber="1" pageNumber="6" updateTime="1674753466069" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of Gerrhopilus inornatus (Squamata: Gerrhopilidae) reveals a multi-species complex</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kraus, Fred</mods:namePart>
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<mods:affiliation>Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, U. S. A</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5231</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>23</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5231.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5231.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7571485</mods:identifier>
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<mods:identifier type="ZooBank">9C66A388-C221-4D17-B2AA-F9EB7F1EC940</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571493" ID-Zenodo-Dep="7571493" LSID="urn:lsid:plazi:treatment:0381A239FFF1FFD67E8DFB3DFF73FDCE" httpUri="http://treatment.plazi.org/id/0381A239FFF1FFD67E8DFB3DFF73FDCE" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
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<subSubSection box="[151,540,1247,1274]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph blockId="5.[151,540,1247,1310]" box="[151,540,1247,1274]" pageId="5" pageNumber="6">
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<heading bold="true" box="[151,540,1247,1274]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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<emphasis bold="true" box="[151,540,1247,1274]" pageId="5" pageNumber="6">
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||
<taxonomicName authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[151,439,1247,1274]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="slapcinskyi" status="sp. nov.">
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<emphasis bold="true" box="[151,439,1247,1274]" italics="true" pageId="5" pageNumber="6">Gerrhopilus slapcinskyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[446,540,1248,1274]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,280,1283,1310]" pageId="5" pageNumber="6" type="description">
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<paragraph blockId="5.[151,540,1247,1310]" box="[151,280,1283,1310]" pageId="5" pageNumber="6">
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<figureCitation box="[151,280,1283,1310]" captionStart="FIGURE 1" captionStartId="4.[152,255,1787,1811]" captionTargetBox="[170,1419,198,1748]" captionTargetId="figure-27@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 20 midbody scale rows. A, B holotype of Gerrhopilus inornatus (BMNH 1946.1.11.80); C, D holotype of Gerrhopilus slapcinskyi (BPBM 18944)." figureDoi="http://doi.org/10.5281/zenodo.7571487" httpUri="https://zenodo.org/record/7571487/files/figure.png" pageId="5" pageNumber="6">Figs. 1C, D</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="5" pageNumber="6" type="materials_examined">
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||
<paragraph blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="6">
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||
<materialsCitation collectingDate="2003-10-20" collectionCode="BPBM" country="Papua New Guinea" elevation="750" latitude="-6.816" location="John Slapcinsky" longLatPrecision="78" longitude="146.692" municipality="Dunch River" pageId="5" pageNumber="6" specimenCode="BPBM 18944" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
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<emphasis bold="true" box="[151,267,1356,1382]" pageId="5" pageNumber="6">
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<typeStatus box="[151,260,1356,1382]" pageId="5" pageNumber="6">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode box="[275,432,1356,1382]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="6" type="Museum">BPBM 18944</specimenCode>
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(field tag FK 8467), collected
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<date box="[786,977,1355,1382]" pageId="5" pageNumber="6" value="2003-10-20">
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<collectingDate box="[786,977,1355,1382]" pageId="5" pageNumber="6" value="2003-10-20">20 October 2003</collectingDate>
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</date>
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by
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<location LSID="urn:lsid:plazi:treatment:0381A239FFF1FFD67E8DFB3DFF73FDCE:8EF745F4FFF1FFD47DE4FAA9FB0CFAB2" box="[1022,1206,1355,1381]" country="Papua New Guinea" latitude="-6.816" longLatPrecision="78" longitude="146.692" municipality="Dunch River" name="John Slapcinsky" pageId="5" pageNumber="6" stateProvince="Morobe">John Slapcinsky</location>
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along
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<collectingMunicipality box="[1285,1432,1355,1381]" pageId="5" pageNumber="6">Dunch River</collectingMunicipality>
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,
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||
<quantity box="[151,229,1391,1418]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="6" unit="km" value="5.6">5.6 km</quantity>
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||
(by air)
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<locationDeviation box="[327,544,1391,1418]" location="Mt Shungol" pageId="5" pageNumber="6">NW of Mt Shungol</locationDeviation>
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summit,
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||
<geoCoordinate box="[650,749,1391,1418]" degrees="6.816" direction="south" orientation="latitude" pageId="5" pageNumber="6" precision="55" value="-6.816">6.816° S</geoCoordinate>
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,
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||
<geoCoordinate box="[762,890,1391,1418]" degrees="146.692" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="55" value="146.692">146.692° E</geoCoordinate>
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||
,
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||
<quantity box="[903,974,1391,1418]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="6" unit="m" value="750.0">
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||
<elevation box="[903,974,1391,1418]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="6" unit="m" value="750.0">750 m</elevation>
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||
</quantity>
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elevation,
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<collectingRegion box="[1098,1296,1391,1417]" country="Papua New Guinea" name="Morobe" pageId="5" pageNumber="6">Morobe Province</collectingRegion>
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,
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||
<collectingCountry name="Papua New Guinea" pageId="5" pageNumber="6">Papua New Guinea</collectingCountry>
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||
.
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||
</materialsCitation>
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</paragraph>
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||
<paragraph blockId="5.[151,1437,1355,2030]" box="[199,745,1463,1490]" pageId="5" pageNumber="6">
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<materialsCitation box="[199,741,1463,1490]" collectingDate="2003-10-20" collectionCode="BPBM" country="Papua New Guinea" elevation="750" latitude="-6.816" location="John Slapcinsky" longLatPrecision="78" longitude="146.692" municipality="Dunch River" pageId="5" pageNumber="6" specimenCode="BPBM 18945" specimenCount="1" stateProvince="Morobe" typeStatus="paratype">
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<emphasis bold="true" box="[199,315,1464,1490]" pageId="5" pageNumber="6">
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<typeStatus box="[199,310,1464,1490]" pageId="5" pageNumber="6">Paratype</typeStatus>
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||
.
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</emphasis>
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||
<specimenCode box="[322,480,1463,1490]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="6" type="Museum">BPBM 18945</specimenCode>
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, same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="diagnosis">
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||
<paragraph blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="6">
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||
<emphasis bold="true" box="[199,322,1499,1525]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
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A
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<taxonomicName box="[351,575,1499,1525]" pageId="5" pageNumber="6">
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<emphasis box="[351,487,1499,1525]" italics="true" pageId="5" pageNumber="6">Gerrhopilus</emphasis>
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species
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</taxonomicName>
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||
characterized by the unique combination of having a rounded snout in lateral view; LSR = 20 at midbody; TSR = 298–318; loreal absent; supralabial imbrication pattern T-V; subocular scale one; presubocular scale absent; a sharp, protruding, tail spine that is black basally and white distally; 1–2 glands in the prefrontal,
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||
<quantity box="[314,384,1607,1634]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="7.62" metricValueMin="0.0" pageId="5" pageNumber="6" unit="in" value="1.5" valueMax="3.0" valueMin="0.0">0–3 in</quantity>
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||
the supraocular,
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||
<quantity box="[572,642,1607,1634]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.286" metricValueMin="1.016" pageId="5" pageNumber="6" unit="in" value="6.5" valueMax="9.0" valueMin="4.0">4–9 in</quantity>
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||
the ocular,
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||
<quantity box="[769,867,1607,1634]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="9.906" metricValueMin="8.89" pageId="5" pageNumber="6" unit="in" value="37.0" valueMax="39.0" valueMin="35.0">35–39 in</quantity>
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||
the preocular, 0 in the frontal,
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||
<quantity box="[1204,1274,1607,1634]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="5" pageNumber="6" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
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||
the subocular; L/W = 35.5–38.6; TL/SVL =
|
||
<quantity box="[476,567,1643,1670]" metricMagnitude="-4" metricUnit="m" metricValue="8.636" pageId="5" pageNumber="6" unit="in" value="0.034">0.034 in</quantity>
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||
<specimenCount box="[573,695,1643,1670]" pageId="5" pageNumber="6" type="female">one female</specimenCount>
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||
and
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||
<quantity box="[747,838,1643,1670]" metricMagnitude="-3" metricUnit="m" metricValue="1.0922" pageId="5" pageNumber="6" unit="in" value="0.043">0.043 in</quantity>
|
||
<specimenCount box="[844,948,1643,1670]" pageId="5" pageNumber="6" type="male">one male</specimenCount>
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||
; and a uniformly dark-brown venter. Its size is small to moderate for this species group (
|
||
<tableCitation box="[639,723,1679,1706]" captionStart="TABLE 1" captionStartId="17.[151,244,552,578]" captionTargetPageId="17" captionText="TABLE 1. Mensural and count data for members of the Gerrhopilus inornatus Group. Forward slashes separate counts on the right and left sides, respectively." pageId="5" pageNumber="6">Table 1</tableCitation>
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||
).
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||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,1355,2030]" pageId="5" pageNumber="6">
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||
<emphasis bold="true" box="[199,584,1715,1742]" pageId="5" pageNumber="6">Comparisons with other species.</emphasis>
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[591,858,1715,1741]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="slapcinskyi">
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||
<emphasis box="[591,858,1715,1741]" italics="true" pageId="5" pageNumber="6">Gerrhopilus slapcinskyi</emphasis>
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||
</taxonomicName>
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||
is distinguished from
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[1108,1268,1715,1742]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fredparkeri">
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||
<emphasis box="[1108,1268,1715,1742]" italics="true" pageId="5" pageNumber="6">G. fredparkeri</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[1283,1382,1715,1742]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hades">
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||
<emphasis box="[1283,1382,1715,1742]" italics="true" pageId="5" pageNumber="6">G. hades</emphasis>
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||
</taxonomicName>
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||
, and
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||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[151,285,1751,1778]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="suturalis">
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<emphasis box="[151,285,1751,1778]" italics="true" pageId="5" pageNumber="6">G. suturalis</emphasis>
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||
</taxonomicName>
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in having 20 longitudinal scale rows at midbody (vs
|
||
<quantity box="[896,955,1751,1777]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="5" pageNumber="6" unit="in" value="16.0">16 in</quantity>
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||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[963,1126,1751,1778]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fredparkeri">
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||
<emphasis box="[963,1126,1751,1778]" italics="true" pageId="5" pageNumber="6">G. fredparkeri</emphasis>
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||
</taxonomicName>
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||
and
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||
<quantity box="[1183,1242,1751,1778]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="5" pageNumber="6" unit="in" value="18.0">18 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[1250,1351,1751,1778]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hades">
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||
<emphasis box="[1250,1351,1751,1778]" italics="true" pageId="5" pageNumber="6">G. hades</emphasis>
|
||
</taxonomicName>
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||
and
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis italics="true" pageId="5" pageNumber="6">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
). It is further distinguished from
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[625,787,1787,1814]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fredparkeri">
|
||
<emphasis box="[625,787,1787,1814]" italics="true" pageId="5" pageNumber="6">G. fredparkeri</emphasis>
|
||
</taxonomicName>
|
||
in its lower number of transverse scale rows (298–318 vs
|
||
<quantity box="[151,222,1823,1850]" metricMagnitude="1" metricUnit="m" metricValue="1.36906" pageId="5" pageNumber="6" unit="in" value="539.0">539 in</quantity>
|
||
<taxonomicName baseAuthorityName="Wallach" baseAuthorityYear="1996" box="[229,390,1823,1850]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fredparkeri">
|
||
<emphasis box="[229,390,1823,1850]" italics="true" pageId="5" pageNumber="6">G. fredparkeri</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[473,572,1823,1850]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[473,572,1823,1850]" italics="true" pageId="5" pageNumber="6">G. hades</emphasis>
|
||
</taxonomicName>
|
||
in having 22 longitudinal scale rows behind the head (vs
|
||
<quantity box="[1217,1274,1823,1850]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="5" pageNumber="6" unit="in" value="18.0">18 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[1280,1379,1823,1850]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[1280,1379,1823,1850]" italics="true" pageId="5" pageNumber="6">G. hades</emphasis>
|
||
</taxonomicName>
|
||
) and more and smaller glands in the center of the preocular (35–39 vs
|
||
<quantity box="[866,963,1859,1886]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="5" pageNumber="6" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kraus" baseAuthorityYear="2005" box="[969,1067,1859,1886]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hades">
|
||
<emphasis box="[969,1067,1859,1886]" italics="true" pageId="5" pageNumber="6">G. hades</emphasis>
|
||
</taxonomicName>
|
||
), and from
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[1196,1327,1859,1886]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[1196,1327,1859,1886]" italics="true" pageId="5" pageNumber="6">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
in having a single postocular on each side of the head (vs two in
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" box="[781,915,1895,1922]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis box="[781,915,1895,1922]" italics="true" pageId="5" pageNumber="6">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
) and lacking a presubocular (vs present in
|
||
<taxonomicName baseAuthorityName="Brongersma" baseAuthorityYear="1934" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="suturalis">
|
||
<emphasis italics="true" pageId="5" pageNumber="6">G. suturalis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[273,541,1931,1957]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="slapcinskyi">
|
||
<emphasis box="[273,541,1931,1957]" italics="true" pageId="5" pageNumber="6">Gerrhopilus slapcinskyi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName authorityName="Vidal, Wynn, Donnellan & Hedges" authorityYear="2010" box="[692,834,1932,1958]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[692,834,1932,1958]" italics="true" pageId="5" pageNumber="6">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
in having fewer or more transverse scale rows (298– 318 vs
|
||
<quantity box="[231,357,1967,1994]" metricMagnitude="0" metricUnit="m" metricValue="9.5123" metricValueMax="9.525" metricValueMin="9.4996" pageId="5" pageNumber="6" unit="in" value="374.5" valueMax="375.0" valueMin="374.0">374–375 in</quantity>
|
||
<taxonomicName authorityName="Vidal, Wynn, Donnellan & Hedges" authorityYear="2010" box="[363,504,1968,1994]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[363,504,1968,1994]" italics="true" pageId="5" pageNumber="6">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
), a greater number of glands in the preocular (35–39 vs
|
||
<quantity box="[1128,1226,1967,1994]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="5" pageNumber="6" unit="in" value="23.0" valueMax="24.0" valueMin="22.0">22–24 in</quantity>
|
||
<taxonomicName authorityName="Vidal, Wynn, Donnellan & Hedges" authorityYear="2010" box="[1232,1373,1968,1994]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[1232,1373,1968,1994]" italics="true" pageId="5" pageNumber="6">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
), and longer tail (TL/SVL = 0.034 vs
|
||
<quantity box="[507,599,2003,2029]" metricMagnitude="-4" metricUnit="m" metricValue="6.604" pageId="5" pageNumber="6" unit="in" value="0.026">0.026 in</quantity>
|
||
female and 0.043 vs
|
||
<quantity box="[839,931,2003,2030]" metricMagnitude="-4" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="6" unit="in" value="0.03">0.030 in</quantity>
|
||
male
|
||
<taxonomicName authorityName="Vidal, Wynn, Donnellan & Hedges" authorityYear="2010" box="[999,1140,2004,2030]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[999,1140,2004,2030]" italics="true" pageId="5" pageNumber="6">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
|
||
<paragraph blockId="6.[151,1437,150,1293]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[199,529,150,177]" pageId="6" pageNumber="7">
|
||
Description of the
|
||
<typeStatus box="[420,523,151,177]" pageId="6" pageNumber="7">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. L =
|
||
<quantity box="[679,796,150,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.955" pageId="6" pageNumber="7" unit="mm" value="195.5">195.5 mm</quantity>
|
||
, SVL =
|
||
<quantity box="[889,985,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.89" pageId="6" pageNumber="7" unit="mm" value="189.0">189 mm</quantity>
|
||
, TL =
|
||
<quantity box="[1059,1147,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="7" unit="mm" value="6.5">6.5 mm</quantity>
|
||
, HW =
|
||
<quantity box="[1235,1323,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="7" unit="mm" value="4.3">4.3 mm</quantity>
|
||
, SN =
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="7" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, SW =
|
||
<quantity box="[278,365,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, PSN = 1.0 mm, RW =
|
||
<quantity box="[625,712,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="7" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, EW =
|
||
<quantity box="[795,882,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="mm" value="0.4">0.4 mm</quantity>
|
||
, W =
|
||
<quantity box="[946,1034,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="7" unit="mm" value="5.5">5.5 mm</quantity>
|
||
, VW =
|
||
<quantity box="[1118,1206,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="7" unit="mm" value="4.3">4.3 mm</quantity>
|
||
, TW =
|
||
<quantity box="[1287,1375,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="7" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, L/W = 35.5, TL/SVL = 0.034. Head slightly wider than neck. Snout rounded in dorsal and lateral views. Snout anterior to lower jaw angled slightly ventrally. Rostral moderately large (RW/HW = 0.42), oval dorsally, lateral margins convex, posterior border extending approximately two-thirds way between naris and eye, posterior margin straight; ventrally surface papillose, with straight sides and posterior margin concave. Nasals separated dorsally by prefrontal (
|
||
<figureCitation box="[159,244,367,393]" captionStart="FIGURE 1" captionStartId="4.[152,255,1787,1811]" captionTargetBox="[170,1419,198,1748]" captionTargetId="figure-27@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 20 midbody scale rows. A, B holotype of Gerrhopilus inornatus (BMNH 1946.1.11.80); C, D holotype of Gerrhopilus slapcinskyi (BPBM 18944)." figureDoi="http://doi.org/10.5281/zenodo.7571487" httpUri="https://zenodo.org/record/7571487/files/figure.png" pageId="6" pageNumber="7">Fig. 1C</figureCitation>
|
||
); superior nasal large, with sinuous posterior margin, concave dorsally, convex ventrally (
|
||
<figureCitation box="[1235,1322,367,393]" captionStart="FIGURE 1" captionStartId="4.[152,255,1787,1811]" captionTargetBox="[170,1419,198,1748]" captionTargetId="figure-27@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal and lateral views of heads for members of the Gerrhopilus inornatus Group having 20 midbody scale rows. A, B holotype of Gerrhopilus inornatus (BMNH 1946.1.11.80); C, D holotype of Gerrhopilus slapcinskyi (BPBM 18944)." figureDoi="http://doi.org/10.5281/zenodo.7571487" httpUri="https://zenodo.org/record/7571487/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
|
||
). External naris semicircular, oriented obliquely, close to rostral, anterior half covered by inferior nasal; superior nasal suture extending anterodorsally from naris to rostral; inferior nasal suture complete, contacting second supralabial well posterior to latter’s contact with first supralabial. Prefrontal and supraoculars subequal in size, latter wider, same size as frontal but larger than parietals and interparietal, which are also subequal in size. Preocular large, triangular; larger than ocular but smaller than superior nasal. Ocular large, smaller than preocular, extending dorsally well above preocular, extending ventrally to ~2/3 depth of preocular, bordered posteroventrally by subocular of less than half its size. Eye distinct, with large distinct pupil, situated at widest point of ocular and closer to its dorsal than ventral margin, anterior 40% covered by preocular plate in lateral view. Four postoculars bordering ocular and subocular between parietal and fourth supralabial. Four supralabials, third the largest, all with long axis oblique to long axis of body, first approximately rectangular. Supralabial imbrication pattern T-V, posterior border of second supralabial overlaps anteroventral margin of preocular, that of third supralabial overlaps anteroventral margin of subocular and ends just posterior to rear margin of preocular. Mental crescentic, wider than long, projecting slightly beyond curve of lower jaw and fitting into notch on upper lip when mouth is closed. Infralabials two on each side, second much longer.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1293]" pageId="6" pageNumber="7">
|
||
Longitudinal scale rows 22 behind head, 20 at midbody, and 20 anterior to vent; transverse scale rows 318,
|
||
<specimenCount pageId="6" pageNumber="7" type="intercalary">ten intercalary</specimenCount>
|
||
scales along vertebral row; subcaudals 15; dorsocaudals 15; apical region with sharp, slightly downturned spine that extends
|
||
<quantity box="[359,445,979,1005]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="mm" value="0.4">0.4 mm</quantity>
|
||
past last scales.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1293]" pageId="6" pageNumber="7">Rostral, nasals, and preoculars densely covered in pale glands; oculars with 4 (R) and 9 (L) glands, preoculars with 35 (R) and 37 (L), supraoculars with 0 (R) and 1 (L), prefrontal with 2; suboculars with 4 (R) and 3 (L), and frontal without glands.</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1293]" pageId="6" pageNumber="7">
|
||
In preservative, 19 years after preservation, body dark brown above and below but somewhat paler below; no sharp distinction between dorsal and ventral shading; each scale uniformly dark. Anterior tip and margins of rostral, area around nares, first and second supralabials, mental, infralabials, and center of throat to three scales behind mental white; second supralabials white with brown along dorsal margins. Head glands white; tail spine dark brown basally, white distally. Iris dark gray; pupil paler gray. In life, the
|
||
<typeStatus box="[884,982,1267,1293]" pageId="6" pageNumber="7">holotype</typeStatus>
|
||
was black.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571495" ID-Zenodo-Dep="7571495" httpUri="https://zenodo.org/record/7571495/files/figure.png" pageId="6" pageNumber="7" startId="6.[152,255,1767,1791]" targetBox="[152,1436,1317,1744]" targetPageId="6">
|
||
<paragraph blockId="6.[151,1437,1767,1864]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[152,281,1767,1791]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
|
||
Habitats for members of the
|
||
<taxonomicName authorityName="Vidal, Wynn, Donnellan & Hedges" authorityYear="2010" box="[580,808,1767,1791]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[580,808,1767,1791]" italics="true" pageId="6" pageNumber="7">Gerrhopilus inornatus</emphasis>
|
||
</taxonomicName>
|
||
Group.
|
||
<emphasis bold="true" box="[892,910,1767,1791]" pageId="6" pageNumber="7">A</emphasis>
|
||
Forest at Varirata National Park, very near the type locality of
|
||
<taxonomicName authorityName="Vidal, Wynn, Donnellan & Hedges" authorityYear="2010" box="[263,392,1804,1827]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis box="[263,392,1804,1827]" italics="true" pageId="6" pageNumber="7">G. inornatus</emphasis>
|
||
</taxonomicName>
|
||
(photo M. O’Shea);
|
||
<emphasis bold="true" box="[608,625,1804,1828]" pageId="6" pageNumber="7">B</emphasis>
|
||
the hills around Bunisi and Siyomu from which
|
||
<taxonomicName authorityName="Kraus" authorityYear="2023" box="[1129,1290,1804,1827]" class="Reptilia" family="Gerrhopilidae" genus="Gerrhopilus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="papuanorum">
|
||
<emphasis box="[1129,1290,1804,1827]" italics="true" pageId="6" pageNumber="7">G. papuanorum</emphasis>
|
||
</taxonomicName>
|
||
was collected (photo F. Kraus).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[151,1436,1909,2008]" lastBlockId="7.[151,1437,151,537]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[199,319,1909,1935]" pageId="6" pageNumber="7">Variation.</emphasis>
|
||
The
|
||
<typeStatus box="[378,474,1910,1936]" pageId="6" pageNumber="7">paratype</typeStatus>
|
||
(
|
||
<specimenCode box="[491,649,1910,1937]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="7" type="Museum">BPBM 18945</specimenCode>
|
||
) differs from the holotype in being male, somewhat shorter (L =
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="7" unit="mm" value="170.0">170 mm</quantity>
|
||
, SVL =
|
||
<quantity box="[303,402,1946,1972]" metricMagnitude="-1" metricUnit="m" metricValue="1.63" pageId="6" pageNumber="7" unit="mm" value="163.0">163 mm</quantity>
|
||
, TL =
|
||
<quantity box="[488,559,1946,1972]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="7" unit="mm" value="7.0">7 mm</quantity>
|
||
,
|
||
<tableCitation box="[574,660,1946,1972]" captionStart="TABLE 1" captionStartId="17.[151,244,552,578]" captionTargetPageId="17" captionText="TABLE 1. Mensural and count data for members of the Gerrhopilus inornatus Group. Forward slashes separate counts on the right and left sides, respectively." pageId="6" pageNumber="7">Table 1</tableCitation>
|
||
), approximately the same width (W =
|
||
<quantity box="[1123,1215,1946,1972]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="7" unit="mm" value="4.4">4.4 mm</quantity>
|
||
, L/W = 38.6), and with a longer tail (TL/SVL = 0.043). It too has 22-20-20 longitudinal scale rows, 298 transverse scale rows,
|
||
<specimenCount lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="intercalary">nine intercalary</specimenCount>
|
||
scales, 16 subcaudals, 16 dorsocaudals, the eye one-third covered by the preocular, oculars with 8 (
|
||
<collectionCode box="[1407,1428,151,177]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="8">R</collectionCode>
|
||
) and 9 (L) glands, preocular with 36 (
|
||
<collectionCode box="[576,597,187,213]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="8">R</collectionCode>
|
||
) and 39 (L), supraoculars with 3 (
|
||
<collectionCode box="[992,1013,187,213]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="8">R</collectionCode>
|
||
) and 2 (L), prefrontal with 1, frontal with 0, and suboculars with 4 (
|
||
<collectionCode box="[495,516,223,249]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="8">R</collectionCode>
|
||
) and 3 (L) glands. Like the
|
||
<typeStatus box="[828,929,223,249]" pageId="7" pageNumber="8">holotype</typeStatus>
|
||
, it was black in life. The
|
||
<typeStatus box="[1210,1308,223,249]" pageId="7" pageNumber="8">holotype</typeStatus>
|
||
has greater asymmetry in ocular gland numbers and slightly more mid-dorsal scale rows, but otherwise variation between the
|
||
<specimenCount box="[151,317,295,321]" pageId="7" pageNumber="8" type="generic">two specimens</specimenCount>
|
||
is slight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="8" type="etymology">
|
||
<paragraph blockId="7.[151,1437,151,537]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[199,334,331,357]" pageId="7" pageNumber="8">Etymology.</emphasis>
|
||
This species is named for malacologist John Slapcinsky, who collected the
|
||
<typeStatus box="[1212,1260,331,357]" pageId="7" pageNumber="8">type</typeStatus>
|
||
specimens and proved a great field companion on several of my expeditions to
|
||
<collectingCountry box="[867,1088,367,393]" name="Papua New Guinea" pageId="7" pageNumber="8">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,988,402,429]" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph blockId="7.[151,1437,151,537]" box="[199,988,402,429]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[199,353,402,428]" pageId="7" pageNumber="8">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus box="[608,656,403,429]" pageId="7" pageNumber="8">type</typeStatus>
|
||
locality, west of Lae (
|
||
<figureCitation box="[906,975,403,429]" captionStart="FIGURE 2" captionStartId="5.[152,255,1069,1093]" captionTargetBox="[181,1408,456,1045]" captionTargetId="figure-148@5.[179,1408,453,1045]" captionTargetPageId="5" captionText="FIGURE 2. Map of southern Papua New Guinea showing collecting localities for Gerrhopilus inornatus (star), G. flavinotatus (diamond), G. lorealis (hexagon), G. papuanorum (square), G. polyadenus (open circles), G. slapcinskyi (filled circle), and G. wallachi (triangle)." figureDoi="http://doi.org/10.5281/zenodo.7571491" httpUri="https://zenodo.org/record/7571491/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="8" type="biology_ecology">
|
||
<paragraph blockId="7.[151,1437,151,537]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[199,300,439,465]" pageId="7" pageNumber="8">Ecology.</emphasis>
|
||
Both specimens were taken at night in primary lowland hill rainforest with a canopy of approximately
|
||
<quantity box="[151,209,475,501]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="m" value="30.0">30 m</quantity>
|
||
height and a moderately dense understory. One of the specimens was found while climbing on the trunk of a tree.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |