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1.
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Aviorrhynchidae
<bibRefCitation id="03241F9A333C724EFE61F932FC8579B0" author="Nel, A. &amp; Roques, P. &amp; Nel, P. &amp; Prokin, A. A. &amp; Bourgoin, T. &amp; Prokop, J. &amp; Szwedo, J. &amp; Azar, D. &amp; Desutter-Grandcolas, L. &amp; Wappler, T. &amp; Garrouste, R. &amp; Coty, D. &amp; Huang, D. Y. &amp; Engel, M. S. &amp; Kirejtshuk, A. G." box="[398,866,1713,1739]" pageId="1" pageNumber="563" pagination="257 - 261" refId="ref3745" refString="Nel, A., Roques, P., Nel, P., Prokin, A. A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D. Y., Engel, M. S. &amp; Kirejtshuk, A. G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257 - 261. https: // doi. org / 10.1038 / nature 12629" type="journal article" year="2013">Nel, Bourgoin, Engel &amp; Szwedo, 2013</bibRefCitation>
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<taxonomicName id="A0B519E8333C724EFF78F975FEA67875" authorityName="Nel, Bourgoin, Engel &amp; Szwedo" authorityYear="2013" box="[151,321,1782,1806]" class="Insecta" family="Aviorrhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="563" phylum="Arthropoda" rank="family">Aviorrhynchidae</taxonomicName>
sec.
<bibRefCitation id="03241F9A333C724EFE9BF975FDF17875" author="Nel, A. &amp; Roques, P. &amp; Nel, P. &amp; Prokin, A. A. &amp; Bourgoin, T. &amp; Prokop, J. &amp; Szwedo, J. &amp; Azar, D. &amp; Desutter-Grandcolas, L. &amp; Wappler, T. &amp; Garrouste, R. &amp; Coty, D. &amp; Huang, D. Y. &amp; Engel, M. S. &amp; Kirejtshuk, A. G." box="[372,534,1782,1806]" pageId="1" pageNumber="563" pagination="257 - 261" refId="ref3745" refString="Nel, A., Roques, P., Nel, P., Prokin, A. A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D. Y., Engel, M. S. &amp; Kirejtshuk, A. G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257 - 261. https: // doi. org / 10.1038 / nature 12629" type="journal article" year="2013">
Nel
<emphasis id="55C1BE79333C724EFE70F974FE347875" box="[415,467,1782,1806]" italics="true" pageId="1" pageNumber="563">et al.</emphasis>
, 2013
</bibRefCitation>
: family: 258.
</paragraph>
<paragraph id="670A626B333C724EFF78F8BEFB72782C" blockId="1.[151,1437,1852,2023]" box="[151,1173,1852,1879]" pageId="1" pageNumber="563">
Type-genus:
<taxonomicName id="A0B519E8333C724EFEC7F8BFFC9E782D" authority="Nel, Bourgoin, Engel &amp; Szwedo, 2013" authorityName="Nel, Bourgoin, Engel &amp; Szwedo" authorityYear="2013" box="[296,889,1852,1878]" class="Insecta" family="Aviorrhynchidae" genus="Aviorrhyncha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="563" phylum="Arthropoda" rank="genus">
<emphasis id="55C1BE79333C724EFEC7F8BFFE27782D" box="[296,448,1852,1878]" italics="true" pageId="1" pageNumber="563">Aviorrhyncha</emphasis>
<bibRefCitation id="03241F9A333C724EFE28F8BFFC9E782D" author="Nel, A. &amp; Roques, P. &amp; Nel, P. &amp; Prokin, A. A. &amp; Bourgoin, T. &amp; Prokop, J. &amp; Szwedo, J. &amp; Azar, D. &amp; Desutter-Grandcolas, L. &amp; Wappler, T. &amp; Garrouste, R. &amp; Coty, D. &amp; Huang, D. Y. &amp; Engel, M. S. &amp; Kirejtshuk, A. G." box="[455,889,1852,1878]" pageId="1" pageNumber="563" pagination="257 - 261" refId="ref3745" refString="Nel, A., Roques, P., Nel, P., Prokin, A. A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D. Y., Engel, M. S. &amp; Kirejtshuk, A. G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257 - 261. https: // doi. org / 10.1038 / nature 12629" type="journal article" year="2013">Nel, Bourgoin, Engel &amp; Szwedo, 2013</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="03241F9A333C724EFC49F8BFFB6A782C" author="Nel, A. &amp; Roques, P. &amp; Nel, P. &amp; Prokin, A. A. &amp; Bourgoin, T. &amp; Prokop, J. &amp; Szwedo, J. &amp; Azar, D. &amp; Desutter-Grandcolas, L. &amp; Wappler, T. &amp; Garrouste, R. &amp; Coty, D. &amp; Huang, D. Y. &amp; Engel, M. S. &amp; Kirejtshuk, A. G." box="[934,1165,1852,1879]" pageId="1" pageNumber="563" pagination="257 - 261" refId="ref3745" refString="Nel, A., Roques, P., Nel, P., Prokin, A. A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D. Y., Engel, M. S. &amp; Kirejtshuk, A. G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257 - 261. https: // doi. org / 10.1038 / nature 12629" type="journal article" year="2013">
Nel
<emphasis id="55C1BE79333C724EFC3AF8BEFBEA782D" box="[981,1037,1852,1878]" italics="true" pageId="1" pageNumber="563">et al.</emphasis>
, 2013: 258
</bibRefCitation>
)
</paragraph>
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Type-species:
<taxonomicName id="A0B519E8333C724EFE86F8E3FBDD7801" authority="Nel, Bourgoin, Engel &amp; Szwedo, 2013" authorityName="Nel, Bourgoin, Engel &amp; Szwedo" authorityYear="2013" box="[361,1082,1888,1914]" class="Insecta" family="Aviorrhynchidae" genus="Aviorrhyncha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="563" phylum="Arthropoda" rank="species" species="magnifica">
<emphasis id="55C1BE79333C724EFE86F8E3FD9E7801" box="[361,633,1888,1914]" italics="true" pageId="1" pageNumber="563">Aviorrhyncha magnifica</emphasis>
<bibRefCitation id="03241F9A333C724EFD6DF8E3FBDD7801" author="Nel, A. &amp; Roques, P. &amp; Nel, P. &amp; Prokin, A. A. &amp; Bourgoin, T. &amp; Prokop, J. &amp; Szwedo, J. &amp; Azar, D. &amp; Desutter-Grandcolas, L. &amp; Wappler, T. &amp; Garrouste, R. &amp; Coty, D. &amp; Huang, D. Y. &amp; Engel, M. S. &amp; Kirejtshuk, A. G." box="[642,1082,1888,1914]" pageId="1" pageNumber="563" pagination="257 - 261" refId="ref3745" refString="Nel, A., Roques, P., Nel, P., Prokin, A. A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D. Y., Engel, M. S. &amp; Kirejtshuk, A. G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257 - 261. https: // doi. org / 10.1038 / nature 12629" type="journal article" year="2013">Nel, Bourgoin, Engel &amp; Szwedo, 2013</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="03241F9A333C724EFB86F8E3FABF7800" author="Nel, A. &amp; Roques, P. &amp; Nel, P. &amp; Prokin, A. A. &amp; Bourgoin, T. &amp; Prokop, J. &amp; Szwedo, J. &amp; Azar, D. &amp; Desutter-Grandcolas, L. &amp; Wappler, T. &amp; Garrouste, R. &amp; Coty, D. &amp; Huang, D. Y. &amp; Engel, M. S. &amp; Kirejtshuk, A. G." box="[1129,1368,1888,1915]" pageId="1" pageNumber="563" pagination="257 - 261" refId="ref3745" refString="Nel, A., Roques, P., Nel, P., Prokin, A. A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D. Y., Engel, M. S. &amp; Kirejtshuk, A. G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257 - 261. https: // doi. org / 10.1038 / nature 12629" type="journal article" year="2013">
Nel
<emphasis id="55C1BE79333C724EFB75F8E2FB337801" box="[1178,1236,1888,1914]" italics="true" pageId="1" pageNumber="563">et al.</emphasis>
, 2013: 258
</bibRefCitation>
, 2c-e; Extended data figs. 5a, b; 6a)
</paragraph>
<paragraph id="670A626B333C724EFF28F82BFB26789D" blockId="1.[151,1437,1852,2023]" pageId="1" pageNumber="563">
Chronostratigraphic occurrence: Carboniferous, Moscovian (315.2-308 Ma). Known from single fossil occurrence:
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<emphasis id="55C1BE79333C724EFECEF84FFDD7789D" box="[289,560,1996,2022]" italics="true" pageId="1" pageNumber="563">Aviorrhyncha magnifica</emphasis>
<bibRefCitation id="03241F9A333C724EFDD8F84FFC0F789D" author="Nel, A. &amp; Roques, P. &amp; Nel, P. &amp; Prokin, A. A. &amp; Bourgoin, T. &amp; Prokop, J. &amp; Szwedo, J. &amp; Azar, D. &amp; Desutter-Grandcolas, L. &amp; Wappler, T. &amp; Garrouste, R. &amp; Coty, D. &amp; Huang, D. Y. &amp; Engel, M. S. &amp; Kirejtshuk, A. G." box="[567,1000,1996,2022]" pageId="1" pageNumber="563" pagination="257 - 261" refId="ref3745" refString="Nel, A., Roques, P., Nel, P., Prokin, A. A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D. Y., Engel, M. S. &amp; Kirejtshuk, A. G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257 - 261. https: // doi. org / 10.1038 / nature 12629" type="journal article" year="2013">
Nel, Bourgoin, Engel &amp; Szwedo,
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in
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Nel
<emphasis id="55C1BE79333C724EFBD4F84EFB93789D" box="[1083,1140,1996,2022]" italics="true" pageId="1" pageNumber="563">et al.</emphasis>
, 2013
</bibRefCitation>
.
</paragraph>
<paragraph id="670A626B333F724DFF28FF14FC317F82" blockId="2.[151,1437,151,249]" pageId="2" pageNumber="564">
Note. The monospecific
<taxonomicName id="A0B519E8333F724DFE0BFF14FD477FCA" authorityName="Nel, Bourgoin, Engel &amp; Szwedo" authorityYear="2013" box="[484,672,151,177]" class="Insecta" family="Aviorrhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="564" phylum="Arthropoda" rank="family">Aviorrhynchidae</taxonomicName>
belong to a distinct lineage from the following Eucixioidian one. Both lineages would need a more specific analysis before gaining a formal rank within the clade Euhemiptera sec. Zrzavý, 1990 (synonym of Hemelytrata sec. Fallén, 1829;
<bibRefCitation id="03241F9A333F724DFCC8FF5CFC2D7F82" author="Szwedo, J." box="[807,970,223,249]" pageId="2" pageNumber="564" pagination="109 - 128" refId="ref4159" refString="Szwedo, J. (2018) The unity, diversity and conformity of bugs (Hemiptera) through time. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 107 (2 - 3), 109 - 128. [2017] https: // doi. org / 10.1017 / S 175569101700038 X" type="journal article" year="2018">Szwedo, 2018</bibRefCitation>
).
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