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<document id="FB79179FB22D8B801200D3D6D82971EE" ID-DOI="10.1080/00222933.2011.590947" ID-ISSN="1464-5262" ID-Zenodo-Dep="5204287" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629052250660" checkinUser="felipe" docAuthor="Rueda Martín, Paola A. &amp; Sganga, Julieta V." docDate="2011" docId="BF5487DEFFC19C2209114715FD19FEF6" docLanguage="en" docName="JNatHist.45.35-36.2215-2230.pdf" docOrigin="Journal of Natural History 45 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222933.2011.590947" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Smicridea (Rhyacophylax) valeni Rueda Martin and Sganga 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2225" masterDocId="436DFFA6FFCB9C2E0825463FFFD4FF8B" masterDocTitle="Smicridea McLachlan (Trichoptera: Hydropsychidae) from northwestern Argentina and Bolivia: new species, redescription, association and new records" masterLastPageNumber="2230" masterPageNumber="2215" pageNumber="2223" updateTime="1699058564112" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1B46EEDB6B4DC8970AC59BD1B4C8520F">Smicridea McLachlan (Trichoptera: Hydropsychidae) from northwestern Argentina and Bolivia: new species, redescription, association and new records</mods:title>
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<mods:name id="E50241A737560016C9B66D2C9FAA5C78" type="personal">
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<mods:namePart id="BA2D1AEFD8FEC5B663F5A6FD7AD73B1F">Rueda Martín, Paola A.</mods:namePart>
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<mods:name id="E7165357F930AC64E274992287D2F789" type="personal">
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<mods:namePart id="9559105F9244AE88CDF32AC2C0044FCD">Sganga, Julieta V.</mods:namePart>
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<mods:typeOfResource id="501B8FB5672A25082EDFFEF78BEC4854">text</mods:typeOfResource>
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<mods:title id="BCAABC95B10E0CE4B2F5BC35F8EBC441">Journal of Natural History</mods:title>
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<mods:date id="0921CCF1D3C3773539641C63B04D45ED">2011</mods:date>
<mods:detail id="751968EFBA1A5213A4143E458AA75FC7" type="pubDate">
<mods:number id="57E2830F7E1CFB426B570CA34B0FD7CE">2011-09-30</mods:number>
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<mods:number id="58258ADAA0866EA57044543FFF819372">45</mods:number>
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<mods:number id="C171215B51DDBD8CFB05CBA96156E8E8">35 - 36</mods:number>
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<mods:url id="21FC5BAE6FBB8BB38D4FA50073FE4752">http://dx.doi.org/10.1080/00222933.2011.590947</mods:url>
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<mods:identifier id="80F70EFD03FABB253291691851DCCC07" type="DOI">10.1080/00222933.2011.590947</mods:identifier>
<mods:identifier id="C275950AB5A668BAE54B276BFEA737D8" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="FA9D2DA0021072AA7037715923B9F014" type="Zenodo-Dep">5204287</mods:identifier>
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<treatment id="BF5487DEFFC19C2209114715FD19FEF6" LSID="urn:lsid:plazi:treatment:BF5487DEFFC19C2209114715FD19FEF6" httpUri="http://treatment.plazi.org/id/BF5487DEFFC19C2209114715FD19FEF6" lastPageId="12" lastPageNumber="2225" pageId="10" pageNumber="2223">
<subSubSection id="7FE76543FFC19C2409114715FBE7FEC8" box="[308,1075,298,323]" pageId="10" pageNumber="2223" type="nomenclature">
<paragraph id="374236C8FFC19C2409114715FBE7FEC8" blockId="10.[308,1075,298,355]" box="[308,1075,298,323]" pageId="10" pageNumber="2223">
<heading id="6C0A81A4FFC19C2409114715FBE7FEC8" box="[308,1075,298,323]" centered="true" fontSize="36" level="2" pageId="10" pageNumber="2223" reason="7">
<taxonomicName id="F0FD4D4BFFC19C2409114715FC00FEC8" authority="Rueda Martin and Sganga" authorityName="Rueda Martin and Sganga" authorityYear="2011" box="[308,980,298,323]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="2223" phylum="Arthropoda" rank="species" species="valeni" status="sp. nov." subGenus="Rhyacophylax">
<emphasis id="0589EADAFFC19C2409114715FD4BFEC8" bold="true" box="[308,671,298,323]" pageId="10" pageNumber="2223">
<emphasis id="0589EADAFFC19C2409114715FE77FEC8" bold="true" box="[308,419,298,323]" italics="true" pageId="10" pageNumber="2223">Smicridea</emphasis>
(
<emphasis id="0589EADAFFC19C2409964715FD84FEC8" bold="true" box="[435,592,298,323]" italics="true" pageId="10" pageNumber="2223">Rhyacophylax</emphasis>
)
<emphasis id="0589EADAFFC19C240A7A4715FD4BFEC8" bold="true" box="[607,671,298,323]" italics="true" pageId="10" pageNumber="2223">valeni</emphasis>
</emphasis>
Rueda Martín and Sganga
</taxonomicName>
,
<taxonomicNameLabel id="1EBA57A1FFC19C240BFA4713FBE7FEC8" box="[991,1075,300,323]" pageId="10" pageNumber="2223" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FE76543FFC19C220A794774FD19FEF6" lastPageId="12" lastPageNumber="2225" pageId="10" pageNumber="2223" type="description">
<paragraph id="374236C8FFC19C240A794774FCD0FEE8" blockId="10.[308,1075,298,355]" box="[604,772,331,355]" pageId="10" pageNumber="2223">
(
<figureCitation id="AFC62A4DFFC19C240A414774FD28FEE8" box="[612,764,331,355]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4AG</figureCitation>
)
</paragraph>
<paragraph id="374236C8FFC19C2408F0479CFE73FE30" blockId="10.[213,423,419,443]" box="[213,423,419,443]" pageId="10" pageNumber="2223">
<emphasis id="0589EADAFFC19C2408F0479CFE73FE30" box="[213,423,419,443]" italics="true" pageId="10" pageNumber="2223">Material examined</emphasis>
</paragraph>
<paragraph id="374236C8FFC19C2408F047F4FE2FFD89" blockId="10.[213,1170,458,514]" pageId="10" pageNumber="2223">
<materialsCitation id="87953C95FFC19C2408F047F4FE2FFD89" collectingDate="2008-11-01" collectionCode="IML" collectorName="El Siambon &amp; Rio el Siambon &amp; Rueda Martin" country="Argentina" location="El Siambon" pageId="10" pageNumber="2223" specimenCount="1" stateProvince="Tucuman" typeStatus="holotype">
<emphasis id="0589EADAFFC19C2408F047F4FEA9FE68" box="[213,381,459,483]" italics="true" pageId="10" pageNumber="2223">
<typeStatus id="E846886AFFC19C2408F047F4FEE8FE68" box="[213,316,459,483]" pageId="10" pageNumber="2223" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<collectingCountry id="4FEA7658FFC19C2409A147F5FE2DFE69" box="[388,505,458,482]" name="Argentina" pageId="10" pageNumber="2223">Argentina</collectingCountry>
:
<collectingRegion id="F539F82AFFC19C240A2647F5FDA5FE69" box="[515,625,458,482]" country="Argentina" name="Tucuman" pageId="10" pageNumber="2223">Tucumán</collectingRegion>
,
<collectorName id="9A08531EFFC19C240A5F47F5FCD5FE69" box="[634,769,458,482]" pageId="10" pageNumber="2223">
<location id="32226013FFC19C240A5F47F5FCD5FE69" LSID="urn:lsid:plazi:treatment:BF5487DEFFC19C2209114715FD19FEF6:32226013FFC19C240A5F47F5FCD5FE69" box="[634,769,458,482]" country="Argentina" name="El Siambon" pageId="10" pageNumber="2223" stateProvince="Tucuman">El Siambón</location>
</collectorName>
,
<collectorName id="9A08531EFFC19C240B2E47F5FC68FE69" box="[779,956,458,482]" pageId="10" pageNumber="2223">Río el Siambón</collectorName>
,
<date id="43431008FFC19C240BE347F5FB59FE69" box="[966,1165,458,482]" pageId="10" pageNumber="2223" value="2008-11-01">
<collectingDate id="5307E9E0FFC19C240BE347F5FB59FE69" box="[966,1165,458,482]" pageId="10" pageNumber="2223" value="2008-11-01">1 November 2008</collectingDate>
</date>
,
<collectorName id="9A08531EFFC19C2408F047D5FEA2FD89" box="[213,374,490,514]" pageId="10" pageNumber="2223">Rueda Martín</collectorName>
col. (
<collectionCode id="51ECAE0DFFC19C24099147D5FE3BFD89" box="[436,495,490,514]" country="Argentina" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="10" pageNumber="2223">IML</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="374236C8FFC19C2408F04414FCD0FD2A" blockId="10.[213,1170,554,673]" pageId="10" pageNumber="2223">
<materialsCitation id="87953C95FFC19C2408F04414FDCAFDE9" collectingDate="2008-11-01" collectorName="Rio el Siambon &amp; Rueda Martin" country="Argentina" location="El Siambon" pageId="10" pageNumber="2223" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman" typeStatus="paratype">
<emphasis id="0589EADAFFC19C2408F04414FE9EFDC9" box="[213,330,555,578]" italics="true" pageId="10" pageNumber="2223">
<typeStatus id="E846886AFFC19C2408F04414FE91FDC9" box="[213,325,555,578]" pageId="10" pageNumber="2223" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="4FEA7658FFC19C2409744415FE12FDC9" box="[337,454,554,578]" name="Argentina" pageId="10" pageNumber="2223">Argentina</collectingCountry>
:
<collectingRegion id="F539F82AFFC19C2409F04415FD97FDC9" box="[469,579,554,578]" country="Argentina" name="Tucuman" pageId="10" pageNumber="2223">Tucumán</collectingRegion>
,
<location id="32226013FFC19C240A774415FD0BFDC9" LSID="urn:lsid:plazi:treatment:BF5487DEFFC19C2209114715FD19FEF6:32226013FFC19C240A774415FD0BFDC9" box="[594,735,554,578]" country="Argentina" name="El Siambon" pageId="10" pageNumber="2223" stateProvince="Tucuman">El Siambón</location>
,
<collectorName id="9A08531EFFC19C240ACA4415FC7FFDC9" box="[751,939,554,578]" pageId="10" pageNumber="2223">Río el Siambón</collectorName>
,
<date id="43431008FFC19C240B9F4415FB59FDC9" box="[954,1165,554,578]" pageId="10" pageNumber="2223" value="2008-11-01">
<collectingDate id="5307E9E0FFC19C240B9F4415FB59FDC9" box="[954,1165,554,578]" pageId="10" pageNumber="2223" value="2008-11-01">1 November 2008</collectingDate>
</date>
,
<collectorName id="9A08531EFFC19C2408F04475FEA2FDE9" box="[213,374,586,610]" pageId="10" pageNumber="2223">Rueda Martín</collectorName>
col.:
<specimenCount id="21FBFD41FFC19C2409914474FDCAFDE9" box="[436,542,586,610]" pageId="10" pageNumber="2223" type="male">one male</specimenCount>
</materialsCitation>
;
<materialsCitation id="87953C95FFC19C240A0C4474FBFBFDE9" box="[553,1071,586,610]" collectingDate="1999-10-29" country="Argentina" location="Gonzalo" municipality="Trancas" pageId="10" pageNumber="2223" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman" typeStatus="paratype">
<collectingMunicipality id="D726ACB2FFC19C240A0C4474FD57FDE9" box="[553,643,587,610]" pageId="10" pageNumber="2223">Trancas</collectingMunicipality>
,
<location id="32226013FFC19C240AAA4475FD27FDE9" LSID="urn:lsid:plazi:treatment:BF5487DEFFC19C2209114715FD19FEF6:32226013FFC19C240AAA4475FD27FDE9" box="[655,755,586,610]" country="Argentina" municipality="Trancas" name="Gonzalo" pageId="10" pageNumber="2223" stateProvince="Tucuman">Gonzalo</location>
,
<date id="43431008FFC19C240ADB4475FC68FDE9" box="[766,956,586,610]" pageId="10" pageNumber="2223" value="1999-10-29">
<collectingDate id="5307E9E0FFC19C240ADB4475FC68FDE9" box="[766,956,586,610]" pageId="10" pageNumber="2223" value="1999-10-29">29 October 1999</collectingDate>
</date>
:
<specimenCount id="21FBFD41FFC19C240BE24474FBFBFDE9" box="[967,1071,586,610]" pageId="10" pageNumber="2223" type="male">one male</specimenCount>
</materialsCitation>
.
<materialsCitation id="87953C95FFC19C240C1F4475FC3FFD09" collectorName="Ferbuary &amp; Rueda Martin" country="Bolivia" location="Rio Camacho" pageId="10" pageNumber="2223" specimenCount="1" specimenCount-male="1" stateProvince="Tarija" typeStatus="paratype">
<collectingCountry id="4FEA7658FFC19C240C1F4475FB5AFDE9" box="[1082,1166,586,610]" name="Bolivia" pageId="10" pageNumber="2223">Bolivia</collectingCountry>
:
<collectingRegion id="F539F82AFFC19C2408F04455FECFFD09" box="[213,283,618,642]" country="Bolivia" name="Tarija" pageId="10" pageNumber="2223">Tarija</collectingRegion>
,
<location id="32226013FFC19C2409034455FE12FD09" LSID="urn:lsid:plazi:treatment:BF5487DEFFC19C2209114715FD19FEF6:32226013FFC19C2409034455FE12FD09" box="[294,454,618,642]" country="Bolivia" name="Rio Camacho" pageId="10" pageNumber="2223" stateProvince="Tarija">Río Camacho</location>
, 27
<collectorName id="9A08531EFFC19C2409D74455FD8EFD09" box="[498,602,618,642]" pageId="10" pageNumber="2223">Ferbuary</collectorName>
2008,
<collectorName id="9A08531EFFC19C240A804455FC91FD09" box="[677,837,618,642]" pageId="10" pageNumber="2223">Rueda Martín</collectorName>
col.:
<specimenCount id="21FBFD41FFC19C240BA74455FC3FFD09" box="[898,1003,618,642]" pageId="10" pageNumber="2223" type="male">one male</specimenCount>
</materialsCitation>
;
<materialsCitation id="87953C95FFC19C240BD34455FD2BFD2A" collectingDate="2006-03-06" collectionCode="IML" collectorName="Rueda Martin" country="Bolivia" location="Rio Salinas" pageId="10" pageNumber="2223" specimenCount="1" specimenCount-male="1" stateProvince="Tarija" typeStatus="paratype">
<location id="32226013FFC19C240BD34455FBADFD09" LSID="urn:lsid:plazi:treatment:BF5487DEFFC19C2209114715FD19FEF6:32226013FFC19C240BD34455FBADFD09" box="[1014,1145,618,642]" country="Bolivia" name="Rio Salinas" pageId="10" pageNumber="2223" stateProvince="Tarija">Río Salinas</location>
,
<date id="43431008FFC19C240CA14455FEB4FD2A" pageId="10" pageNumber="2223" value="2006-03-06">
<collectingDate id="5307E9E0FFC19C240CA14455FEB4FD2A" pageId="10" pageNumber="2223" value="2006-03-06">6 March 2006</collectingDate>
</date>
,
<collectorName id="9A08531EFFC19C24094E44B6FDDFFD2A" box="[363,523,649,673]" pageId="10" pageNumber="2223">Rueda Martín</collectorName>
col.:
<specimenCount id="21FBFD41FFC19C240A6D44B5FD79FD2A" box="[584,685,649,673]" pageId="10" pageNumber="2223" type="male">one male</specimenCount>
(
<collectionCode id="51ECAE0DFFC19C240A9944B6FD23FD2A" box="[700,759,649,673]" country="Argentina" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="10" pageNumber="2223">IML</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="374236C8FFC19C2408F044DEFE72FD72" blockId="10.[213,422,737,761]" box="[213,422,737,761]" pageId="10" pageNumber="2223">
<emphasis id="0589EADAFFC19C2408F044DEFE72FD72" box="[213,422,737,761]" italics="true" pageId="10" pageNumber="2223">Species description</emphasis>
</paragraph>
<paragraph id="374236C8FFC19C2408F04536FE6BFC2A" blockId="10.[213,1170,777,992]" pageId="10" pageNumber="2223">
<emphasis id="0589EADAFFC19C2408F04536FEC2FCAA" box="[213,278,777,801]" italics="true" pageId="10" pageNumber="2223">Male.</emphasis>
Length of forewings
<quantity id="F0059B2DFFC19C240A364536FDBAFCAA" box="[531,622,777,801]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="2223" unit="mm" value="4.9">4.9 mm</quantity>
. Colouration of head and thorax light reddish brown, setal warts and abdomen stramineous. Forewing yellowish, with an oval pale macula parallel to the costal margin (pterostigma), and light brown stripes over transversal veins. The colouration of the body and wings was observed in specimens preserved in alcohol.
</paragraph>
<paragraph id="374236C8FFC19C2409254597FC26FC6B" blockId="10.[213,1170,777,992]" pageId="10" pageNumber="2223">Diameter of eye, in dorsal view, 2.3 times the length of the interocular distance. Anterolateral process of sternum V 1.5 times the length of the sternum.</paragraph>
<paragraph id="374236C8FFC19C2408F04237FE8BFA34" blockId="10.[213,1170,1032,1471]" pageId="10" pageNumber="2223">
<emphasis id="0589EADAFFC19C2408F04237FE96FBAB" box="[213,322,1032,1056]" italics="true" pageId="10" pageNumber="2223">Genitalia.</emphasis>
Segment IX with anterolateral margin rounded (
<figureCitation id="AFC62A4DFFC19C240B484237FC33FBAB" box="[877,999,1032,1056]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4A</figureCitation>
). Tergum X, in dorsal view, deeply divided mesally, internal margin of each hemitergite concave with apex slightly curved to the midline bearing a small fold preapically (
<figureCitation id="AFC62A4DFFC19C240BCB4277FBB1FBEB" box="[1006,1125,1096,1120]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4B</figureCitation>
); in lateral view with ventral margin almost straight, apex pointed (
<figureCitation id="AFC62A4DFFC19C240B8E4257FBF1FB0B" box="[939,1061,1128,1152]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4A</figureCitation>
). Inferior appendages two-segmented, setose; basal portion long (2.5 times the length of the apical segment), narrow basally, broad at apex (
<figureCitation id="AFC62A4DFFC19C240AD14298FC53FB34" box="[756,903,1191,1215]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4A,C</figureCitation>
). Phallus long, tubular; basal portion broad, forming an angle of about 90
<superScript id="C0889B80FFC19C240B2642FBFCD8FB5E" attach="left" box="[771,780,1220,1237]" fontSize="7" pageId="10" pageNumber="2223"></superScript>
with the distal part (
<figureCitation id="AFC62A4DFFC19C240BD142F8FB51FB54" box="[1012,1157,1223,1247]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4D,E</figureCitation>
); distal portion with apex produced dorsally, bifid in dorsal view (
<figureCitation id="AFC62A4DFFC19C240BB042D8FBDFFB74" box="[917,1035,1255,1279]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4D</figureCitation>
); ventroapical area of phallus membranous when the endophallus is invaginated (
<figureCitation id="AFC62A4DFFC19C240C284338FB51FA94" box="[1037,1157,1287,1311]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4E</figureCitation>
). Endophallus membranous, tubular, truncated at apex, without spines or processes; when evaginated the tip can be seen through the dorsal process of the phallus as an oval membranous structure, because of the transversal wrinkles that cover its surface. Internal sclerite long and slender, in lateral view slightly curved in the apex (
<figureCitation id="AFC62A4DFFC19C2408F84398FE85FA34" box="[221,337,1447,1471]" captionStart="Figure 4" captionStartId="11.[144,211,1460,1482]" captionTargetBox="[144,1102,161,1425]" captionTargetId="figure-7@11.[144,1102,159,1426]" captionTargetPageId="11" captionText="Figure 4. Smicridea (Rhyacophylax) valeni sp. nov. (A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view. Smicridea (Rhyacophylax) atrobasis. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view. Smicridea (Rhyacophylax) dithyra. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="10" pageNumber="2223">Figure 4E</figureCitation>
).
</paragraph>
<paragraph id="374236C8FFC19C2408F043D8FE78FA75" blockId="10.[213,428,1510,1535]" box="[213,428,1510,1535]" pageId="10" pageNumber="2223">
<emphasis id="0589EADAFFC19C2408F043D8FEFFFA74" box="[213,299,1511,1535]" italics="true" pageId="10" pageNumber="2223">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="374236C8FFC19C2408F04006FE25F9DA" blockId="10.[213,497,1593,1617]" box="[213,497,1593,1617]" pageId="10" pageNumber="2223">
<emphasis id="0589EADAFFC19C2408F04006FE25F9DA" box="[213,497,1593,1617]" italics="true" pageId="10" pageNumber="2223">Systematic considerations</emphasis>
</paragraph>
<paragraph id="374236C8FFC19C2408F0405FFB46F913" blockId="10.[213,1170,1632,1689]" pageId="10" pageNumber="2223">
This species is related to
<taxonomicName id="F0FD4D4BFFC19C2409CF405EFDA3F9F2" authority="Rueda Martín &amp; Sganga, 2011" authorityName="Rueda Martín &amp; Sganga" authorityYear="2011" box="[490,631,1632,1657]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="2223" phylum="Arthropoda" rank="species" species="elisae" status="sp. nov." subGenus="Rhyacophylax">
<emphasis id="0589EADAFFC19C2409CF405EFE2EF9F2" box="[490,506,1633,1657]" italics="true" pageId="10" pageNumber="2223">S</emphasis>
. (
<emphasis id="0589EADAFFC19C240A34405DFDFFF9F2" box="[529,555,1634,1657]" italics="true" pageId="10" pageNumber="2223">R.</emphasis>
)
<emphasis id="0589EADAFFC19C240A1F405EFDA3F9F2" box="[570,631,1633,1657]" italics="true" pageId="10" pageNumber="2223">elisae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EBA57A1FFC19C240A5B405EFD06F9F3" box="[638,722,1633,1656]" pageId="10" pageNumber="2223" rank="species">sp. nov.</taxonomicNameLabel>
Both species share the bifid dorsoapical area of phallus. The
<emphasis id="0589EADAFFC19C2409E240BEFE03F912" box="[455,471,1665,1689]" italics="true" pageId="10" pageNumber="2223">S</emphasis>
.
<taxonomicName id="F0FD4D4BFFC19C2409CC40BFFD8BF912" authorityName="Rueda Martin and Sganga" authorityYear="2011" box="[489,607,1664,1689]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="2223" phylum="Arthropoda" rank="species" species="valeni" subGenus="Rhyacophylax">
(
<emphasis id="0589EADAFFC19C2409D740BEFDD1F913" box="[498,517,1665,1688]" italics="true" pageId="10" pageNumber="2223">R</emphasis>
.)
<emphasis id="0589EADAFFC19C240A3A40BEFD8BF912" box="[543,607,1665,1689]" italics="true" pageId="10" pageNumber="2223">valeni</emphasis>
</taxonomicName>
is distinguished from other species by the tip of
</paragraph>
<caption id="63826640FFC09C2508B5438BFEDCF9F2" ID-DOI="http://doi.org/10.5281/zenodo.5204295" ID-Zenodo-Dep="5204295" httpUri="https://zenodo.org/record/5204295/files/figure.png" pageId="11" pageNumber="2224" startId="11.[144,211,1460,1482]" targetBox="[144,1102,161,1425]" targetPageId="11">
<paragraph id="374236C8FFC09C2508B5438BFEDCF9F2" blockId="11.[144,1101,1460,1657]" pageId="11" pageNumber="2224">
Figure 4.
<taxonomicName id="F0FD4D4BFFC09C2508DF438BFD90FA41" authority="Rueda Martín &amp; Sganga, 2011" authorityName="Rueda Martín &amp; Sganga" authorityYear="2011" box="[250,580,1460,1482]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="2224" phylum="Arthropoda" rank="species" species="valeni" status="sp. nov." subGenus="Rhyacophylax">
<emphasis id="0589EADAFFC09C2508DF438BFE8AFA41" box="[250,350,1460,1482]" italics="true" pageId="11" pageNumber="2224">Smicridea</emphasis>
(
<emphasis id="0589EADAFFC09C25094B438BFE2FFA41" box="[366,507,1460,1482]" italics="true" pageId="11" pageNumber="2224">Rhyacophylax</emphasis>
)
<emphasis id="0589EADAFFC09C250A2E438BFD90FA41" box="[523,580,1460,1482]" italics="true" pageId="11" pageNumber="2224">valeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EBA57A1FFC09C250A68438AFD4FFA41" box="[589,667,1461,1482]" pageId="11" pageNumber="2224" rank="species">sp. nov.</taxonomicNameLabel>
(A) Male genitalia, lateral view; (B) male genitalia, dorsal view; (C) inferior appendages, ventral view; (D) phallus, dorsal view; (E) phallus, lateral view.
<taxonomicName id="F0FD4D4BFFC09C25091D43D1FD74F98F" authorityName=": Flint" authorityYear="1983" box="[312,672,1518,1540]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="2224" phylum="Arthropoda" rank="species" species="atrobasis" subGenus="Rhyacophylax">
<emphasis id="0589EADAFFC09C25091D43D1FE48F98F" box="[312,412,1518,1540]" italics="true" pageId="11" pageNumber="2224">Smicridea</emphasis>
(
<emphasis id="0589EADAFFC09C25098E43D1FDECF98F" box="[427,568,1518,1540]" italics="true" pageId="11" pageNumber="2224">Rhyacophylax</emphasis>
)
<emphasis id="0589EADAFFC09C250A6243D1FD74F98F" box="[583,672,1518,1540]" italics="true" pageId="11" pageNumber="2224">atrobasis</emphasis>
</taxonomicName>
. (F) male genitalia, lateral view; (G) male genitalia, dorsal view; (H) inferior appendages, ventral view; (I) phallus, dorsal view; (J) phallus, lateral view.
<taxonomicName id="F0FD4D4BFFC09C2509354016FDB1F9B4" authorityName=", Flint" authorityYear="1974" box="[272,613,1577,1599]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="2224" phylum="Arthropoda" rank="species" species="dithyra" subGenus="Rhyacophylax">
<emphasis id="0589EADAFFC09C2509354016FEA0F9B4" box="[272,372,1577,1599]" italics="true" pageId="11" pageNumber="2224">Smicridea</emphasis>
(
<emphasis id="0589EADAFFC09C2509A64016FDC4F9B4" box="[387,528,1577,1599]" italics="true" pageId="11" pageNumber="2224">Rhyacophylax</emphasis>
)
<emphasis id="0589EADAFFC09C250A3A4016FDB1F9B4" box="[543,613,1577,1599]" italics="true" pageId="11" pageNumber="2224">dithyra</emphasis>
</taxonomicName>
. (K) male genitalia, lateral view; (L) male genitalia, dorsal view; (M) inferior appendages, ventral view; (N) phallus, lateral view; (O) phallus, dorsal view.
</paragraph>
</caption>
<paragraph id="374236C8FFC79C2208F04699FCA4FF55" blockId="12.[213,1170,166,222]" pageId="12" pageNumber="2225">phallus. The bifid apex and the membranous tubular endophallus truncated at apex are the main characters for the identification of this species.</paragraph>
<paragraph id="374236C8FFC79C2208F04721FE99FEBD" blockId="12.[213,333,286,310]" box="[213,333,286,310]" pageId="12" pageNumber="2225">
<emphasis id="0589EADAFFC79C2208F04721FE99FEBD" box="[213,333,286,310]" italics="true" pageId="12" pageNumber="2225">Etymology</emphasis>
</paragraph>
<paragraph id="374236C8FFC79C2208F0477AFD19FEF6" blockId="12.[213,1169,325,382]" pageId="12" pageNumber="2225">
This species is dedicated to Valentino, son of one of the authors (JVS). The name
<taxonomicName id="F0FD4D4BFFC79C2208F04759FEC1FEF5" authorityName="Rueda Martín &amp; Sganga" authorityYear="2011" box="[213,277,358,382]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="2225" phylum="Arthropoda" rank="species" species="valeni">
<emphasis id="0589EADAFFC79C2208F04759FEC1FEF5" box="[213,277,358,382]" italics="true" pageId="12" pageNumber="2225">valeni</emphasis>
</taxonomicName>
is derived from his nickname, “Valen”.
</paragraph>
</subSubSection>
</treatment>
</document>