288 lines
37 KiB
XML
288 lines
37 KiB
XML
<document id="E6E8AFF72F0CB8B942639B2AB141292B" ID-DOI="10.5281/zenodo.5354295" ID-ISSN="0374-1036" ID-Zenodo-Dep="5354295" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630213702006" checkinUser="felipe" docAuthor="Halstead, David G. H." docDate="2012" docId="BB492919FFAB4C0AFE18DD6CFBF9A8A1" docLanguage="en" docName="ActaEntMusNatPra.2012.52.suppl.223-231.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Oryzaephilus socotraensis Halstead 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="227" masterDocId="47705161FFAA4C0EFF84D877FFE0A915" masterDocTitle="New species of Oryzaephilus and Silvanolomus from Socotra Island (Coleoptera: Silvanidae: Silvaninae)" masterLastPageNumber="231" masterPageNumber="223" pageNumber="224" updateTime="1699188462142" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="204D4CEB94DE340FC69B3A248792DA28" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0D5F45A431F025E51436121A54040471">
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<mods:title id="A0F74F1FA1ED939300C4C2E35E3E3475">New species of Oryzaephilus and Silvanolomus from Socotra Island (Coleoptera: Silvanidae: Silvaninae)</mods:title>
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</mods:titleInfo>
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<mods:name id="D9D63136F746772807A6FE4A09F5A289" type="personal">
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<mods:role id="29B36C1DCFF6564D60607CC38A6E4BD2">
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<mods:roleTerm id="004906C04659340C14C112AAEE69E7B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FA599F71FF9CC13B5C7BE4AA14ACD7A1">Halstead, David G. H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="79F0D48B809BA58CDD053A0634934955">text</mods:typeOfResource>
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<mods:relatedItem id="1A505F17812B3988ADE3B3F3E945E60B" type="host">
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<mods:titleInfo id="874EC5882D1D0FF22AB2C142198C932A">
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<mods:title id="BD0B611221883B56AD89FDECA966FB56">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="9A558C4C7C2852CD458ACE609DC7D10D">
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<mods:date id="FD18360581C7D7295EC6EE973E2FFE29">2012</mods:date>
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<mods:detail id="55A5BF8C381AC01719CBDD45CAFE3DF6" type="pubDate">
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<mods:number id="BAC501D4CE7DA6FBC26803B992B7F1AB">2012-12-17</mods:number>
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</mods:detail>
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<mods:detail id="0DD49122D86E3B756E963B428B91597A" type="volume">
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<mods:number id="AB57DB482BC66BF04F95D9221A08BE8E">52</mods:number>
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</mods:detail>
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<mods:extent id="79D3888CABFAF29FA894BA665857849D" unit="page">
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<mods:start id="E5D59BB86AFAC5B6AF88E13D06068BF5">223</mods:start>
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<mods:end id="7F6429FE4F623FEED5FF158E41EAC714">231</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="7A6827B0DF1EB47B191AA2EB54586948">journal article</mods:classification>
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<mods:identifier id="48A3B2E8E4BC880D00F6BE3A1A5E9DE1" type="DOI">10.5281/zenodo.5354295</mods:identifier>
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<mods:identifier id="E15279A664FED694E9275591163949E5" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="0FA3550BC16F23CA05270D2CC472F45F" type="Zenodo-Dep">5354295</mods:identifier>
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</mods:mods>
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<treatment id="BB492919FFAB4C0AFE18DD6CFBF9A8A1" LSID="urn:lsid:plazi:treatment:BB492919FFAB4C0AFE18DD6CFBF9A8A1" httpUri="http://treatment.plazi.org/id/BB492919FFAB4C0AFE18DD6CFBF9A8A1" lastPageId="4" lastPageNumber="227" pageId="1" pageNumber="224">
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<subSubSection id="7BFACB84FFAB4C0FFE18DD6CFCF7AC21" box="[412,791,1307,1332]" pageId="1" pageNumber="224" type="nomenclature">
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<paragraph id="335F980FFFAB4C0FFE18DD6CFCF7AC21" blockId="1.[412,791,1307,1332]" box="[412,791,1307,1332]" pageId="1" pageNumber="224">
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<heading id="68172F63FFAB4C0FFE18DD6CFCF7AC21" bold="true" box="[412,791,1307,1332]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="224" reason="3">
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||
<emphasis id="0194441DFFAB4C0FFE18DD6CFCF7AC21" bold="true" box="[412,791,1307,1332]" pageId="1" pageNumber="224">
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||
<taxonomicName id="F4E0E38CFFAB4C0FFE18DD6CFD5BAC21" authority="Halstead, 2012" authorityName="Halstead" authorityYear="2012" box="[412,699,1307,1332]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="socotraensis" status="sp. nov.">
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<emphasis id="0194441DFFAB4C0FFE18DD6CFD5BAC21" bold="true" box="[412,699,1307,1332]" italics="true" pageId="1" pageNumber="224">Oryzaephilus socotraensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1AA7F966FFAB4C0FFD45DD6BFCF7AC21" box="[705,791,1308,1332]" pageId="1" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="7BFACB84FFAB4C0FFDA6DD4DFD6FAC58" box="[546,655,1338,1357]" pageId="1" pageNumber="224" type="description">
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||
<paragraph id="335F980FFFAB4C0FFDA6DD4DFD6FAC58" blockId="1.[546,655,1338,1357]" box="[546,655,1338,1357]" pageId="1" pageNumber="224">
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||
(
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<figureCitation id="ABDB848AFFAB4C0FFDADDD4DFD80AC58" box="[553,608,1338,1357]" captionStart="Figs" captionStartId="2.[151,190,1094,1113]" captionTargetBox="[204,1077,227,1070]" captionTargetId="figure-165@2.[197,1080,219,1073]" captionTargetPageId="2" captionText="Figs.1–2. Habitus, dorsal view. 1 – Oryzaephilus socotraensis sp. nov. (paratype, male); 2 – Silvanolomus depressus sp. nov. (holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.5354297" httpUri="https://zenodo.org/record/5354297/files/figure.png" pageId="1" pageNumber="224">Figs.1</figureCitation>
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,
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<figureCitation id="ABDB848AFFAB4C0FFDEDDD4DFD6AAC58" box="[617,650,1338,1357]" captionStart="Figs" captionStartId="4.[151,190,1520,1539]" captionTargetBox="[170,1110,465,1498]" captionTargetId="figure-12@4.[167,1111,461,1504]" captionTargetPageId="4" captionText="Figs. 3–7. Oryzaephilus spp. 3–6 – O. socotraensis sp. nov.: 3 – head, pronotum and elytra of male; 4 – head and pronotum of female; 5–6 – male genitalia, 5 – internal sac, 6 – genitalia without sac. 7 – O. gibbosus Aitken, 1965, median lobe and parameres of male genitalia (after HALSTEAD 1980). Scale bars = 1 mm (Figs. 3–4), 0.1 mm (Figs. 5–7)." figureDoi="http://doi.org/10.5281/zenodo.5354299" httpUri="https://zenodo.org/record/5354299/files/figure.png" pageId="1" pageNumber="224">3–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="7BFACB84FFAB4C0FFFF5DD12FCD6AC62" box="[113,822,1380,1399]" pageId="1" pageNumber="224" type="reference_group">
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<paragraph id="335F980FFFAB4C0FFFF5DD12FCD6AC62" blockId="1.[113,822,1380,1399]" box="[113,822,1380,1399]" pageId="1" pageNumber="224">
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<treatmentCitationGroup id="13F0BF21FFAB4C0FFFF5DD12FCD6AC62" box="[113,822,1380,1399]" pageId="1" pageNumber="224">
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<taxonomicName id="F4E0E38CFFAB4C0FFFF5DD12FEFAAC62" authorityName="Halstead" authorityYear="1980" box="[113,282,1381,1399]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="canus">
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<emphasis id="0194441DFFAB4C0FFFF5DD12FEFAAC62" box="[113,282,1381,1399]" italics="true" pageId="1" pageNumber="224">Oryzaephilus canus</emphasis>
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</taxonomicName>
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non
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<bibRefCitation id="5771E5FEFFAB4C0FFEC1DD13FE28AC62" author="HALSTEAD D. G. H." box="[325,456,1380,1399]" pageId="1" pageNumber="224" pagination="271 - 374" refId="ref3804" refString="HALSTEAD D. G. H. 1980: A revision of the genus Oryzaephilus Ganglbauer, including descriptions of related genera (Coleoptera: Silvanidae). Zoological Journal of the Linnaean Society 69: 271 - 374." type="journal article" year="1980">Halstead, 1980</bibRefCitation>
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:
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<treatmentCitation id="B241BE1EFFAB4C0FFE54DD13FD6CAC62" author="HALSTEAD D. G. H." box="[464,652,1380,1399]" page="234" pageId="1" pageNumber="224" year="2011">
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<bibRefCitation id="5771E5FEFFAB4C0FFE54DD13FD84AC62" author="HALSTEAD D. G. H." box="[464,612,1380,1399]" pageId="1" pageNumber="224" pagination="233 - 245" refId="ref3917" refString="HALSTEAD D. G. H. 2011: Order Coleoptera, family Silvanidae. Pp. 233 - 245. In: HARTEN A. VAN (ed.): Arthropod Fauna of the United Arab Emirates. Volume 4. Multiply Marketing Consultancy Services, Abu Dhabi, 832 pp." type="book chapter" year="2011">HALSTEAD (2011)</bibRefCitation>
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: 234
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</treatmentCitation>
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(misidentification).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="7BFACB84FFAB4C0FFFF5DDF9FBDCAF4E" pageId="1" pageNumber="224" type="materials_examined">
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||
<paragraph id="335F980FFFAB4C0FFFF5DDF9FC0FACB4" blockId="1.[113,1090,1421,1628]" box="[113,1007,1421,1441]" pageId="1" pageNumber="224">
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<emphasis id="0194441DFFAB4C0FFFF5DDF9FF0AACB4" bold="true" box="[113,234,1421,1441]" pageId="1" pageNumber="224">
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||
<typeStatus id="EC5B26ADFFAB4C0FFFF5DDF9FF7EACB4" box="[113,158,1422,1441]" pageId="1" pageNumber="224">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="4BF7D89FFFAB4C0FFF6ADDF9FECCACB4" box="[238,300,1422,1441]" name="Yemen" pageId="1" pageNumber="224">Yemen</collectingCountry>
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,
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<collectingCountry id="4BF7D89FFFAB4C0FFEB1DDF9FE51ACB4" box="[309,433,1422,1441]" name="Socotra Island" pageId="1" pageNumber="224">Socotra Island</collectingCountry>
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, Noged plain, Sharet Halma vill. env.,
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<geoCoordinate id="56D4FEC8FFAB4C0FFD78DDF9FCB5ACB4" box="[764,853,1421,1441]" degrees="12" direction="north" minutes="21.9" orientation="latitude" pageId="1" pageNumber="224" precision="92" value="12.365">12°21.9′N</geoCoordinate>
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,
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<geoCoordinate id="56D4FEC8FFAB4C0FFCDADDF9FC55ACB4" box="[862,949,1421,1441]" degrees="54" direction="east" minutes="05.3" orientation="longitude" pageId="1" pageNumber="224" precision="92" value="54.088333">54°05.3′E</geoCoordinate>
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,
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<specimenCount id="25E65386FFAB4C0FFC39DDF9FC0FACB4" box="[957,1007,1422,1441]" pageId="1" pageNumber="224" type="generic">
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||
<quantity id="F41835EAFFAB4C0FFC39DDF9FC0BACB4" box="[957,1003,1422,1441]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="224" unit="m" value="20.0">20 m</quantity>
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.
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</specimenCount>
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</paragraph>
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<paragraph id="335F980FFFAB4C0FFFF5DDDEFC89ACE4" blockId="1.[113,1090,1421,1628]" pageId="1" pageNumber="224">
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||
<emphasis id="0194441DFFAB4C0FFFF5DDDEFF15ACA9" bold="true" box="[113,245,1448,1468]" pageId="1" pageNumber="224">Type material.</emphasis>
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||
<materialsCitation id="83889252FFAB4C0FFF7EDDDEFCE6ACC3" collectingDate="2010-11-10" collectingDateMax="2010-11-11" collectingDateMin="2010-11-10" collectionCode="NMPC, SHARET, HALMA" collectorName="J. Bezdek" country="Yemen" elevation="20" latitude="12.365" location="Island" longLatPrecision="129" longitude="54.088333" municipality="Noged" pageId="1" pageNumber="224" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="EC5B26ADFFAB4C0FFF7EDDDEFEADACAF" box="[250,333,1449,1468]" pageId="1" pageNumber="224" type="holotype">HOLOTYPE</typeStatus>
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||
: dissected J, (
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<collectionCode id="55F100CAFFAB4C0FFE4EDDDEFDECACA9" box="[458,524,1449,1468]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="224">NMPC</collectionCode>
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||
): ‘
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<collectingCountry id="4BF7D89FFFAB4C0FFD98DDDEFD8EACA9" box="[540,622,1449,1468]" name="Yemen" pageId="1" pageNumber="224">YEMEN</collectingCountry>
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, SOCOTRA
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<location id="363FCED4FFAB4C0FFD59DDDFFCF1ACAE" LSID="urn:lsid:plazi:treatment:BB492919FFAB4C0AFE18DD6CFBF9A8A1:363FCED4FFAB4C0FFD59DDDFFCF1ACAE" box="[733,785,1448,1467]" country="Yemen" latitude="12.365" longLatPrecision="129" longitude="54.088333" municipality="Noged" name="Island" pageId="1" pageNumber="224">Island</location>
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/
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<collectingMunicipality id="D33B0275FFAB4C0FFC9FDDDFFCB4ACAE" box="[795,852,1448,1467]" pageId="1" pageNumber="224">Noged</collectingMunicipality>
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plain (sand dunes) /
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<collectionCode id="55F100CAFFAB4C0FFB87DDDEFF4DACC0" pageId="1" pageNumber="224">SHARET HALMA</collectionCode>
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vill. env. /
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<geoCoordinate id="56D4FEC8FFAB4C0FFE89DDB4FE86ACC2" box="[269,358,1474,1495]" degrees="12" direction="north" minutes="21.9" orientation="latitude" pageId="1" pageNumber="224" precision="92" value="12.365">12°21.9′N</geoCoordinate>
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,
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<geoCoordinate id="56D4FEC8FFAB4C0FFEEBDDB4FE26ACC2" box="[367,454,1474,1495]" degrees="54" direction="east" minutes="05.3" orientation="longitude" pageId="1" pageNumber="224" precision="92" value="54.088333">54°05.3′E</geoCoordinate>
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,
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<quantity id="F41835EAFFAB4C0FFE4BDDB4FE15ACC3" box="[463,501,1475,1494]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="224" unit="m" value="20.0">
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||
<elevation id="B8CD7F3CFFAB4C0FFE4BDDB4FE15ACC3" box="[463,501,1475,1494]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="224" unit="m" value="20.0">20m</elevation>
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</quantity>
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/
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<collectorName id="9E15FDD9FFAB4C0FFD82DDB3FDBAACC3" box="[518,602,1475,1495]" pageId="1" pageNumber="224">J. Bezděk</collectorName>
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||
leg.,
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||
<date id="475EBECFFFAB4C0FFD0DDDB4FCE6ACC3" box="[649,774,1475,1494]" pageId="1" pageNumber="224" value="2010-11-10" valueMax="2010-11-11" valueMin="2010-11-10">
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||
<collectingDate id="571A4727FFAB4C0FFD0DDDB4FCE6ACC3" box="[649,774,1475,1494]" pageId="1" pageNumber="224" value="2010-11-10" valueMax="2010-11-11" valueMin="2010-11-10">10-11.xi.2010</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
’.
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<typeStatus id="EC5B26ADFFAB4C0FFC96DDB3FC8AACC0" box="[786,874,1476,1495]" pageId="1" pageNumber="224" type="paratype">PARATYPES</typeStatus>
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||
: 4 JJ
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<specimenCount id="25E65386FFAB4C0FFC2EDDB4FC29ACC3" box="[938,969,1469,1494]" pageId="1" pageNumber="224" type="female">1 ♀</specimenCount>
|
||
, same data as
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<typeStatus id="EC5B26ADFFAB4C0FFFF5DDA9FF5CACE4" box="[113,188,1502,1521]" pageId="1" pageNumber="224" type="holotype">holotype</typeStatus>
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||
but JJ collected by Jiří Hájek and
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<specimenCount id="25E65386FFAB4C0FFE75DDAFFDE1ACE4" box="[497,513,1496,1521]" pageId="1" pageNumber="224" type="female">♀</specimenCount>
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by Luboš Purchart (
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||
<collectionCode id="55F100CAFFAB4C0FFD2ADDA9FD0EACE4" box="[686,750,1502,1521]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="224">NMPC</collectionCode>
|
||
, 1 J
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||
<collectionCode id="55F100CAFFAB4C0FFC98DDA9FC80ACE4" box="[796,864,1502,1521]" pageId="1" pageNumber="224">BMNH</collectionCode>
|
||
).
|
||
</paragraph>
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||
<paragraph id="335F980FFFAB4C0FFFF5DD8FFBDCAF4E" blockId="1.[113,1090,1421,1628]" pageId="1" pageNumber="224">
|
||
<emphasis id="0194441DFFAB4C0FFFF5DD8FFE65AF19" bold="true" box="[113,389,1528,1548]" pageId="1" pageNumber="224">Additional material examined.</emphasis>
|
||
1 dissected J: ‘
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||
<collectingCountry id="4BF7D89FFFAB4C0FFD82DD8EFDB8AF19" box="[518,600,1529,1548]" name="Yemen" pageId="1" pageNumber="224">YEMEN</collectingCountry>
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, SOQOTRA-ARCHIPEL, SOQOTRA/ Noged.Farmihin, Nähe Strand, Om./ 12˚ 24′ 41″ N, 54˚ 13′ 35″ E, 24.-
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<date id="475EBECFFFAB4C0FFDB7DE64FD77AF33" box="[563,663,1555,1574]" pageId="1" pageNumber="224" value="2000-10-25">25.10.2000</date>
|
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/ leg.: H. Pohl, SOQ 2000/04 // HLMD-Col-934’. Although this specimen has all the characters of the species and falls within the measurements given, it has not been included in the
|
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<typeStatus id="EC5B26ADFFAB4C0FFF72DE3EFEFBAF49" box="[246,283,1609,1628]" pageId="1" pageNumber="224">type</typeStatus>
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||
series because it is badly damaged, particularly its pronotum, which is split down the middle.
|
||
</paragraph>
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||
</subSubSection>
|
||
<caption id="679FC887FFA84C0CFF13DC31FE92AD66" ID-DOI="http://doi.org/10.5281/zenodo.5354297" ID-Zenodo-Dep="5354297" httpUri="https://zenodo.org/record/5354297/files/figure.png" pageId="2" pageNumber="225" startId="2.[151,190,1094,1113]" targetBox="[204,1077,227,1070]" targetPageId="2">
|
||
<paragraph id="335F980FFFA84C0CFF13DC31FE92AD66" blockId="2.[151,1126,1094,1140]" pageId="2" pageNumber="225">
|
||
Figs.1–2. Habitus, dorsal view. 1 –
|
||
<taxonomicName id="F4E0E38CFFA84C0CFE45DC30FD42AD4C" authority="Halstead, 2012" authorityName="Halstead" authorityYear="2012" box="[449,674,1095,1113]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="socotraensis" status="sp. nov.">
|
||
<emphasis id="0194441DFFA84C0CFE45DC30FD42AD4C" box="[449,674,1095,1113]" italics="true" pageId="2" pageNumber="225">Oryzaephilus socotraensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA7F966FFA84C0CFD22DC31FD07AD4C" box="[678,743,1094,1113]" pageId="2" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(paratype, male); 2 –
|
||
<taxonomicName id="F4E0E38CFFA84C0CFC18DC30FB86AD4C" authority="Halstead, 2012" authorityName="Halstead" authorityYear="2012" box="[924,1126,1095,1113]" class="Insecta" family="Silvanidae" genus="Silvanolomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="depressus" status="sp. nov.">
|
||
<emphasis id="0194441DFFA84C0CFC18DC30FB86AD4C" box="[924,1126,1095,1113]" italics="true" pageId="2" pageNumber="225">Silvanolomus depressus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA7F966FFA84C0CFF13DC16FF3AAD61" box="[151,218,1121,1140]" pageId="2" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(holotype, male).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7BFACB84FFA84C0DFF13DC93FE12AA86" lastPageId="3" lastPageNumber="226" pageId="2" pageNumber="225" type="description">
|
||
<paragraph id="335F980FFFA84C0CFF13DC93FE8EAC49" blockId="2.[151,1127,1252,1628]" pageId="2" pageNumber="225">
|
||
<emphasis id="0194441DFFA84C0CFF13DC93FEC2ADE9" bold="true" box="[151,290,1252,1276]" pageId="2" pageNumber="225">Description.</emphasis>
|
||
Dark brown to blackish-brown, moderately depressed and elongate (as usual for the genus). Length
|
||
<quantity id="F41835EAFFA84C0CFE15DD73FDF2AC0E" box="[401,530,1284,1308]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="2" pageNumber="225" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
|
||
(
|
||
<typeStatus id="EC5B26ADFFA84C0CFDA6DD73FD65AC09" box="[546,645,1284,1308]" pageId="2" pageNumber="225" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="F41835EAFFA84C0CFD14DD73FD08AC09" box="[656,744,1284,1308]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="225" unit="mm" value="2.4">2.4 mm</quantity>
|
||
); length: maximum elytral breadth ratio, 32.4–38.3: 10. Pubescence golden or greyish. Head and pronotum appearing dull, elytra more shining.
|
||
</paragraph>
|
||
<paragraph id="335F980FFFA84C0CFF31DD13FC22AF4E" blockId="2.[151,1127,1252,1628]" pageId="2" pageNumber="225">
|
||
Head. As long or slightly longer than breadth across temples; genae (sides of head in front of eyes) obviously raised forming conspicuous rim (as in many other
|
||
<taxonomicName id="F4E0E38CFFA84C0CFC19DDF2FBCBAC89" authorityName="Ganglbauer" authorityYear="1899" box="[925,1067,1413,1436]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0194441DFFA84C0CFC19DDF2FBCBAC89" box="[925,1067,1413,1436]" italics="true" pageId="2" pageNumber="225">Oryzaephilus</emphasis>
|
||
</taxonomicName>
|
||
), not sharply angled and without horns; small depressions above antennal insertions; eyes large, separated medially across head by about ×5–×7 breadth, separated from front of head by about ×1.5 length; temple length: eye length ratio, 10: 50–60; antennal length: body length ratio, males 10: 31.5–33.2, female 10: 35; head of usual size compared with pronotum (breadth across temples: pronotal maximum breadth, excluding anterior angles and other teeth, males 10: 11.3–11.5, female 10: 12.3); puncturation strongly reticulate on vertex.
|
||
</paragraph>
|
||
<paragraph id="335F980FFFA94C0DFF14D8ABFD8EA8A6" blockId="3.[113,1089,219,1619]" pageId="3" pageNumber="226">
|
||
Pronotum. Moderately elongate, length: maximum breadth (excluding anterior angles and other teeth) ratio, males 13.2–14.0: 10, female 13.0: 10 (probably tending to be less elongate in all females); lateral ridges well developed, somewhat curved in female, more or less parallel for greater length in males, median ridge higher than lateral ridges; anterior angles strongly produced forming very prominent laterally directed teeth (slightly more so in males than in female seen), see
|
||
<figureCitation id="ABDB848AFFA94C0DFE06D90CFE07A881" box="[386,487,379,404]" captionStart="Figs" captionStartId="4.[151,190,1520,1539]" captionTargetBox="[170,1110,465,1498]" captionTargetId="figure-12@4.[167,1111,461,1504]" captionTargetPageId="4" captionText="Figs. 3–7. Oryzaephilus spp. 3–6 – O. socotraensis sp. nov.: 3 – head, pronotum and elytra of male; 4 – head and pronotum of female; 5–6 – male genitalia, 5 – internal sac, 6 – genitalia without sac. 7 – O. gibbosus Aitken, 1965, median lobe and parameres of male genitalia (after HALSTEAD 1980). Scale bars = 1 mm (Figs. 3–4), 0.1 mm (Figs. 5–7)." figureDoi="http://doi.org/10.5281/zenodo.5354299" httpUri="https://zenodo.org/record/5354299/files/figure.png" pageId="3" pageNumber="226">Figs. 3–4</figureCitation>
|
||
, other teeth moderately developed; puncturation coarse, reticulate as on head, a few punctures reniform.
|
||
</paragraph>
|
||
<paragraph id="335F980FFFA94C0DFF14D9CCFD35AB01" blockId="3.[113,1089,219,1619]" pageId="3" pageNumber="226">Elytra. Length: maximum breadth ratio, 18.7–22.6: 10; sides gradually curved to apex (apices curved to suture, not produced before it); third and alternate interstriae with setae arranged in three rows (usual herring-bone arrangement).</paragraph>
|
||
<paragraph id="335F980FFFA94C0DFF14DA6CFEA2AB61" blockId="3.[113,1089,219,1619]" pageId="3" pageNumber="226">
|
||
Metathoracic legs. Male, metatrochanter with spine, metatibia with minute spine near apex (metafemur without spine); female, without secondary sexual characters (as in other
|
||
<taxonomicName id="F4E0E38CFFA94C0DFFF5DA2AFED6AB61" box="[113,310,604,628]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="0194441DFFA94C0DFFF5DA2AFF1FAB61" box="[113,255,605,628]" italics="true" pageId="3" pageNumber="226">Oryzaephilus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="335F980FFFA94C0DFF14DA0CFE12AA86" blockId="3.[113,1089,219,1619]" pageId="3" pageNumber="226">
|
||
Male genitalia. (
|
||
<figureCitation id="ABDB848AFFA94C0DFEC5DA0CFE49AB86" box="[321,425,635,659]" captionStart="Figs" captionStartId="4.[151,190,1520,1539]" captionTargetBox="[170,1110,465,1498]" captionTargetId="figure-12@4.[167,1111,461,1504]" captionTargetPageId="4" captionText="Figs. 3–7. Oryzaephilus spp. 3–6 – O. socotraensis sp. nov.: 3 – head, pronotum and elytra of male; 4 – head and pronotum of female; 5–6 – male genitalia, 5 – internal sac, 6 – genitalia without sac. 7 – O. gibbosus Aitken, 1965, median lobe and parameres of male genitalia (after HALSTEAD 1980). Scale bars = 1 mm (Figs. 3–4), 0.1 mm (Figs. 5–7)." figureDoi="http://doi.org/10.5281/zenodo.5354299" httpUri="https://zenodo.org/record/5354299/files/figure.png" pageId="3" pageNumber="226">Figs. 5–6</figureCitation>
|
||
). Internal sac with armature (
|
||
<figureCitation id="ABDB848AFFA94C0DFD6CDA0CFCCAAB86" box="[744,810,635,659]" captionStart="Figs" captionStartId="4.[151,190,1520,1539]" captionTargetBox="[170,1110,465,1498]" captionTargetId="figure-12@4.[167,1111,461,1504]" captionTargetPageId="4" captionText="Figs. 3–7. Oryzaephilus spp. 3–6 – O. socotraensis sp. nov.: 3 – head, pronotum and elytra of male; 4 – head and pronotum of female; 5–6 – male genitalia, 5 – internal sac, 6 – genitalia without sac. 7 – O. gibbosus Aitken, 1965, median lobe and parameres of male genitalia (after HALSTEAD 1980). Scale bars = 1 mm (Figs. 3–4), 0.1 mm (Figs. 5–7)." figureDoi="http://doi.org/10.5281/zenodo.5354299" httpUri="https://zenodo.org/record/5354299/files/figure.png" pageId="3" pageNumber="226">Fig. 5</figureCitation>
|
||
, somewhat fragmented in specimen drawn); 11–13 rods on each side towards ostium; median strut strongly narrowed to basal half; median lobe with a few short setae on basal third, sinuate before broad, rounded apex; parameres elongate, more or less parallel sided, slightly curved to base from apical half, apices truncate to slightly rounded to outer margin, bearing four long apically forked setae plus two or three thinner, shorter simple setae towards outer margin, outer and inner margins below the apices with several short inconspicuous setae (
|
||
<figureCitation id="ABDB848AFFA94C0DFCB6DB4CFC95AA46" box="[818,885,827,851]" captionStart="Figs" captionStartId="4.[151,190,1520,1539]" captionTargetBox="[170,1110,465,1498]" captionTargetId="figure-12@4.[167,1111,461,1504]" captionTargetPageId="4" captionText="Figs. 3–7. Oryzaephilus spp. 3–6 – O. socotraensis sp. nov.: 3 – head, pronotum and elytra of male; 4 – head and pronotum of female; 5–6 – male genitalia, 5 – internal sac, 6 – genitalia without sac. 7 – O. gibbosus Aitken, 1965, median lobe and parameres of male genitalia (after HALSTEAD 1980). Scale bars = 1 mm (Figs. 3–4), 0.1 mm (Figs. 5–7)." figureDoi="http://doi.org/10.5281/zenodo.5354299" httpUri="https://zenodo.org/record/5354299/files/figure.png" pageId="3" pageNumber="226">Fig. 6</figureCitation>
|
||
); sternites VIII–IX (not illustrated), sternite VIII with three longer setae present on outer half of margin of each side, finer very short setae medially.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFACB84FFA94C0AFFF5DBECFD40A841" lastPageId="4" lastPageNumber="227" pageId="3" pageNumber="226" type="diagnosis">
|
||
<paragraph id="335F980FFFA94C0DFFF5DBECFE45AD66" blockId="3.[113,1089,219,1619]" pageId="3" pageNumber="226">
|
||
<emphasis id="0194441DFFA94C0DFFF5DBECFE90AAA6" bold="true" box="[113,368,923,947]" pageId="3" pageNumber="226">Differential diagnosis.</emphasis>
|
||
The following combination of characters distinguishes this from other known species: Eyes large and prominent; temples short, about a fifth or less as long as eye; anterior angles of pronotum strongly produced laterally to form a prominent, narrow tooth, obviously more strongly developed than all other teeth; pronotum not gibbous; male genitalia: internal sac with armature; median lobe without a ventral tooth, sides sinuate before broad apex; parameres elongate more or less parallel-sided, only a few short setae along outer margin (no long setae there).
|
||
</paragraph>
|
||
<paragraph id="335F980FFFA94C0DFF14DC0BFC06AC46" blockId="3.[113,1089,219,1619]" pageId="3" pageNumber="226">
|
||
A strongly developed tooth at the anterior pronotal angle, more prominent than the other teeth especially in larger males, is also present in
|
||
<taxonomicName id="F4E0E38CFFA94C0DFDB5DCEAFD4CADA1" authorityName="Aitken" authorityYear="1965" box="[561,684,1181,1204]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="gibbosus">
|
||
<emphasis id="0194441DFFA94C0DFDB5DCEAFD4CADA1" box="[561,684,1181,1204]" italics="true" pageId="3" pageNumber="226">O. gibbosus</emphasis>
|
||
</taxonomicName>
|
||
, an African species that has been found in
|
||
<collectingCountry id="4BF7D89FFFA94C0DFF0FDCCBFF3BADC1" box="[139,219,1212,1236]" name="Yemen" pageId="3" pageNumber="226">Socotra</collectingCountry>
|
||
(see below), and
|
||
<taxonomicName id="F4E0E38CFFA94C0DFE0BDCCAFD29ADC1" authority="Halstead, 1980" authorityName="Halstead" authorityYear="1980" box="[399,713,1211,1236]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="acuminatus">
|
||
<emphasis id="0194441DFFA94C0DFE0BDCCAFDC6ADC1" box="[399,550,1213,1236]" italics="true" pageId="3" pageNumber="226">O. acuminatus</emphasis>
|
||
<bibRefCitation id="5771E5FEFFA94C0DFDAFDCCCFD29ADC1" author="HALSTEAD D. G. H." box="[555,713,1211,1236]" pageId="3" pageNumber="226" pagination="271 - 374" refId="ref3804" refString="HALSTEAD D. G. H. 1980: A revision of the genus Oryzaephilus Ganglbauer, including descriptions of related genera (Coleoptera: Silvanidae). Zoological Journal of the Linnaean Society 69: 271 - 374." type="journal article" year="1980">Halstead, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
, an Oriental species. However, apart from having obviously different male genitalia (parameres and median lobe are quite different) the elytra of these two species appear more elongate, more parallel sided and at their apices usually have the margin slightly produced before meeting the suture, also the pronota of these species are to some extent gibbous, inconspicuously to (in large males) obviously so.
|
||
</paragraph>
|
||
<paragraph id="335F980FFFA94C0AFF14DD2CFD40A841" blockId="3.[113,1089,219,1619]" lastBlockId="4.[151,1127,220,436]" lastPageId="4" lastPageNumber="227" pageId="3" pageNumber="226">
|
||
Males of
|
||
<taxonomicName id="F4E0E38CFFA94C0DFF75DD2AFD84AC61" authority="Halstead, 1980" authorityName="Halstead" authorityYear="1980" box="[241,612,1371,1396]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="canus">
|
||
<emphasis id="0194441DFFA94C0DFF75DD2AFE21AC61" box="[241,449,1373,1396]" italics="true" pageId="3" pageNumber="226">Oryzaephilus canus</emphasis>
|
||
<bibRefCitation id="5771E5FEFFA94C0DFE42DD2CFD84AC61" author="HALSTEAD D. G. H." box="[454,612,1371,1396]" pageId="3" pageNumber="226" pagination="271 - 374" refId="ref3804" refString="HALSTEAD D. G. H. 1980: A revision of the genus Oryzaephilus Ganglbauer, including descriptions of related genera (Coleoptera: Silvanidae). Zoological Journal of the Linnaean Society 69: 271 - 374." type="journal article" year="1980">Halstead, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
and the new species have the same secondary sexual characters of the hind legs; in both the metafemora are without a spine. Also their genitalia are similar although in
|
||
<taxonomicName id="F4E0E38CFFA94C0DFE55DDEAFD98ACA1" authority="Halstead, 2012" authorityName="Halstead" authorityYear="2012" box="[465,632,1437,1460]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="socotraensis" status="sp. nov.">
|
||
<emphasis id="0194441DFFA94C0DFE55DDEAFD98ACA1" box="[465,632,1437,1460]" italics="true" pageId="3" pageNumber="226">O. socotraensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA7F966FFA94C0DFD04DDEBFD34ACA6" box="[640,724,1436,1459]" pageId="3" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the parameres, apart from having long setae limited to the apices, are more elongate. In
|
||
<taxonomicName id="F4E0E38CFFA94C0DFD39DDCAFCFCACC1" authorityName="Halstead" authorityYear="1980" box="[701,796,1469,1492]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="canus">
|
||
<emphasis id="0194441DFFA94C0DFD39DDCAFCFCACC1" box="[701,796,1469,1492]" italics="true" pageId="3" pageNumber="226">O. canus</emphasis>
|
||
</taxonomicName>
|
||
long setae are also present along the apical half of the outer margin of the parameres (see
|
||
<bibRefCitation id="5771E5FEFFA94C0DFD76DDABFC73ACE1" author="HALSTEAD D. G. H." box="[754,915,1500,1524]" pageId="3" pageNumber="226" pagination="271 - 374" refId="ref3804" refString="HALSTEAD D. G. H. 1980: A revision of the genus Oryzaephilus Ganglbauer, including descriptions of related genera (Coleoptera: Silvanidae). Zoological Journal of the Linnaean Society 69: 271 - 374." type="journal article" year="1980">HALSTEAD 1980</bibRefCitation>
|
||
for figures). The damaged J specimen of
|
||
<taxonomicName id="F4E0E38CFFA94C0DFEF2DD8AFDFAAF01" authority="Halstead, 2012" authorityName="Halstead" authorityYear="2012" box="[374,538,1533,1556]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="socotraensis" status="sp. nov.">
|
||
<emphasis id="0194441DFFA94C0DFEF2DD8AFDFAAF01" box="[374,538,1533,1556]" italics="true" pageId="3" pageNumber="226">O. socotraensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA7F966FFA94C0DFDA4DD8BFD92AF06" box="[544,626,1532,1555]" pageId="3" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Farmihin was first seen in 2002, when it was mistakenly thought to be a form of
|
||
<taxonomicName id="F4E0E38CFFA94C0DFDBEDE6AFD7AAF21" authorityName="Halstead" authorityYear="1980" box="[570,666,1565,1588]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="canus">
|
||
<emphasis id="0194441DFFA94C0DFDBEDE6AFD7AAF21" box="[570,666,1565,1588]" italics="true" pageId="3" pageNumber="226">O. canus</emphasis>
|
||
</taxonomicName>
|
||
with more prominent anterior pronotal angles, etc. Its true status became apparent when the author received additional specimens last year. [Unfortunately, the record of
|
||
<taxonomicName id="F4E0E38CFFAE4C0AFDB4D8AAFD6EA9E1" authorityName="Halstead" authorityYear="1980" box="[560,654,221,244]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="canus">
|
||
<emphasis id="0194441DFFAE4C0AFDB4D8AAFD6EA9E1" box="[560,654,221,244]" italics="true" pageId="4" pageNumber="227">O. canus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="4BF7D89FFFAE4C0AFD4AD8ABFCFFA9E1" box="[718,799,220,244]" name="Yemen" pageId="4" pageNumber="227">Socotra</collectingCountry>
|
||
that was included in
|
||
<bibRefCitation id="5771E5FEFFAE4C0AFC7AD8ABFF3CA801" author="HALSTEAD D. G. H." pageId="4" pageNumber="227" pagination="233 - 245" refId="ref3917" refString="HALSTEAD D. G. H. 2011: Order Coleoptera, family Silvanidae. Pp. 233 - 245. In: HARTEN A. VAN (ed.): Arthropod Fauna of the United Arab Emirates. Volume 4. Multiply Marketing Consultancy Services, Abu Dhabi, 832 pp." type="book chapter" year="2011">HALSTEAD (2011)</bibRefCitation>
|
||
was based on the Farmihin specimen and consequently is wrong. However,
|
||
<taxonomicName id="F4E0E38CFFAE4C0AFB8CD88AFB86A801" authorityName="Halstead" authorityYear="1980" box="[1032,1126,253,276]" class="Insecta" family="Silvanidae" genus="Oryzaephilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="canus">
|
||
<emphasis id="0194441DFFAE4C0AFB8CD88AFB86A801" box="[1032,1126,253,276]" italics="true" pageId="4" pageNumber="227">O. canus</emphasis>
|
||
</taxonomicName>
|
||
occurs in
|
||
<collectingCountry id="4BF7D89FFFAE4C0AFF7BD96BFEB9A821" box="[255,345,284,308]" name="Somalia" pageId="4" pageNumber="227">Somalia</collectingCountry>
|
||
, as well as other parts of East Africa, and in the
|
||
<collectingCountry id="4BF7D89FFFAE4C0AFCDDD96BFBA5A821" box="[857,1093,284,308]" name="United Arab Emirates" pageId="4" pageNumber="227">United Arab Emirates</collectingCountry>
|
||
, so perhaps it will be found in
|
||
<collectingCountry id="4BF7D89FFFAE4C0AFE33D94BFDE8A841" box="[439,520,316,340]" name="Yemen" pageId="4" pageNumber="227">Socotra</collectingCountry>
|
||
in the future.]
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFACB84FFAE4C0AFF13D92BFEA2A881" pageId="4" pageNumber="227" type="etymology">
|
||
<paragraph id="335F980FFFAE4C0AFF13D92BFEA2A881" blockId="4.[151,1127,220,436]" pageId="4" pageNumber="227">
|
||
<emphasis id="0194441DFFAE4C0AFF13D92BFEF8A861" bold="true" box="[151,280,348,372]" pageId="4" pageNumber="227">Etymology.</emphasis>
|
||
The geographic species name has been used because
|
||
<collectingCountry id="4BF7D89FFFAE4C0AFCE3D92BFC58A861" box="[871,952,348,372]" name="Yemen" pageId="4" pageNumber="227">Socotra</collectingCountry>
|
||
is where it was first discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFACB84FFAE4C0AFF13D9ECFBF9A8A1" box="[151,1049,411,436]" pageId="4" pageNumber="227" type="distribution">
|
||
<paragraph id="335F980FFFAE4C0AFF13D9ECFBF9A8A1" blockId="4.[151,1127,220,436]" box="[151,1049,411,436]" pageId="4" pageNumber="227">
|
||
<emphasis id="0194441DFFAE4C0AFF13D9ECFECAA8A6" bold="true" box="[151,298,411,435]" pageId="4" pageNumber="227">Distribution.</emphasis>
|
||
So far known only from two localities in Noged plain,
|
||
<collectingCountry id="4BF7D89FFFAE4C0AFCFCD9EBFBF5A8A1" box="[888,1045,412,436]" name="Socotra Island" pageId="4" pageNumber="227">Socotra Island</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |