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<document ID-CLB-Dataset="33275" ID-DOI="http://dx.doi.org/10.3897/zookeys.666.9191" ID-PMC="PMC5534527" ID-Pensoft-Pub="1313-2970-666-1" ID-Pensoft-UUID="C43EEB41A94B930FFE439D1FAD29FF9C" ID-PubMed="28769631" ID-Zenodo-Dep="579453" ID-ZooBank="B3C377E8BBB14F328AECA2C22D1E625A" ModsDocID="1313-2970-666-1" checkinTime="1491551283800" checkinUser="pensoft" docAuthor="Moore, Matthew R., Jameson, Mary L., Garner, Beulah H., Audibert, Cedric, Smith, Andrew B. T. &amp; Seidel, Matthias" docDate="2017" docId="9513C4F64A64D44155701CA1760136F9" docLanguage="en" docName="C43EEB41A94B930FFE439D1FAD29FF9C" docOrigin="ZooKeys 666" docPubDate="2017-04-06" docSource="http://dx.doi.org/10.3897/zookeys.666.9191" docTitle="Homonyx planicostatus Blanchard 1851" docType="treatment" docVersion="9" id="C43EEB41A94B930FFE439D1FAD29FF9C" lastPageNumber="1" masterDocId="C43EEB41A94B930FFE439D1FAD29FF9C" masterDocTitle="Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies" masterLastPageNumber="349" masterPageNumber="1" pageNumber="1" updateTime="1719493172494" updateUser="carolina">
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<mods:title id="ED5346837311AEC7EBA34D60FD84E910">Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies</mods:title>
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<mods:namePart id="6841E1FC17C2FBC61440284BB5257189">Moore, Matthew R.</mods:namePart>
<mods:affiliation id="5C7A8DEC2D224AA9C6AFE8A0E1FAF58D">Department of Entomology and Nematology, University of Florida Building 1881 Natural Drive Area, Steinmetz Hall, Box 110620, Gainesville, FL 32611 - 0620, USA</mods:affiliation>
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<mods:namePart id="415B6B707B08E095729F7AE4F6EAD36B">Jameson, Mary L.</mods:namePart>
<mods:affiliation id="BA5A590A75A78CC4E635099D45114A10">Department of Biological Sciences, Wichita State University 1845 Fairmount, Box 26, Wichita, KS 67260 - 0026, USA</mods:affiliation>
<mods:nameIdentifier id="91C86471138F2A91F064BE8F66DDC631" type="email">maryliz.jameson@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3F447CA364918272F0FD49C317017674">Garner, Beulah H.</mods:namePart>
<mods:affiliation id="570584DD75C8D627BAD488B7238261B4">Natural History Museum, Insects Division, Department of Life Sciences, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="93865E3E1EC60475EAB57030596C3403">Audibert, Cedric</mods:namePart>
<mods:affiliation id="50AFDD549759786B3DD65483E5EE6021">Musee des Confluences, Centre de Conservation et d'Etude des Collections, 13 A Rue Bancel, F- 69007 Lyon, France</mods:affiliation>
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<mods:namePart id="ABDF9B1B5EBD319D33C73F94B01F269D">Smith, Andrew B. T.</mods:namePart>
<mods:affiliation id="0CF2C41112381DC3A1F86D2257AEFC5D">Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada</mods:affiliation>
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<mods:namePart id="9A3F09A9DAD6BC1EF326F463526D506F">Seidel, Matthias</mods:namePart>
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<treatment id="9513C4F64A64D44155701CA1760136F9" ID-GBIF-Taxon="128344629" LSID="urn:lsid:plazi:treatment:9513C4F64A64D44155701CA1760136F9" httpUri="http://treatment.plazi.org/id/9513C4F64A64D44155701CA1760136F9" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="3A65D1A6D2E7E4A83F4F9FF226B1D0D6" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="FDD1BCF1E7BFADF5EC127C73A2076D7A" pageId="0" pageNumber="1">
<taxonomicName id="AB11C90B793CA16B3BFC04AB30465E4F" ID-CoL="6MD44" LSID="9513C4F6-4A64-D441-5570-1CA1760136F9" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Homonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homonyx planicostatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="planicostatus">Homonyx planicostatus Blanchard, 1851</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A21610AE548A75894FC4AA1217E27E9E" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="34159D2875B64835DA4BF06E3A94D6FB" pageId="0" pageNumber="1">
<taxonomicName id="C028A36F8A94AD7848A771DFB7DF2C91" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Homonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homonyx planicostatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="planicostatus">Homonyx planicostatus</taxonomicName>
Blanchard, 1851: 214 [original combination].
</paragraph>
</subSubSection>
<subSubSection id="255C4BB54F0F0028513FD49AC3E18A96" pageId="0" pageNumber="1" type="distribution">
<paragraph id="BBA5ED635A666085480081FE6709A826" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="0255BA0A37A74F22A0381CE0E9600D3E" pageId="0" pageNumber="1">
ARGENTINA: Mendoza,
<normalizedToken id="D9AD65289D4C6AED60D4BB77514D2085" originalValue="Tucumán">Tucuman</normalizedToken>
(
<bibRefCitation id="C6FFB3315A2491D14A4950F1EF2ED967" author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" pagination="487 - 511" refId="B274" refString="Ohaus, F, 1913. XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift 1913: 487 - 511" title="XI. Beitrag zur Kenntnis der Ruteliden. (Col.)." volume="1913" year="1913">Ohaus 1913</bibRefCitation>
,
<bibRefCitation id="4047376C5DF1AE273EBE4CF28590F497" author="Ohaus, F" journalOrPublisher="Mitteilungen der Deutschen Entomologischen Gesellschaft" pageId="0" pageNumber="1" refId="B286" refString="Ohaus, F, 1934b. Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934 b">1934b</bibRefCitation>
,
<bibRefCitation id="807595525AFFE863C700EF0BBA766F49" author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="189 - 265" refId="B14" refString="Blackwelder, RE, 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin 185: 189 - 265" title="Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
,
<bibRefCitation id="FCD35C9CCD5B5C515FAB02356D84E836" author="Gutierrez, AR" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="1" pagination="207 - 227" refId="B105" refString="Gutierrez, AR, 1952. Notas sobre Scarabaeidae neotropicos (III) (Coleoptera). Revista Chilena de Entomologia 2: 207 - 227" title="Notas sobre Scarabaeidae neotropicos (III) (Coleoptera)." volume="2" year="1952">
<normalizedToken id="74708FFE7585FF49192F9D082E111920" originalValue="Gutiérrez">Gutierrez</normalizedToken>
1952
</bibRefCitation>
,
<bibRefCitation id="0F89261B3C90DEEB8357C82EE1EC7A4C" author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="1" pagination="1 - 361" refId="B188" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
). BOLIVIA (
<bibRefCitation id="9DF31AAC4A7998EBF4D84E94BBAFCA8C" author="Blanchard, CE" editor="Milne-Edwards, H" journalOrPublisher="Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris" pageId="0" pageNumber="1" pagination="129 - 240" refId="B17" refString="Blanchard, CE, 1851. Ordre des Coleoptera. In: Milne-Edwards, H, Blanchard, CE, Lucas, H, Eds., Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris: 129 - 240" title="Ordre des Coleoptera." volumeTitle="Museum d'Histoire Naturelle de Paris." year="1851">Blanchard 1851</bibRefCitation>
,
<bibRefCitation id="9990B85DD7F67395306579213CDD4ACB" author="Burmeister, HCC" journalOrPublisher="Enslin, Berlin" pageId="0" pageNumber="1" refId="B25" refString="Burmeister, HCC, 1855. Handbuch der Entomologie (ColeopteraLamellicorniaPhyllophaga chaenochela). Volume 4, Part 2. T. C. F. Enslin, Berlin" title="Handbuch der Entomologie (ColeopteraLamellicorniaPhyllophaga chaenochela). Volume 4, Part 2. T. C. F." year="1855">Burmeister 1855</bibRefCitation>
,
<bibRefCitation id="E61134627C4F825B83183E1EC9BEAB90" author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" pagination="487 - 511" refId="B274" refString="Ohaus, F, 1913. XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift 1913: 487 - 511" title="XI. Beitrag zur Kenntnis der Ruteliden. (Col.)." volume="1913" year="1913">Ohaus 1913</bibRefCitation>
,
<bibRefCitation id="4B2B5F8AA24F8D3AE8A94D8BF501C490" author="Ohaus, F" journalOrPublisher="Mitteilungen der Deutschen Entomologischen Gesellschaft" pageId="0" pageNumber="1" refId="B286" refString="Ohaus, F, 1934b. Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934 b">1934b</bibRefCitation>
,
<bibRefCitation id="7D23225A3C3EC0DC377F3E969756F074" author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="189 - 265" refId="B14" refString="Blackwelder, RE, 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin 185: 189 - 265" title="Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
,
<bibRefCitation id="F3497D05265F52E8534D6A58C00FF5F5" author="Gutierrez, AR" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="1" pagination="207 - 227" refId="B105" refString="Gutierrez, AR, 1952. Notas sobre Scarabaeidae neotropicos (III) (Coleoptera). Revista Chilena de Entomologia 2: 207 - 227" title="Notas sobre Scarabaeidae neotropicos (III) (Coleoptera)." volume="2" year="1952">
<normalizedToken id="5AA1B6B72EF4A294A0E9FC16F7058EF7" originalValue="Gutiérrez">Gutierrez</normalizedToken>
1952
</bibRefCitation>
,
<bibRefCitation id="37CB76D1FD969A1016ECCDF79EB71D75" author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="1" pagination="1 - 361" refId="B188" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
,
<bibRefCitation id="98A28FB3664EB2BC6632A99EDFB0FF9E" author="Krajcik, M" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="0" pageNumber="1" refId="B155" refString="Krajcik, M, 2008. Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Second Edition. Plzen, 142 pp." title="Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Second Edition. Plzen, 142 pp." year="2008">Krajcik 2008</bibRefCitation>
,
<bibRefCitation id="A578162D7DCE4A6760D514F25846EEBB" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B361" refString="Soula, M, 2010a. Les Coleopteres du Nouveau Monde. Volume 4: Rutelini 4. Revision des Pelidnotina 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 4: Rutelini 4. Revision des Pelidnotina 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2010 a">Soula 2010a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2C33AB576939511FFC06CED48D999C3" pageId="0" pageNumber="1" type="types">
<paragraph id="AE8FD9332E5CC1557E545254EDF4AD54" pageId="0" pageNumber="1">Types.</paragraph>
<paragraph id="EF93B85CE2FE7903D178EB189388C412" pageId="0" pageNumber="1">
1 ♀ syntype at MNHN (
<bibRefCitation id="4B9A6DF509331D5E16813C6FDF91ED35" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B361" refString="Soula, M, 2010a. Les Coleopteres du Nouveau Monde. Volume 4: Rutelini 4. Revision des Pelidnotina 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 4: Rutelini 4. Revision des Pelidnotina 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2010 a">Soula 2010a</bibRefCitation>
). An exemplar specimen identified by Ohaus and compared with
<normalizedToken id="E09FC70909D10A6567A147541E2F37C6" originalValue="Blanchards">Blanchard's</normalizedToken>
type specimen is figured (Fig.
<figureCitation id="AE622E4EAEEC7745544ABE6E93E56D20" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Homonyx planicostatus Blanchard (male specimen compared [by Ohaus] with Blanchard's type from MNHN). A Dorsal habitus B Lateral habitus C Specimen labels and male genitalia D Male genitalia, lateral view E Male parameres, caudal view." figureDoi="10.3897/zookeys.666.9191.figure31" httpUri="https://binary.pensoft.net/fig/130459" pageId="0" pageNumber="1">31</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9E708C72C0E884FA2107BD6A98FA2161" pageId="0" pageNumber="1" type="remarks">
<paragraph id="997C97E30196E84CBFAEB86EBDC81E74" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="F671D3E77D75394A1000AF8D3B35B315" pageId="0" pageNumber="1">
CCECL contains a
<taxonomicName id="88B8E5DF96A2FF9C6A1A7E7C196C8910" family="Scarabaeidae" kingdom="Animalia" lsidName="H. planicostatus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="planicostatus">
<emphasis id="073B3BAFCF8C2AD13C873FE3B39FFA1D" italics="true" pageId="0" pageNumber="1">H. planicostatus</emphasis>
</taxonomicName>
specimen labeled as a male ♂
<normalizedToken id="E53597F964E0FFA3DD00E255D336893C" originalValue="alloréférent">alloreferent</normalizedToken>
with the following data: &quot;Vaccaguzman [arrow] Camiri coll. - SOULA [obverse] 1615m//
<normalizedToken id="AC8F500F240FA99A675AD7744C19457B" originalValue="Alloréférent">Alloreferent</normalizedToken>
♂ de
<taxonomicName id="7298350243126D7897D7AA45F232F48A" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Homonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homonyx planicostatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="planicostatus">
<emphasis id="63863C415F1B6FB9A0B9E2F74715882D" italics="true" pageId="0" pageNumber="1">Homonyx planicostatus</emphasis>
</taxonomicName>
Bl. M. SOULA det 19 (47030995). Genitalia card-mounted underneath specimen. Box 4618689 SOULA.
</paragraph>
<caption id="7CDD465E37E0B549DC87A05E5997712B" ID-Zenodo-Dep="982359" doi="10.3897/zookeys.666.9191.figure31" httpUri="https://binary.pensoft.net/fig/130459" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
<paragraph id="4A145A80608C844DC659F426C9D82B5D" pageId="0" pageNumber="1">
<emphasis id="BB80B037DED47FED9647ABAD1FBCDE60" bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
<taxonomicName id="1EBCFA8351DFE9C561A8C1E17CDFF891" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Homonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homonyx planicostatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="planicostatus">
<emphasis id="130190BED03159A7EA940E9B26955B55" italics="true" pageId="0" pageNumber="1">Homonyx planicostatus</emphasis>
</taxonomicName>
Blanchard (male specimen compared [by Ohaus] with
<normalizedToken id="CCC1C7810EB83433583D697AF32C626A" originalValue="Blanchards">Blanchard's</normalizedToken>
type from MNHN).
<emphasis id="10676D608AC08FD5582FFF8E71829197" bold="true" pageId="0" pageNumber="1">A</emphasis>
Dorsal habitus
<emphasis id="3FD1AF8AB3DF7DDDD84020BD686ADEBC" bold="true" pageId="0" pageNumber="1">B</emphasis>
Lateral habitus
<emphasis id="55001E80BEF7AA3A33E92BCCBA349326" bold="true" pageId="0" pageNumber="1">C</emphasis>
Specimen labels and male genitalia
<emphasis id="4E3C1E8367C53084EFC62AA532DE0537" bold="true" pageId="0" pageNumber="1">D</emphasis>
Male genitalia, lateral view
<emphasis id="227F3D343AF076FE6DCC2E61A5F9271E" bold="true" pageId="0" pageNumber="1">E</emphasis>
Male parameres, caudal view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>