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<document id="7F612824E1DE5F70880C26D02D119746" ID-CLB-Dataset="2379" ID-DOI="10.5281/zenodo.5314482" ID-GBIF-Dataset="fccd2191-5b05-4b4c-93e2-6427cbd89bee" ID-ISSN="0374-1036" ID-Zenodo-Dep="5314482" ID-ZooBank="AA858A7C-0AA8-4C16-8848-60707709CE4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1630121610907" checkinUser="marcus" docAuthor="Batelka, Jan &amp; Bologna, Marco A." docDate="2014" docId="1118F66DDE70FFE5FE67FA06BE15F9FB" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.241-268.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Zonitoschema arabica Batelka &amp; Bologna 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="246" masterDocId="ED218E15DE72FFE0FFB1FF96BD7FFFA1" masterDocTitle="A review of the Saharo-Sindian species of the genus Zonitoschema (Coleoptera: Meloidae), with description of new species from Tunisia, Yemen and Socotra Island" masterLastPageNumber="268" masterPageNumber="241" pageNumber="243" updateTime="1699165552238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="77D991CBACB5291958A1346504EAA72F">A review of the Saharo-Sindian species of the genus Zonitoschema (Coleoptera: Meloidae), with description of new species from Tunisia, Yemen and Socotra Island</mods:title>
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<mods:name id="BECC52A00BEC937E2B57C5E4B820E87A" type="personal">
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<mods:namePart id="7813E8F4866AD4FC541557BACEF0DA41">Batelka, Jan</mods:namePart>
<mods:affiliation id="CF69D76069A4533CE2FE19EBCD2E0031">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="AF3E573FD547BF5AF2E706941B2A6E33" type="email">janbat@centrum.cz</mods:nameIdentifier>
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<mods:namePart id="99FA3AE9D7030E70658EADF5E9064B96">Bologna, Marco A.</mods:namePart>
<mods:affiliation id="4E2C121F81E40D71405FEABBBEA2DF71">Dipartimento di Scienze, Università Roma Tre, Viale G. Marconi 446, 00146 Roma, Italy</mods:affiliation>
<mods:nameIdentifier id="51A87E083A58F57141AC7D50DD55C2D9" type="email">marcoalberto.bologna@uniroma3.it</mods:nameIdentifier>
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<mods:title id="F79E90D51F448ABE4F61307E0B2CA4BB">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="AC854D28F95BE6613DFE3B2B0C83E427">2014</mods:date>
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<treatment id="1118F66DDE70FFE5FE67FA06BE15F9FB" ID-DOI="http://doi.org/10.5281/zenodo.5457255" ID-GBIF-Taxon="192663954" ID-Zenodo-Dep="5457255" LSID="urn:lsid:plazi:treatment:1118F66DDE70FFE5FE67FA06BE15F9FB" httpUri="http://treatment.plazi.org/id/1118F66DDE70FFE5FE67FA06BE15F9FB" lastPageId="5" lastPageNumber="246" pageId="2" pageNumber="243">
<subSubSection id="D1AB14F0DE70FFE2FE67FA06BE57FA09" box="[470,808,1424,1448]" pageId="2" pageNumber="243" type="nomenclature">
<paragraph id="990E477BDE70FFE2FE67FA06BE57FA09" blockId="2.[470,808,1424,1448]" box="[470,808,1424,1448]" pageId="2" pageNumber="243">
<heading id="C246F017DE70FFE2FE67FA06BE57FA09" bold="true" box="[470,808,1424,1448]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="243" reason="3">
<emphasis id="ABC59B69DE70FFE2FE67FA06BE57FA09" bold="true" box="[470,808,1424,1448]" pageId="2" pageNumber="243">
<taxonomicName id="5EB13CF8DE70FFE2FE67FA06BFB4FA09" authority="Batelka &amp; Bologna, 2014" authorityName="Batelka &amp; Bologna" authorityYear="2014" box="[470,715,1424,1448]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="ABC59B69DE70FFE2FE67FA06BFB4FA09" bold="true" box="[470,715,1424,1448]" italics="true" pageId="2" pageNumber="243">Zonitoschema arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0F62612DE70FFE2FD63FA07BE57FA09" box="[722,808,1425,1448]" pageId="2" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D1AB14F0DE70FFE2FDACFA39BF9FFA63" box="[541,736,1455,1474]" pageId="2" pageNumber="243" type="description">
<paragraph id="990E477BDE70FFE2FDACFA39BF9FFA63" blockId="2.[541,736,1455,1474]" box="[541,736,1455,1474]" pageId="2" pageNumber="243">
(
<figureCitation id="018A5BFEDE70FFE2FD92FA39BF25FA63" box="[547,602,1455,1474]" captionStart="Figs 12. 1" captionStartId="4.[151,188,1070,1089]" captionTargetBox="[249,1028,233,1047]" captionTargetId="figure-169@4.[237,1041,219,1058]" captionTargetPageId="4" captionText="Figs 12. 1 Zonitoschema arabica sp.nov., dorsal habitus of holotype (male); 2 Z.kaszabi sp. nov., dorsal habitus of female paratype. Not on the same scale." figureDoi="http://doi.org/10.5281/zenodo.5314484" httpUri="https://zenodo.org/record/5314484/files/figure.png" pageId="2" pageNumber="243">Figs 1</figureCitation>
,
<figureCitation id="018A5BFEDE70FFE2FDD5FA39BF04FA63" box="[612,635,1455,1474]" captionStart="Figs 2834" captionStartId="13.[113,150,1520,1539]" captionTargetBox="[113,1088,219,1485]" captionTargetId="figure-11@13.[113,1091,219,1486]" captionTargetPageId="13" captionText="Figs 2834. Heads. 28 Zonitoschema iranica Kaszab, 1959 from Iran. 29 Z. griseohirta Pic, 1914 from Yemen. 30 Z. pallidissima (Reitter, 1908) from Morocco. 3132 Z. pallidissima from Egypt: 31 female; 32 male. 33 Z. arabica sp. nov., holotype (male). 34 Z. kaszabi sp. nov., paratype (female). Not on the same scale." figureDoi="http://doi.org/10.5281/zenodo.5314492" httpUri="https://zenodo.org/record/5314492/files/figure.png" pageId="2" pageNumber="243">33</figureCitation>
,
<figureCitation id="018A5BFEDE70FFE2FD35FA39BFE4FA63" box="[644,667,1455,1474]" captionStart="Figs 3549" captionStartId="14.[151,188,1439,1458]" captionTargetBox="[211,1088,223,1380]" captionTargetId="figure-11@14.[187,1092,219,1402]" captionTargetPageId="14" captionText="Figs 3549. Pronotal disc, dorsal view. 3536 Zonitoschema iranica Kaszab, 1959: 35 male from the U.A.E.; 36 female from Iran. 37 Z. kaszabi sp. nov., paratype. 3839 Z. sp. cf. genicularis (Wellman, 1910): 38 specimen from Somalia; 39 specimen from Yemen. 4041 Z. pallidissima (Reitter, 1908) from Egypt: 40 male; 41 female. 42 Z. pallidissima from Morocco. 43 Z. rubricolor Pic, 1924 from Kenya. 4445 Z. chourriba sp. nov.: 44 male paratype; 45 female paratype. 46 Z. arabica sp. nov., holotype. 4748 Z. oculatissima Peyerimhoff, 1929: 47 holotype; 48 specimen from Chad. 49 Z. cf. pallidissima from Algeria. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.5314494" httpUri="https://zenodo.org/record/5314494/files/figure.png" pageId="2" pageNumber="243">46</figureCitation>
,
<figureCitation id="018A5BFEDE70FFE2FD15FA39BFA5FA63" box="[676,730,1455,1474]" captionStart="Figs 5059" captionStartId="15.[113,150,1457,1476]" captionTargetBox="[127,1088,247,1307]" captionTargetId="figure-11@15.[113,1091,219,1320]" captionTargetPageId="15" captionText="Figs 5059. Male genitalia, lateral view (57 and 59, dorsal view). 5051 Z. genicularis (Wellman, 1910): 50 specimen from Zambia; 51 specimen from Somalia (parameres only). 52 Z. iranica Kaszab, 1959 from Israel. 53 Z. pallidissima (Reitter, 1908). 54 Z. gibdoana (Kaszab, 1956) from Kenya.55 Z. sp. from Kenya (a species similar in colouration to Z. sp. cf. genicularis (Wellman, 1910)). 5657 Zonitoschema arabica sp. nov., holotype. 5859 Zonitoschema chourriba sp. nov., paratype. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.5314496" httpUri="https://zenodo.org/record/5314496/files/figure.png" pageId="2" pageNumber="243">56, 57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D1AB14F0DE70FFE2FF26FA4FBC31F9F7" pageId="2" pageNumber="243" type="materials_examined">
<paragraph id="990E477BDE70FFE2FF26FA4FBE40FA4D" blockId="2.[151,1128,1497,1622]" box="[151,831,1497,1516]" pageId="2" pageNumber="243">
<emphasis id="ABC59B69DE70FFE2FF26FA4FBC6FFA4D" bold="true" box="[151,272,1497,1516]" pageId="2" pageNumber="243">
<typeStatus id="460AF9D9DE70FFE2FF26FA4FBDBBFA4D" box="[151,196,1497,1516]" pageId="2" pageNumber="243">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E1A607EBDE70FFE2FEA4FA4FBC2CFA4D" box="[277,339,1497,1516]" name="Yemen" pageId="2" pageNumber="243">Yemen</collectingCountry>
,
<quantity id="5E49EA9EDE70FFE2FEEAFA4FBCEEFA4D" box="[347,401,1497,1516]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="243" unit="km" value="20.0">20 km</quantity>
NW of Dhawran,
<geoCoordinate id="FC8521BCDE70FFE2FD9EFA4FBF07FA4D" box="[559,632,1497,1516]" degrees="14" direction="north" minutes="40" orientation="latitude" pageId="2" pageNumber="243" precision="925" value="14.666667">14°40ʹN</geoCoordinate>
,
<geoCoordinate id="FC8521BCDE70FFE2FD30FA4FBFB7FA4D" box="[641,712,1497,1516]" degrees="44" direction="east" minutes="13" orientation="longitude" pageId="2" pageNumber="243" precision="925" value="44.216667">44°13ʹE</geoCoordinate>
,
<quantity id="5E49EA9EDE70FFE2FD61FA4FBE6EFA4D" box="[720,785,1497,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="2" pageNumber="243" unit="m" value="1794.0">1794 m</quantity>
a.s.l.
</paragraph>
<paragraph id="990E477BDE70FFE2FF26FA65BC31F9F7" blockId="2.[151,1128,1497,1622]" pageId="2" pageNumber="243">
<emphasis id="ABC59B69DE70FFE2FF26FA65BC64F9A7" bold="true" box="[151,283,1523,1542]" pageId="2" pageNumber="243">Type material.</emphasis>
<materialsCitation id="29D94D26DE70FFE2FE91FA65BC77F980" ID-GBIF-Occurrence="3458440303" collectingDate="2005-10-29" collectionCode="NMPC" collectorName="S. Kadlec" country="Yemen" elevation="1794" latitude="14.666667" location="Dhawran" longLatPrecision="1287" longitude="44.216667" municipality="Dhawran" pageId="2" pageNumber="243" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="460AF9D9DE70FFE2FE91FA65BC0CF9A4" box="[288,371,1523,1542]" pageId="2" pageNumber="243" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="8FB78CF2DE70FFE2FECFFA78BCF1F9A6" box="[382,398,1518,1543]" pageId="2" pageNumber="243" type="male"></specimenCount>
, W.
<collectingCountry id="E1A607EBDE70FFE2FE02FA65BC8EF9A7" box="[435,497,1523,1542]" name="Yemen" pageId="2" pageNumber="243">Yemen</collectingCountry>
,
<locationDeviation id="88BC5E59DE70FFE2FE48FA65BF12F9A7" bearing="315.0" box="[505,621,1523,1542]" distance="20000.0" pageId="2" pageNumber="243">
<quantity id="5E49EA9EDE70FFE2FE48FA65BF51F9A7" box="[505,558,1523,1542]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="243" unit="km" value="20.0">20 km</quantity>
NW of
</locationDeviation>
<collectingMunicipality id="796ADD01DE70FFE2FDC3FA65BFBDF9A7" box="[626,706,1523,1542]" country="Yemen" latitude="14.666667" longLatPrecision="1287" longitude="44.216667" name="Dhawran" pageId="2" pageNumber="243">Dhawran</collectingMunicipality>
,
<geoCoordinate id="FC8521BCDE70FFE2FD7BFA65BE6CF9A7" box="[714,787,1523,1542]" degrees="14" direction="north" minutes="40" orientation="latitude" pageId="2" pageNumber="243" precision="925" value="14.666667">14°40ʹN</geoCoordinate>
,
<geoCoordinate id="FC8521BCDE70FFE2FCAAFA65BE1DF9A7" box="[795,866,1523,1542]" degrees="44" direction="east" minutes="13" orientation="longitude" pageId="2" pageNumber="243" precision="925" value="44.216667">44°13ʹE</geoCoordinate>
,
<quantity id="5E49EA9EDE70FFE2FCDBFA65BED2F9A7" box="[874,941,1523,1542]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="2" pageNumber="243" unit="m" value="1794.0">
<elevation id="129CA048DE70FFE2FCDBFA65BED2F9A7" box="[874,941,1523,1542]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="2" pageNumber="243" unit="m" value="1794.0">1794 m</elevation>
</quantity>
,
<date id="ED0F61BBDE70FFE2FC04FA65B973F9A7" box="[949,1036,1523,1542]" pageId="2" pageNumber="243" value="2005-10-29">
<collectingDate id="FD4B9853DE70FFE2FC04FA65B973F9A7" box="[949,1036,1523,1542]" pageId="2" pageNumber="243" value="2005-10-29">29.x.2005</collectingDate>
</date>
,
<collectorName id="344422ADDE70FFE2FBA5FA65B919F9A7" box="[1044,1126,1523,1542]" pageId="2" pageNumber="243">S. Kadlec</collectorName>
lgt. (
<collectionCode id="FFA0DFBEDE70FFE2FF0CF998BD81F980" box="[189,254,1550,1569]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="243">NMPC</collectionCode>
).
</materialsCitation>
<materialsCitation id="29D94D26DE70FFE2FEBAF998BC31F9F7" ID-GBIF-Occurrence="3458440364" pageId="2" pageNumber="243" typeStatus="paratype">
<typeStatus id="460AF9D9DE70FFE2FEBAF998BC26F9BE" box="[267,345,1550,1569]" pageId="2" pageNumber="243" type="paratype">PARATYPE</typeStatus>
:
<specimenCount id="8FB78CF2DE70FFE2FED2F998BCFFF980" box="[355,384,1544,1569]" pageId="2" pageNumber="243" type="female">1♀</specimenCount>
(
<collectionCode id="FFA0DFBEDE70FFE2FE3AF998BCB0F980" box="[395,463,1550,1569]" pageId="2" pageNumber="243">BMNH</collectionCode>
),Arabia [
<collectingCountry id="E1A607EBDE70FFE2FDAFF998BF1FF980" box="[542,608,1550,1569]" name="Yemen" pageId="2" pageNumber="243">Yemen</collectingCountry>
],
<collectingRegion id="5B758999DE70FFE2FDDDF998BFDEF980" box="[620,673,1550,1569]" country="Yemen" name="Ad Dali'" pageId="2" pageNumber="243">Dhala</collectingRegion>
,
<date id="ED0F61BBDE70FFE2FD19F998BF9AF980" box="[680,741,1550,1569]" pageId="2" pageNumber="243" value="1935-10">x.1935</date>
, at light, R.C.M. Darling [see remarks below]. The
<typeStatus id="460AF9D9DE70FFE2FF0CF9BFBC79F99D" box="[189,262,1577,1596]" pageId="2" pageNumber="243" type="paratype">paratype</typeStatus>
lacks totally both antennae, left foretarsi and right foreleg and is probably teneral because of the light colour of black parts.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D1AB14F0DE71FFE3FFC0FF4ABC47F992" pageId="3" pageNumber="244" type="description">
<paragraph id="990E477BDE71FFE3FFC0FF4ABE69FEF2" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="244">
<emphasis id="ABC59B69DE71FFE3FFC0FF4ABC3EFF52" bold="true" box="[113,321,219,243]" pageId="3" pageNumber="244">
Description.
<emphasis id="ABC59B69DE71FFE3FEB3FF4DBC3EFF52" bold="true" box="[258,321,219,243]" italics="true" pageId="3" pageNumber="244">Male.</emphasis>
</emphasis>
Body subopaque yellowish-brown, darker dorsally; eyes, apical third of mandibles, maxillary palpomeres IIIIV, labial palpomere III, antennomeres IIIXI, tibiae and tarsi black. Setation light golden-yellow, recumbent, not dense except on tibiae, longer on elytral sides, tibiae and tarsi. Body length:
<quantity id="5E49EA9EDE71FFE3FDEDFEAABFE6FEF2" box="[604,665,316,339]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="244" unit="mm" value="8.0">8 mm</quantity>
(
<typeStatus id="460AF9D9DE71FFE3FD16FEAABE74FEF2" box="[679,779,316,339]" pageId="3" pageNumber="244" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="990E477BDE71FFE3FF21FECABF9CFC92" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="244">
Head (
<figureCitation id="018A5BFEDE71FFE3FF68FECABC55FED2" box="[217,298,348,371]" captionStart="Figs 2834" captionStartId="13.[113,150,1520,1539]" captionTargetBox="[113,1088,219,1485]" captionTargetId="figure-11@13.[113,1091,219,1486]" captionTargetPageId="13" captionText="Figs 2834. Heads. 28 Zonitoschema iranica Kaszab, 1959 from Iran. 29 Z. griseohirta Pic, 1914 from Yemen. 30 Z. pallidissima (Reitter, 1908) from Morocco. 3132 Z. pallidissima from Egypt: 31 female; 32 male. 33 Z. arabica sp. nov., holotype (male). 34 Z. kaszabi sp. nov., paratype (female). Not on the same scale." figureDoi="http://doi.org/10.5281/zenodo.5314492" httpUri="https://zenodo.org/record/5314492/files/figure.png" pageId="3" pageNumber="244">Fig. 33</figureCitation>
) slightly wider than pronotum, maximum width at eye level. Eyes very large, bulged, anterior portion distinctly emarginated near antennal socket, extended ventrally and reaching medial margin of maxillae on underside of head, almost contiguous; frontally 1.4 times as wide as frontal narrowest space between eyes. Frontal suture curved; frontal longitudinal middle line only weakly depressed, regularly punctuate. Head punctures wide, deep, intermediate surface very narrow and shiny; setae suberected. Temples very short and distinctly narrowed posteriad, ca. 0.25 times as long as longitudinal middle diameter of eye. Clypeus flat, subtrapezoidal; anterior half glabrous and impunctate, shagreened; posterior half with punctuation and setation as on head capsule. Labrum suboval, depressed anteriorly and medially; anterior margin scarcely arcuate, about as long as clypeus. Mandibles slightly longer than apex of labrum, straight but curved in apical third. Galeae penicillate, about as long as maxillary palpi; palpomeres IIIII slender and subcylindrical, IV scarcely widened in anterior half and subtruncate at apex. Antennae elongate, ca. 5.3 times as long as pronotum disc; ratio between each antennomere and shortest one (II) as follows: I: 1.1; III: 1.1; IV: 1.5; V: 1.7; VI: 1.8; VII: 1.7; VIII: 1.7; IX: 1.7; X: 1.7; XI: 1.8.
</paragraph>
<paragraph id="990E477BDE71FFE3FF21FCAABF42FA92" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="244">
Pronotum (
<figureCitation id="018A5BFEDE71FFE3FEB8FCAABC25FCF2" box="[265,346,828,851]" captionStart="Figs 3549" captionStartId="14.[151,188,1439,1458]" captionTargetBox="[211,1088,223,1380]" captionTargetId="figure-11@14.[187,1092,219,1402]" captionTargetPageId="14" captionText="Figs 3549. Pronotal disc, dorsal view. 3536 Zonitoschema iranica Kaszab, 1959: 35 male from the U.A.E.; 36 female from Iran. 37 Z. kaszabi sp. nov., paratype. 3839 Z. sp. cf. genicularis (Wellman, 1910): 38 specimen from Somalia; 39 specimen from Yemen. 4041 Z. pallidissima (Reitter, 1908) from Egypt: 40 male; 41 female. 42 Z. pallidissima from Morocco. 43 Z. rubricolor Pic, 1924 from Kenya. 4445 Z. chourriba sp. nov.: 44 male paratype; 45 female paratype. 46 Z. arabica sp. nov., holotype. 4748 Z. oculatissima Peyerimhoff, 1929: 47 holotype; 48 specimen from Chad. 49 Z. cf. pallidissima from Algeria. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.5314494" httpUri="https://zenodo.org/record/5314494/files/figure.png" pageId="3" pageNumber="244">Fig. 46</figureCitation>
), 1.061.16 times as wide as long, sides slightly widened from basis to middle and anteriorly evidently narrowed with straight sides; fore third progressively depressed, particularly on sides; base only slightly depressed and posterior margin slightly rebordered; sides scarcely depressed in basal half in lateral view; punctures and setae as on head, setae short; prosternum progressively and weakly narrowed posteriorly. Mesonotum slightly depressed in middle, largely rounded at posterior apex; mesoventrite transverse but laterally narrowed on sides and posteriorly greatly narrowed; metaventrite large, medial line impunctate and glabrous. Elytra densely and subrugosely punctuate, punctures as deep as on head and pronotum, and setae shorter than on pronotum, except at base and on sides; venations scarcely visible. Metathoracic wings present and completely developed. Legs slender, setae of tibiae very dense and silver-yellow, those of femora shorter and more robust, particularly on ventral side. External apex of fore tibiae slightly triangularly expanded; both foretibial spurs slender, inner one more pointed; both mesotibial spurs slender at apex; both metatibial spurs stick-like, parallel and subquadrate at apex; fore and middle tarsi 1.4 times as long as respective tibia, metatarsi as long as metatibia; tarsal claws denticulate, claw teeth thin and close to each other, ventral blade very thin.
</paragraph>
<paragraph id="990E477BDE71FFE3FF21FAAABECEFA32" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="244">
Abdominal ventrites densely setated, surface almost shagreened; last ventrite deeply incised, posterior margin of ventrite IV strongly emarginated in middle. Gonoforceps short, apically narrow and strongly curved upwards, aedeagus short and robust (
<figureCitation id="018A5BFEDE71FFE3FC99FAEABEDBFA32" box="[808,932,1404,1427]" captionStart="Figs 5059" captionStartId="15.[113,150,1457,1476]" captionTargetBox="[127,1088,247,1307]" captionTargetId="figure-11@15.[113,1091,219,1320]" captionTargetPageId="15" captionText="Figs 5059. Male genitalia, lateral view (57 and 59, dorsal view). 5051 Z. genicularis (Wellman, 1910): 50 specimen from Zambia; 51 specimen from Somalia (parameres only). 52 Z. iranica Kaszab, 1959 from Israel. 53 Z. pallidissima (Reitter, 1908). 54 Z. gibdoana (Kaszab, 1956) from Kenya.55 Z. sp. from Kenya (a species similar in colouration to Z. sp. cf. genicularis (Wellman, 1910)). 5657 Zonitoschema arabica sp. nov., holotype. 5859 Zonitoschema chourriba sp. nov., paratype. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.5314496" httpUri="https://zenodo.org/record/5314496/files/figure.png" pageId="3" pageNumber="244">Figs 56, 57</figureCitation>
).
</paragraph>
<paragraph id="990E477BDE71FFE3FF21FA0DBC47F992" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="244">
<emphasis id="ABC59B69DE71FFE3FF21FA0DBD96FA12" bold="true" box="[144,233,1435,1459]" italics="true" pageId="3" pageNumber="244">Female.</emphasis>
Similar to male, except of body colouration lighter; of maxillary palpomeres only palpomere IV black; eyes only 0.9 times as wide as frontal narrowest space between eyes; frontal suture less curved; temples a little longer; pronotal setae longer; elytral venations well visible; claw teeth more robust and well separated, last ventrite V-shaped. Body length:
<quantity id="5E49EA9EDE71FFE3FFC0F98ABDC3F992" box="[113,188,1564,1587]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="244" unit="mm" value="12.0">12 mm</quantity>
(
<typeStatus id="460AF9D9DE71FFE3FF7BF98ABC53F992" box="[202,300,1564,1587]" pageId="3" pageNumber="244" type="paratype">paratype</typeStatus>
).
</paragraph>
</subSubSection>
<caption id="CDCE17F3DE76FFE4FF26FBB8BF7BFBFD" ID-DOI="http://doi.org/10.5281/zenodo.5314484" ID-Zenodo-Dep="5314484" httpUri="https://zenodo.org/record/5314484/files/figure.png" pageId="4" pageNumber="245" startId="4.[151,188,1070,1089]" targetBox="[249,1028,233,1047]" targetPageId="4">
<paragraph id="990E477BDE76FFE4FF26FBB8BF7BFBFD" blockId="4.[151,1127,1070,1116]" pageId="4" pageNumber="245">
Figs 12. 1
<taxonomicName id="5EB13CF8DE76FFE4FEB7FBB8BCBBFBE0" authority="Batelka &amp; Bologna, 2014" authorityName="Batelka &amp; Bologna" authorityYear="2014" box="[262,452,1070,1089]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="ABC59B69DE76FFE4FEB7FBB8BCBBFBE0" box="[262,452,1070,1089]" italics="true" pageId="4" pageNumber="245">Zonitoschema arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0F62612DE76FFE4FE79FBB8BF73FBE0" box="[456,524,1070,1089]" pageId="4" pageNumber="245" rank="species">sp.nov.</taxonomicNameLabel>
, dorsal habitus of holotype (male); 2
<taxonomicName id="5EB13CF8DE76FFE4FCFFFBB8BEDCFBE0" authority="Batelka &amp; Bologna, 2014" authorityName="Batelka &amp; Bologna" authorityYear="2014" box="[846,931,1070,1089]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="kaszabi" status="sp. nov.">
<emphasis id="ABC59B69DE76FFE4FCFFFBB8BEDCFBE0" box="[846,931,1070,1089]" italics="true" pageId="4" pageNumber="245">Z.kaszabi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0F62612DE76FFE4FC16FBB8BE94FBE0" box="[935,1003,1070,1089]" pageId="4" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
, dorsal habitus of female paratype. Not on the same scale.
</paragraph>
</caption>
<subSubSection id="D1AB14F0DE76FFE4FF26FB08BCACF994" pageId="4" pageNumber="245" type="diagnosis">
<paragraph id="990E477BDE76FFE4FF26FB08BF4FFA94" blockId="4.[151,1127,1182,1589]" pageId="4" pageNumber="245">
<emphasis id="ABC59B69DE76FFE4FF26FB08BCEEFB14" bold="true" box="[151,401,1182,1205]" pageId="4" pageNumber="245">Differential diagnosis.</emphasis>
Species characterized by the combination of the following features: elytra unicolour brown-yellow, femora completely orange-yellow, antennomeres III yellow, the remaining black, antennomere II ca. 3 times as long as wide.
<taxonomicName id="5EB13CF8DE76FFE4FCE0FB48B93EFB54" authority="Batelka &amp; Bologna, 2014" authorityName="Batelka &amp; Bologna" authorityYear="2014" box="[849,1089,1246,1269]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="ABC59B69DE76FFE4FCE0FB48B93EFB54" box="[849,1089,1246,1269]" italics="true" pageId="4" pageNumber="245">Zonitoschema arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0F62612DE76FFE4FBF9FB48BDBAFAB4" pageId="4" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="5EB13CF8DE76FFE4FEE1FB68BCAEFAB4" baseAuthorityName="Kaszab" baseAuthorityYear="1956" box="[336,465,1278,1301]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="gibdoana">
<emphasis id="ABC59B69DE76FFE4FEE1FB68BCAEFAB4" box="[336,465,1278,1301]" italics="true" pageId="4" pageNumber="245">Z. gibdoana</emphasis>
</taxonomicName>
, but clearly distinct by legs colour and deeper and wider punctuation of head, pronotum and elytra.
</paragraph>
<paragraph id="990E477BDE76FFE4FF04FAA8BCACF994" blockId="4.[151,1127,1182,1589]" pageId="4" pageNumber="245">
Among the Afrotropical and Palaearctic species with unicolour orange-brown or yellowish elytra,
<taxonomicName id="5EB13CF8DE76FFE4FF53FAC8BC2EFAD4" authority="Batelka &amp; Bologna, 2014" authorityName="Batelka &amp; Bologna" authorityYear="2014" box="[226,337,1374,1397]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="ABC59B69DE76FFE4FF53FAC8BC2EFAD4" box="[226,337,1374,1397]" italics="true" pageId="4" pageNumber="245">Z. arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0F62612DE76FFE4FEE9FAC8BCD4FAD4" box="[344,427,1374,1397]" pageId="4" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
represents an exception because of the completely yellow-brown femora. Only
<taxonomicName id="5EB13CF8DE76FFE4FE9DFAE8BF13FA34" authority="(Fairmaire, 1894)" baseAuthorityName="Fairmaire" baseAuthorityYear="1894" box="[300,620,1406,1429]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="ABC59B69DE76FFE4FE9DFAE8BCD9FA34" box="[300,422,1406,1429]" italics="true" pageId="4" pageNumber="245">Z. gigantea</emphasis>
(Fairmaire, 1894)
</taxonomicName>
, distributed in western Africa and in the
<collectingCountry id="E1A607EBDE76FFE4FBAEFAE8B919FA34" box="[1055,1126,1406,1429]" name="Democratic Republic of the Congo" pageId="4" pageNumber="245">Congo</collectingCountry>
basin has the same femur colouration; that species is easily distinguishable from
<taxonomicName id="5EB13CF8DE76FFE4FC46FA08B919FA14" authority="Batelka &amp; Bologna, 2014" authorityName="Batelka &amp; Bologna" authorityYear="2014" box="[1015,1126,1438,1461]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="ABC59B69DE76FFE4FC46FA08B919FA14" box="[1015,1126,1438,1461]" italics="true" pageId="4" pageNumber="245">Z. arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0F62612DE76FFE4FF26FA28BD94FA74" box="[151,235,1470,1493]" pageId="4" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
because of its greater size (ca.
<quantity id="5E49EA9EDE76FFE4FD8DFA28BFCCFA74" box="[572,691,1470,1493]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="4" pageNumber="245" unit="mm" value="19.0" valueMax="23.0" valueMin="15.0">1523 mm</quantity>
), body colour reddish, only the antennomere I yellow, and shallower dorsal punctures. However, more
<taxonomicName id="5EB13CF8DE76FFE4FCF4FA48B950FA54" box="[837,1071,1502,1525]" class="Insecta" family="Meloidae" genus="Zonitoschema" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="ABC59B69DE76FFE4FCF4FA48BEA4FA54" box="[837,987,1502,1525]" italics="true" pageId="4" pageNumber="245">Zonitoschema</emphasis>
species
</taxonomicName>
with yellow femora occur in
<collectingCountry id="E1A607EBDE76FFE4FE28FA68BCA0F9B4" box="[409,479,1534,1557]" name="Kenya" pageId="4" pageNumber="245">Kenya</collectingCountry>
(two species) and
<collectingCountry id="E1A607EBDE76FFE4FD1BFA68BE79F9B4" box="[682,774,1534,1557]" name="Namibia" pageId="4" pageNumber="245">Namibia</collectingCountry>
(one species) (M. A. Bologna &amp; J. Batelka, unpublished data).
</paragraph>
</subSubSection>
<caption id="CDCE17F3DE77FFE5FFC0FBFFBC4CFB36" ID-DOI="http://doi.org/10.5281/zenodo.5314486" ID-Zenodo-Dep="5314486" httpUri="https://zenodo.org/record/5314486/files/figure.png" pageId="5" pageNumber="246" startId="5.[113,150,1129,1148]" targetBox="[176,1006,228,1112]" targetPageId="5">
<paragraph id="990E477BDE77FFE5FFC0FBFFBC4CFB36" blockId="5.[113,1090,1129,1175]" pageId="5" pageNumber="246">
Figs 34.
<taxonomicName id="5EB13CF8DE77FFE5FF72FBFFBCECFBDD" authority="Batelka &amp; Bologna, 2014" authorityName="Batelka &amp; Bologna" authorityYear="2014" box="[195,403,1129,1148]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="chourriba" status="sp. nov.">
<emphasis id="ABC59B69DE77FFE5FF72FBFFBCECFBDD" box="[195,403,1129,1148]" italics="true" pageId="5" pageNumber="246">Zonitoschema chourriba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0F62612DE77FFE5FE26FBFFBCA6FBDD" box="[407,473,1129,1148]" pageId="5" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
3 dorsal habitus of female paratype; 4 dorsal habitus of male paratype. Not on the same scale.
</paragraph>
</caption>
<subSubSection id="D1AB14F0DE77FFE5FFC0FB55BF63FB5B" pageId="5" pageNumber="246" type="etymology">
<paragraph id="990E477BDE77FFE5FFC0FB55BF63FB5B" blockId="5.[113,1089,1219,1626]" pageId="5" pageNumber="246">
<emphasis id="ABC59B69DE77FFE5FFC0FB55BD8FFB7B" bold="true" box="[113,240,1219,1242]" pageId="5" pageNumber="246">Etymology.</emphasis>
The name of this new species refers to the Arabian Peninsula, in the south-western part of which it is distributed; adjective.
</paragraph>
</subSubSection>
<subSubSection id="D1AB14F0DE77FFE5FFC0FA95BCD3FA7B" pageId="5" pageNumber="246" type="discussion">
<paragraph id="990E477BDE77FFE5FFC0FA95BCD3FA7B" blockId="5.[113,1089,1219,1626]" pageId="5" pageNumber="246">
<emphasis id="ABC59B69DE77FFE5FFC0FA95BCE6FABB" bold="true" box="[113,409,1283,1306]" pageId="5" pageNumber="246">Collecting circumstances.</emphasis>
The type locality is a hill with flowering
<taxonomicName id="5EB13CF8DE77FFE5FCC7FA95BCF9FA9B" authority="Schweinf." authorityName="Schweinf." class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="246" phylum="Tracheophyta" rank="species" species="ammak">
<emphasis id="ABC59B69DE77FFE5FCC7FA95B93FFABB" box="[886,1088,1283,1306]" italics="true" pageId="5" pageNumber="246">Euphorbia ammak</emphasis>
Schweinf. (Euphorbiacae)
</taxonomicName>
(D. Král, pers. comm.). The expedition stayed at the locality overnight and used light to attract nocturnal insects. The
<typeStatus id="460AF9D9DE77FFE5FD28FAD5BF89FAFB" box="[665,758,1347,1370]" pageId="5" pageNumber="246" type="holotype">holotype</typeStatus>
was probably collected at light because it is covered with small white scales of some nocturnal Lepidoptera. The specimen from
<collectingRegion id="5B758999DE77FFE5FF1CFA15BD92FA3B" box="[173,237,1411,1434]" country="Yemen" name="Ad Dali'" pageId="5" pageNumber="246">Dhala</collectingRegion>
was collected at light as it is indicated on the locality label. The
<collectingRegion id="5B758999DE77FFE5FC1BFA15BE95FA3B" box="[938,1002,1411,1434]" country="Yemen" name="Ad Dali'" pageId="5" pageNumber="246">Dhala</collectingRegion>
locality and its vicinity (visited again in
<date id="ED0F61BBDE77FFE5FE79FA35BF0AFA1B" box="[456,629,1443,1466]" pageId="5" pageNumber="246" value="1937-09">September 1937</date>
by another British expedition) is described and pictured in
<bibRefCitation id="FD203A8ADE77FFE5FEA9FA55BCD7FA7B" author="SCOTT H." box="[280,424,1475,1498]" pageId="5" pageNumber="246" pagination="97 - 121" refId="ref16798" refString="SCOTT H. 1939: A journey to the Yemen. Geographical Journal 93: 97 - 121." type="journal article" year="1939">SCOTT (1939)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D1AB14F0DE77FFE5FFC0FA75BE15F9FB" pageId="5" pageNumber="246" type="discussion">
<paragraph id="990E477BDE77FFE5FFC0FA75BE15F9FB" blockId="5.[113,1089,1219,1626]" pageId="5" pageNumber="246">
<emphasis id="ABC59B69DE77FFE5FFC0FA75BD9FFA5B" bold="true" box="[113,224,1507,1530]" pageId="5" pageNumber="246">Remarks.</emphasis>
The
<typeStatus id="460AF9D9DE77FFE5FEA8FA75BC0AFA5B" box="[281,373,1507,1530]" pageId="5" pageNumber="246" type="paratype">paratype</typeStatus>
specimen was erroneously recorded by
<bibRefCitation id="FD203A8ADE77FFE5FC99FA75B93FFA5B" author="BOLOGNA M. A. &amp; TURCO F." box="[808,1088,1507,1530]" pageId="5" pageNumber="246" pagination="1 - 33" refId="ref15914" refString="BOLOGNA M. A. &amp; TURCO F. 2007: The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa 1625: 1 - 33." type="journal article" year="2007">BOLOGNA &amp; TURCO (2007)</bibRefCitation>
as ʻ
<taxonomicName id="5EB13CF8DE77FFE5FF2AF995BC26F9BB" box="[155,345,1539,1562]" class="Insecta" family="Meloidae" genus="Zonitoschema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="ABC59B69DE77FFE5FF2AF995BC4EF9BB" box="[155,305,1539,1562]" italics="true" pageId="5" pageNumber="246">Zonitoschema</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="5EB13CF8DE77FFE5FEDDF995BC8CF9BB" authorityName="Peyerimhoff" authorityYear="1929" box="[364,499,1539,1562]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="oculatissima">
<emphasis id="ABC59B69DE77FFE5FEDDF995BC8CF9BB" box="[364,499,1539,1562]" italics="true" pageId="5" pageNumber="246">oculatissima</emphasis>
</taxonomicName>
?)ʼ from ʻSaudi Arabia: Dhahʼ. After a more detailed analysis, the specimens proved to be different from
<taxonomicName id="5EB13CF8DE77FFE5FD29F9B5BE44F99B" authorityName="Peyerimhoff" authorityYear="1929" box="[664,827,1571,1594]" class="Insecta" family="Meloidae" genus="Zonitoschema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="oculatissima">
<emphasis id="ABC59B69DE77FFE5FD29F9B5BFD8F99B" box="[664,679,1571,1594]" italics="true" pageId="5" pageNumber="246">Z</emphasis>
.
<emphasis id="ABC59B69DE77FFE5FD05F9B5BE44F99B" box="[692,827,1571,1594]" italics="true" pageId="5" pageNumber="246">oculatissima</emphasis>
</taxonomicName>
, and the correct spelling of the locality label is
<collectingRegion id="5B758999DE77FFE5FEEEF9D5BCDEF9FB" box="[351,417,1603,1626]" country="Yemen" name="Ad Dali'" pageId="5" pageNumber="246">Dhala</collectingRegion>
, in SW
<collectingCountry id="E1A607EBDE77FFE5FE44F9D5BF40F9FB" box="[501,575,1603,1626]" name="Yemen" pageId="5" pageNumber="246">Yemen</collectingCountry>
(
<quantity id="5E49EA9EDE77FFE5FDFCF9D5BFE0F9FB" box="[589,671,1603,1626]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="246" unit="km" value="120.0">120 km</quantity>
SSE of Dhawran).
</paragraph>
</subSubSection>
</treatment>
</document>