546 lines
64 KiB
XML
546 lines
64 KiB
XML
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<mods:title id="9DDAB86BA666A18C199155D0CB0C6177">Dragons in neglect: Taxonomic revision of the Sulawesi sailfin lizards of the genus Hydrosaurus Kaup, 1828 (Squamata, Agamidae)</mods:title>
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<mods:namePart id="AD26764F7259C3DE866E1E94B34EE048">Denzer, Wolfgang</mods:namePart>
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<mods:namePart id="6A86B652ABAF92C32DE9D7A6697EB9CC">Campbell, Patrick D.</mods:namePart>
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<mods:namePart id="A40693E5E0478AEEA1C3C2C0B7703175">Manthey, Ulrich</mods:namePart>
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<mods:namePart id="A783219EA73994BCF45471188A68903F">Glässer-Trobisch, Andrea</mods:namePart>
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<mods:namePart id="0892E9DEF1D01AA9B3136C2619BAF044">Koch, André</mods:namePart>
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<mods:title id="A36C6871618540BBEE236761625B394B">Zootaxa</mods:title>
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<mods:date id="857FF5B77FC5EC38EBE982D935ABF321">2020</mods:date>
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<mods:number id="5CDAFD3122F5BCFBCCDD7B087786CE5C">2020-03-03</mods:number>
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<treatment id="FF530825FF947B55FF7A7C4376C2FC52" ID-DOI="http://doi.org/10.5281/zenodo.3703628" ID-GBIF-Taxon="162381185" ID-Zenodo-Dep="3703628" LSID="urn:lsid:plazi:treatment:FF530825FF947B55FF7A7C4376C2FC52" httpUri="http://treatment.plazi.org/id/FF530825FF947B55FF7A7C4376C2FC52" lastPageId="15" lastPageNumber="290" pageId="11" pageNumber="286">
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<subSubSection id="3FE0EAB8FF947B51FF7A7C437665F872" pageId="11" pageNumber="286" type="nomenclature">
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<paragraph id="7745B933FF947B51FF7A7C4375D9F831" blockId="11.[151,662,1906,1933]" box="[151,662,1906,1933]" pageId="11" pageNumber="286">
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<heading id="2C0D0E5FFF947B51FF7A7C4375D9F831" bold="true" box="[151,662,1906,1933]" fontSize="11" level="1" pageId="11" pageNumber="286" reason="1">
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<taxonomicName id="B0FAC2B0FF947B51FF7A7C4375D9F831" authority="(Bleeker, 1860)" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[151,662,1906,1933]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="286" phylum="Chordata" rank="species" species="microlophus">
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<emphasis id="458E6521FF947B51FF7A7C4375D9F831" bold="true" box="[151,662,1906,1933]" pageId="11" pageNumber="286">
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<emphasis id="458E6521FF947B51FF7A7C43769AF830" bold="true" box="[151,469,1906,1933]" italics="true" pageId="11" pageNumber="286">Hydrosaurus microlophus</emphasis>
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(Bleeker, 1860)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="7745B933FF947B51FF7A7C877665F872" blockId="11.[151,685,1974,2030]" box="[151,298,1974,1999]" pageId="11" pageNumber="286">
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<heading id="2C0D0E5FFF947B51FF7A7C877665F872" bold="true" box="[151,298,1974,1999]" fontSize="10" level="2" pageId="11" pageNumber="286" reason="2">
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<emphasis id="458E6521FF947B51FF7A7C877665F872" bold="true" box="[151,298,1974,1999]" pageId="11" pageNumber="286">Chresonymy:</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3FE0EAB8FF947B56FF7A7CE775FDFEB0" lastPageId="12" lastPageNumber="287" pageId="11" pageNumber="286" type="reference_group">
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<paragraph id="7745B933FF947B51FF7A7CE775E2F853" blockId="11.[151,685,1974,2030]" box="[151,685,2005,2030]" pageId="11" pageNumber="286">
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<treatmentCitationGroup id="57EA9E1DFF947B51FF7A7CE775E2F853" box="[151,685,2005,2030]" pageId="11" pageNumber="286">
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<taxonomicName id="B0FAC2B0FF947B51FF7A7CE77626F850" ID-CoL="893WT" authorityName="Bleeker" authorityYear="1860" box="[151,361,2005,2029]" class="Squamata" family="Agamidae" genus="Istiurus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="286" phylum="Chordata" rank="species" species="microlophus">
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<emphasis id="458E6521FF947B51FF7A7CE77626F850" box="[151,361,2005,2029]" italics="true" pageId="11" pageNumber="286">Istiurus microlophus</emphasis>
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</taxonomicName>
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(
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<treatmentCitation id="F65B9F22FF947B51FE9B7CE4757BF850" author="Bleeker, P." box="[374,564,2005,2030]" page="80" pageId="11" pageNumber="286" year="1860">
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<bibRefCitation id="136BC4C2FF947B51FE9B7CE4757BF850" author="Bleeker, P." box="[374,564,2005,2030]" pageId="11" pageNumber="286" pagination="80 - 81" refId="ref16551" refString="Bleeker, P. (1860 a) Reptilien van Makassar. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 22, 80 - 81." type="journal article" year="1860">Bleeker 1860a: 80</bibRefCitation>
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</treatmentCitation>
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,
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<treatmentCitation id="F65B9F22FF947B51FDD37CE475EAF853" author="Bleeker, P." box="[574,677,2005,2030]" page="85" pageId="11" pageNumber="286" year="1860">
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<bibRefCitation id="136BC4C2FF947B51FDD37CE475EAF853" author="Bleeker, P." box="[574,677,2005,2030]" pageId="11" pageNumber="286" pagination="81 - 85" refId="ref16574" refString="Bleeker, P. (1860 b) Reptilien van Boni. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 22, 81 - 85." type="journal article" year="1860">1860b: 85</bibRefCitation>
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</treatmentCitation>
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)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="7745B933FF937B56FF7A7BA87534FF0C" blockId="12.[151,1149,152,269]" box="[151,635,152,177]" pageId="12" pageNumber="287">
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<treatmentCitationGroup id="57EA9E1DFF937B56FF7A7BA87534FF0C" box="[151,635,152,177]" pageId="12" pageNumber="287">
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<taxonomicName id="B0FAC2B0FF937B56FF7A7BA87625FF0C" authorityName="Cuvier" authorityYear="1829" box="[151,362,153,177]" class="Squamata" family="Agamidae" genus="Istiurus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="amboinensis">
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<emphasis id="458E6521FF937B56FF7A7BA87625FF0C" box="[151,362,153,177]" italics="true" pageId="12" pageNumber="287">Istiurus amboinensis</emphasis>
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</taxonomicName>
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(part) (
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<treatmentCitation id="F65B9F22FF937B56FE587BA9753CFF0C" author="Bleeker, P." box="[437,627,152,177]" page="85" pageId="12" pageNumber="287" year="1860">
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<bibRefCitation id="136BC4C2FF937B56FE587BA9753CFF0C" author="Bleeker, P." box="[437,627,152,177]" pageId="12" pageNumber="287" pagination="81 - 85" refId="ref16574" refString="Bleeker, P. (1860 b) Reptilien van Boni. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 22, 81 - 85." type="journal article" year="1860">Bleeker 1860b: 85</bibRefCitation>
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</treatmentCitation>
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)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="7745B933FF937B56FF7A7B867332FF72" blockId="12.[151,1149,152,269]" box="[151,1149,183,208]" pageId="12" pageNumber="287">
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<treatmentCitationGroup id="57EA9E1DFF937B56FF7A7B867332FF72" box="[151,1149,183,208]" pageId="12" pageNumber="287">
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<taxonomicName id="B0FAC2B0FF937B56FF7A7B86763BFF72" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[151,372,183,207]" class="Aves" family="Phasianidae" genus="Lophura" kingdom="Animalia" order="Galliformes" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="amboinensis">
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<emphasis id="458E6521FF937B56FF7A7B86763BFF72" box="[151,372,183,207]" italics="true" pageId="12" pageNumber="287">Lophura amboinensis</emphasis>
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</taxonomicName>
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(part) (
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<treatmentCitation id="F65B9F22FF937B56FE527B8675D5FF72" author="Boulenger, G. A." box="[447,666,183,208]" page="402" pageId="12" pageNumber="287" year="1885">
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<bibRefCitation id="136BC4C2FF937B56FE527B8675D5FF72" author="Boulenger, G. A." box="[447,666,183,208]" pageId="12" pageNumber="287" refId="ref16701" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. Trustees of the British Museum, London, 450 pp." type="book" year="1885">Boulenger 1885: 402</bibRefCitation>
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</treatmentCitation>
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; Casto de Elera 1895: 417;
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<treatmentCitation id="F65B9F22FF937B56FC5F7B867339FF72" author="de Rooij, N." box="[946,1142,183,208]" page="128" pageId="12" pageNumber="287" year="1915">
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<bibRefCitation id="136BC4C2FF937B56FC5F7B867339FF72" author="de Rooij, N." box="[946,1142,183,208]" pageId="12" pageNumber="287" refId="ref17185" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, xiv + 384 pp. https: // doi. org / 10.5962 / bhl. title. 5069" type="book" year="1915">de Rooij 1915: 128</bibRefCitation>
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</treatmentCitation>
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)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="7745B933FF937B56FF7A7BE7744AFF53" blockId="12.[151,1149,152,269]" box="[151,773,213,238]" pageId="12" pageNumber="287">
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<treatmentCitationGroup id="57EA9E1DFF937B56FF7A7BE7744AFF53" box="[151,773,213,238]" pageId="12" pageNumber="287">
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<taxonomicName id="B0FAC2B0FF937B56FF7A7BE77542FF53" authority="Weber 1890" box="[151,525,214,238]" class="Aves" family="Phasianidae" genus="Lophura" kingdom="Animalia" order="Galliformes" pageId="12" pageNumber="287" phylum="Chordata" rank="variety" species="amboinensis" variety="celebensis">
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<emphasis id="458E6521FF937B56FF7A7BE7763BFF53" box="[151,372,214,238]" italics="true" pageId="12" pageNumber="287">Lophura amboinensis</emphasis>
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var.
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<emphasis id="458E6521FF937B56FE487BE77542FF53" box="[421,525,214,238]" italics="true" pageId="12" pageNumber="287">celebensis</emphasis>
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</taxonomicName>
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(
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<treatmentCitation id="F65B9F22FF937B56FDF77BE47583FF53" author="Weber, M." box="[538,716,213,238]" page="160" pageId="12" pageNumber="287" year="1890">
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<bibRefCitation id="136BC4C2FF937B56FDF77BE47583FF53" author="Weber, M." box="[538,716,213,238]" pageId="12" pageNumber="287" pagination="159 - 177" refId="ref20299" refString="Weber, M. (1890) Reptilia from the Malay Archipelago. I. Sauria, Crocodilidae, Chelonia. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. E. J. Brill, Leiden, pp. 159 - 177. https: // doi. org / 10.5962 / bhl. title. 52289" type="book chapter" year="1890">Weber 1890: 160</bibRefCitation>
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</treatmentCitation>
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, 167)
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</paragraph>
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<paragraph id="7745B933FF937B56FF7A7BC475FDFEB0" blockId="12.[151,1149,152,269]" box="[151,690,244,269]" pageId="12" pageNumber="287">
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<treatmentCitationGroup id="57EA9E1DFF937B56FF7A7BC475FDFEB0" box="[151,690,244,269]" pageId="12" pageNumber="287">
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<taxonomicName id="B0FAC2B0FF937B56FF7A7BC476EFFEB0" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[151,416,245,269]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="amboinensis">
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<emphasis id="458E6521FF937B56FF7A7BC476EFFEB0" box="[151,416,245,269]" italics="true" pageId="12" pageNumber="287">Hydrosaurus amboinensis</emphasis>
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</taxonomicName>
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(part) (
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<treatmentCitation id="F65B9F22FF937B56FE067BC575E3FEB0" author="Wermuth, H." box="[491,684,244,269]" page="64" pageId="12" pageNumber="287" year="1967">
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<bibRefCitation id="136BC4C2FF937B56FE067BC575E3FEB0" author="Wermuth, H." box="[491,684,244,269]" pageId="12" pageNumber="287" pagination="1 - 127" refId="ref20367" refString="Wermuth, H. (1967) Liste der rezenten Amphibien und Reptilien. Agamidae. Das Tierreich, 86, i-xiv + 1 - 127." type="journal article" year="1967">Wermuth,1967: 64</bibRefCitation>
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</treatmentCitation>
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)
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="3FE0EAB8FF937B56FF7A7A0A73EEFEC4" pageId="12" pageNumber="287" type="materials_examined">
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<paragraph id="7745B933FF937B56FF7A7A0A7357FEE8" blockId="12.[151,1437,314,2034]" box="[151,1048,314,342]" pageId="12" pageNumber="287">
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<emphasis id="458E6521FF937B56FF7A7A0A7794FEE8" bold="true" box="[151,219,315,341]" pageId="12" pageNumber="287">Type:</emphasis>
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<materialsCitation id="C792B36EFF937B56FF0F7A0A7357FEE8" ID-GBIF-Occurrence="2573218757" box="[226,1048,314,342]" collectionCode="NHMUK" collectorName="P. Bleeker" pageId="12" pageNumber="287" specimenCode="NHMUK 1863.12" specimenCount="1" typeStatus="holotype">
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<typeStatus id="A8410791FF937B56FF0F7A0A7605FEE8" box="[226,330,315,341]" pageId="12" pageNumber="287" type="holotype">Holotype</typeStatus>
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<specimenCode id="275C1148FF937B56FEBC7A0D756EFEE8" box="[337,545,314,342]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="12" pageNumber="287" type="Museum">NHMUK 1863.12</specimenCode>
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.4.35, juvenile, collected by
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<collectorName id="DA0FDCE5FF937B56FCB77A0D7481FEE8" box="[858,974,315,342]" pageId="12" pageNumber="287">P. Bleeker</collectorName>
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, 1855.
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|
<materialsCitation id="C792B36EFF937B56FF2A7A6E73D2FEC4" ID-GBIF-Occurrence="2573218760" box="[199,1181,350,378]" country="Indonesia" location="Makassar" municipality="Southwest" pageId="12" pageNumber="287" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
|
<emphasis id="458E6521FF937B56FF2A7A6E7626FEC4" bold="true" box="[199,361,350,377]" pageId="12" pageNumber="287">
|
|
<typeStatus id="A8410791FF937B56FF2A7A6E764DFEC4" box="[199,258,351,377]" pageId="12" pageNumber="287">Type</typeStatus>
|
|
locality:
|
|
</emphasis>
|
|
<location id="7225EFE8FF937B56FE9D7A6E7694FEC4" LSID="urn:lsid:plazi:treatment:FF530825FF947B55FF7A7C4376C2FC52:7225EFE8FF937B56FE9D7A6E7694FEC4" box="[368,475,351,377]" country="Indonesia" municipality="Southwest" name="Makassar" pageId="12" pageNumber="287" stateProvince="Sulawesi Utara">Makassar</location>
|
|
(= Ujung Pandang),
|
|
<collectingMunicipality id="97212349FF937B56FD2A7A6E7473FEC4" box="[711,828,351,377]" pageId="12" pageNumber="287">Southwest</collectingMunicipality>
|
|
<collectingRegion id="B53E77D1FF937B56FCAE7A6E74E4FEC4" box="[835,939,351,377]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="287">Sulawesi</collectingRegion>
|
|
,
|
|
<collectingCountry id="0FEDF9A3FF937B56FC5B7A6E736CFEC4" box="[950,1059,351,377]" name="Indonesia" pageId="12" pageNumber="287">Indonesia</collectingCountry>
|
|
(
|
|
<figureCitation id="EFC1A5B6FF937B56FBDF7A6E73DCFEC4" box="[1074,1171,351,377]" captionStart="FIGURE 8" captionStartId="18.[151,250,1846,1871]" captionTargetBox="[151,1436,524,1826]" captionTargetId="figure@18.[151,1436,524,1829]" captionTargetPageId="18" captionText="FIGURE 8. Map of Sulawesi showing the distribution of H. microlophus (red) and H. celebensis (grey). Type localities are indicated by stars.. 0 Makassar; 1 Bantimurung; 2 Pattunuang; 3 Bengo-Bengo & Rompegading; 4 Camba; 5 Ujung Lamuru; 6 Belopa; 7 Palopo-Pompengan & Palopo-Siguntu; 8 Toraja; 9 Malele; 10 Poso River, Bay of Tomini; 11 Desa Jelaja; 12 Wasponda; 13 Desa Limbatu; 14 Morowali Nature Reserve; W1 Tempe („at the Minralang river near Tempe“ after Weber1890: 167); W2 Pampanua („at Pampanua on the river Tjinrana“ after Weber 1890: 167)" figureDoi="http://doi.org/10.5281/zenodo.3694822" httpUri="https://zenodo.org/record/3694822/files/figure.png" pageId="12" pageNumber="287">Figure 8</figureCitation>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF937B56FF2A7AB37527FE20" box="[199,616,386,413]" pageId="12" pageNumber="287" type="description">
|
|
<paragraph id="7745B933FF937B56FF2A7AB37527FE20" blockId="12.[151,1437,314,2034]" box="[199,616,386,413]" pageId="12" pageNumber="287">
|
|
<emphasis id="458E6521FF937B56FF2A7AB376A8FE20" bold="true" box="[199,487,386,413]" pageId="12" pageNumber="287">
|
|
Description of
|
|
<typeStatus id="A8410791FF937B56FE9A7AB276ADFE20" box="[375,482,387,413]" pageId="12" pageNumber="287" type="holotype">holotype</typeStatus>
|
|
:
|
|
</emphasis>
|
|
See above.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF937B55FF2A7A9776C1FE4A" lastPageId="15" lastPageNumber="290" pageId="12" pageNumber="287" type="diagnosis">
|
|
<paragraph id="7745B933FF937B56FF2A7A977374FCF0" blockId="12.[151,1437,314,2034]" pageId="12" pageNumber="287">
|
|
<emphasis id="458E6521FF937B56FF2A7A97760BFE7C" bold="true" box="[199,324,422,449]" pageId="12" pageNumber="287">Diagnosis:</emphasis>
|
|
Large species exceeding
|
|
<quantity id="B00214D6FF937B56FD857A96759BFE7F" box="[616,724,423,450]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="287" unit="mm" value="1000.0">1000 mm</quantity>
|
|
in total length; largest specimen, SMF 35996, SVL =
|
|
<quantity id="B00214D6FF937B56FAAD7A9772D7FE7C" box="[1344,1432,422,449]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" pageId="12" pageNumber="287" unit="mm" value="325.0">325mm</quantity>
|
|
, TL =
|
|
<quantity id="B00214D6FF937B56FF3A7AFB7662FE58" box="[215,301,458,485]" metricMagnitude="-1" metricUnit="m" metricValue="7.45" pageId="12" pageNumber="287" unit="mm" value="745.0">745mm</quantity>
|
|
(TL/SVL = 2.29), tip of the tail missing; complete specimen, MZB Lac 5870, SVL
|
|
<quantity id="B00214D6FF937B56FB327AFA7271FE5B" box="[1247,1342,459,486]" metricMagnitude="-1" metricUnit="m" metricValue="1.31" pageId="12" pageNumber="287" unit="mm" value="131.0">131 mm</quantity>
|
|
, TL
|
|
<quantity id="B00214D6FF937B56FA9F7AFB778CFDB7" metricMagnitude="-1" metricUnit="m" metricValue="3.15" pageId="12" pageNumber="287" unit="mm" value="315.0">315 mm</quantity>
|
|
(TL/SVL = 2.4) [Remark: The MVZ database gives
|
|
<quantity id="B00214D6FF937B56FCF67ADE7437FDB7" box="[795,888,495,522]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="12" pageNumber="287" unit="mm" value="370.0">370 mm</quantity>
|
|
as tail length]. Head black; sometimes with yellowish colour around the eyes, the same on the lower half of the neck and in front of the shoulders as well as in the gular region; nuchal region black; a group of enlarged conical or pyramidal scales on either side of the neck; nuchal and dorsal crests continuous; in males, the sides of the body are dirty yellow with three groups of enlarged conical or pyramidal scales, often forming a triangular shape, between fore and hindlimbs that decrease in size ventrally to form transverse bands, the largest scales being black in colour; adult females are black with yellow enlarged scales; some females (presumably subadults or non-reproducing specimens) and juveniles typically grey with the groups of enlarged scales being whitish in colour; yellow ventrally, limbs on the upper side black and on the underside yellow (grey in subadults and juveniles); scales under fourth and fifth toes initially flat or singly keeled; tail black with some irregular yellow markings; sail dirty yellow to light brown with black stripes.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF937B56FF2A78677461FC63" blockId="12.[151,1437,314,2034]" pageId="12" pageNumber="287">
|
|
<emphasis id="458E6521FF937B56FF2A786774B3FCCC" bold="true" box="[199,1020,854,881]" pageId="12" pageNumber="287">
|
|
Allocation of adult museum specimens to
|
|
<taxonomicName id="B0FAC2B0FF937B56FD28786774B9FCCC" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[709,1014,854,881]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF937B56FD28786774B9FCCC" bold="true" box="[709,1014,854,881]" italics="true" pageId="12" pageNumber="287">Hydrosaurus microlophus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
The preserved type material is only represented by a single specimen each. In the case of
|
|
<taxonomicName id="B0FAC2B0FF937B56FD1F784D74C7FC28" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[754,904,891,918]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF937B56FD1F784D7449FC28" box="[754,774,892,917]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FCF8784A74C7FC28" box="[789,904,891,917]" italics="true" pageId="12" pageNumber="287">celebensis</emphasis>
|
|
</taxonomicName>
|
|
we located additional adult voucher specimens that match the
|
|
<typeStatus id="A8410791FF937B56FED278AE76EEFC04" box="[319,417,927,953]" pageId="12" pageNumber="287" type="holotype">holotype</typeStatus>
|
|
of this species (see Appendix 1). However, we were not able to locate any juvenile specimens that can be assigned unambiguously to
|
|
<taxonomicName id="B0FAC2B0FF937B56FD7C78F57468FC60" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[657,807,963,990]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF937B56FD7C78F575EAFC60" box="[657,677,964,989]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FD5978F27468FC60" box="[692,807,963,989]" italics="true" pageId="12" pageNumber="287">celebensis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF937B56FF2A78D975A7FB2F" blockId="12.[151,1437,314,2034]" pageId="12" pageNumber="287">
|
|
However, in the case of
|
|
<taxonomicName id="B0FAC2B0FF937B56FE3F78D97530FBBC" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[466,639,999,1025]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF937B56FE3F78D97530FBBC" box="[466,639,999,1025]" italics="true" pageId="12" pageNumber="287">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
the
|
|
<typeStatus id="A8410791FF937B56FD4078D67440FBBC" box="[685,783,999,1025]" pageId="12" pageNumber="287" type="holotype">holotype</typeStatus>
|
|
is a juvenile and corresponding adult specimens present in museum collections had to be inferred as belonging to this species following morphological comparison. The main diagnostic characters for assignment of adult specimens to
|
|
<taxonomicName id="B0FAC2B0FF937B56FCD47F0174A7FBF4" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[825,1000,1071,1098]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF937B56FCD47F017402FBF4" box="[825,845,1072,1097]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FCB07F1E74A7FBF4" box="[861,1000,1071,1097]" italics="true" pageId="12" pageNumber="287">microlophus</emphasis>
|
|
</taxonomicName>
|
|
is the presence of groups of enlarged, strongly keeled scales dorsolaterally and the highly variable number of enlarged scales on the sides of the neck (≥ 3 up to eleven) as well as their shape and arrangement.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF937B56FF2A7FAA7625F880" blockId="12.[151,1437,314,2034]" pageId="12" pageNumber="287">
|
|
The Senckenberg collection holds three specimens (SMF 35996-98) from
|
|
<collectingRegion id="B53E77D1FF937B56FBE47FAA7321FB08" box="[1033,1134,1179,1205]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="287">Sulawesi</collectingRegion>
|
|
without precise locality or collector details. These specimens were received in 1930 from the Zoological Society Frankfurt and determined under the curatorship of Robert Mertens as
|
|
<taxonomicName id="B0FAC2B0FF937B56FD657FD57484FB40" box="[648,971,1251,1277]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="subSpecies" species="amboinensis" subSpecies="microlophus">
|
|
<emphasis id="458E6521FF937B56FD657FD57484FB40" box="[648,971,1251,1277]" italics="true" pageId="12" pageNumber="287">H. amboinensis microlophus</emphasis>
|
|
</taxonomicName>
|
|
. All three specimens agree well with the characters outlined as specific for the
|
|
<taxonomicName id="B0FAC2B0FF937B56FDD57E3975ABFA9C" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[568,740,1287,1314]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF937B56FDD57E397503FA9C" box="[568,588,1288,1313]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FDB47E3675ABFA9C" box="[601,740,1287,1313]" italics="true" pageId="12" pageNumber="287">microlophus</emphasis>
|
|
</taxonomicName>
|
|
type (i.e., enlarged dorsal scales in groups, ventral scales equal or larger than dorsal scales, 10/11 supra- and infralabials, respectively). Therefore, we consider the Senckenberg material adult representatives of
|
|
<taxonomicName id="B0FAC2B0FF937B56FDEB7E6175FCFAD4" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[518,691,1359,1386]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF937B56FDEB7E617555FAD4" box="[518,538,1360,1385]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FDC57E7E75FCFAD4" box="[552,691,1359,1385]" italics="true" pageId="12" pageNumber="287">microlophus</emphasis>
|
|
</taxonomicName>
|
|
(see also below for a comparison of live specimens). Two of these specimens lack parts of their tails but one has a nearly complete tail (SVL =
|
|
<quantity id="B00214D6FF937B56FC0E7E437374FA30" box="[995,1083,1394,1421]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" pageId="12" pageNumber="287" unit="mm" value="325.0">325mm</quantity>
|
|
, TL =
|
|
<quantity id="B00214D6FF937B56FB6E7E437393FA30" box="[1155,1244,1394,1421]" metricMagnitude="-1" metricUnit="m" metricValue="7.45" pageId="12" pageNumber="287" unit="mm" value="745.0">745mm</quantity>
|
|
, with only the tip of the tail missing according to G. Köhler [SMF, pers. comm.], TL/SVL = 2.3). Currently the specimens, although kept in the same jar, are not individually numbered. For clarity we consider the nearly complete specimen as SMF 35996, the specimen measuring SVL =
|
|
<quantity id="B00214D6FF937B56FDBC7EEF75E5FA44" box="[593,682,1502,1529]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" pageId="12" pageNumber="287" unit="mm" value="295.0">295mm</quantity>
|
|
, TL =
|
|
<quantity id="B00214D6FF937B56FD1E7EEE7475FA44" box="[755,826,1503,1529]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="287" unit="mm" value="90.0">90mm</quantity>
|
|
as SMF 35997 and the one measuring SVL =
|
|
<quantity id="B00214D6FF937B56FAAD7EEE72D7FA44" box="[1344,1432,1503,1529]" metricMagnitude="-1" metricUnit="m" metricValue="3.3299999999999996" pageId="12" pageNumber="287" unit="mm" value="333.0">333mm</quantity>
|
|
, TL =
|
|
<quantity id="B00214D6FF937B56FF3A7D32767AF9A3" box="[215,309,1539,1566]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="287" unit="mm" value="220.0">220 mm</quantity>
|
|
as SMF 35998.
|
|
<bibRefCitation id="136BC4C2FF937B56FE027D3275DDF9A0" author="Gaulke, M." box="[495,658,1539,1565]" pageId="12" pageNumber="287" pagination="6 - 12" refId="ref17586" refString="Gaulke, M. (1989) Einige Bemerkungen uber die philippinische Segelechse Hydrosaurus pustulatus (Eschscholtz, 1829). Herpetofauna, Weinstadt, 11 (62) 6 - 12." type="journal article" year="1989">Gaulke (1989)</bibRefCitation>
|
|
examined these specimens and concluded that they were conspecific with
|
|
<emphasis id="458E6521FF937B56FF397D1977A7F9FC" box="[212,232,1576,1601]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
. (cf.)
|
|
<taxonomicName id="B0FAC2B0FF937B56FEDE7D1676E9F9FC" baseAuthorityName="Eschscholtz" baseAuthorityYear="1829" box="[307,422,1575,1601]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="pustulatus">
|
|
<emphasis id="458E6521FF937B56FEDE7D1676E9F9FC" box="[307,422,1575,1601]" italics="true" pageId="12" pageNumber="287">pustulatus</emphasis>
|
|
</taxonomicName>
|
|
unaware that the Philippine populations of
|
|
<taxonomicName id="B0FAC2B0FF937B56FC457D167376F9FC" authorityName="Kaup" authorityYear="1828" box="[936,1081,1575,1601]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="genus">
|
|
<emphasis id="458E6521FF937B56FC457D167376F9FC" box="[936,1081,1575,1601]" italics="true" pageId="12" pageNumber="287">Hydrosaurus</emphasis>
|
|
</taxonomicName>
|
|
constituted a species complex (
|
|
<bibRefCitation id="136BC4C2FF937B56FF727D7A7611F9D8" author="Siler, C. D. & Lira-Noriega, A. & Brown, R. M." box="[159,350,1611,1638]" pageId="12" pageNumber="287" pagination="100 - 108" refId="ref20070" refString="Siler, C. D., Lira-Noriega, A. & Brown, R. M. (2014) Conservation genetics of Australasian sailfin lizards: Flagship species threatened by coastal development and insufficient protected area coverage. Biological Conservation, 169, 100 - 108 https: // doi. org / 10.1016 / j. biocon. 2013.10.014" type="journal article" year="2014">
|
|
Siler
|
|
<emphasis id="458E6521FF937B56FF337D7D765EF9D8" box="[222,273,1611,1637]" italics="true" pageId="12" pageNumber="287">et al</emphasis>
|
|
. 2014
|
|
</bibRefCitation>
|
|
). A comparison with the juvenile type shows that the enlarged dorsal scales have a much more pronounced keel in adults and that these may exhibit a conical or near pyramidal shape. In the juvenile
|
|
<typeStatus id="A8410791FF937B56FAD77D5E72D3F934" box="[1338,1436,1647,1673]" pageId="12" pageNumber="287" type="holotype">holotype</typeStatus>
|
|
the enlarged dorsal scales are white while they appear to be brown (SMF 35997), dark brown or black (SMF 35996 and SMF 35998) in the preserved adults. This is in contrast to the black and yellow speckled dorsal colouration in
|
|
<taxonomicName id="B0FAC2B0FF937B56FF7A7DED7661F948" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[151,302,1755,1782]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF937B56FF7A7DED77E4F948" box="[151,171,1756,1781]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FF567DEA7661F948" box="[187,302,1755,1781]" italics="true" pageId="12" pageNumber="287">celebensis</emphasis>
|
|
</taxonomicName>
|
|
. Additionally, the enlarged dorsal scales in
|
|
<taxonomicName id="B0FAC2B0FF937B56FCCD7DED74F8F948" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[800,951,1755,1782]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF937B56FCCD7DED747BF948" box="[800,820,1756,1781]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FCA97DEA74F8F948" box="[836,951,1755,1781]" italics="true" pageId="12" pageNumber="287">celebensis</emphasis>
|
|
</taxonomicName>
|
|
are typically light coloured or even white. A comparison between the type specimen of
|
|
<taxonomicName id="B0FAC2B0FF937B56FD7E7C317466F8A4" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[659,809,1791,1818]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF937B56FD7E7C3175E8F8A4" box="[659,679,1792,1817]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FD5B7DCE7466F8A4" box="[694,809,1791,1817]" italics="true" pageId="12" pageNumber="287">celebensis</emphasis>
|
|
</taxonomicName>
|
|
and an adult
|
|
<taxonomicName id="B0FAC2B0FF937B56FC2E7C31733FF8A4" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[963,1136,1791,1818]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF937B56FC2E7C317498F8A4" box="[963,983,1792,1817]" italics="true" pageId="12" pageNumber="287">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF937B56FC087DCE733FF8A4" box="[997,1136,1791,1817]" italics="true" pageId="12" pageNumber="287">microlophus</emphasis>
|
|
</taxonomicName>
|
|
specimen (SMF 35996) is shown in
|
|
<figureCitation id="EFC1A5B6FF937B56FEE97C12762BF880" box="[260,356,1826,1853]" captionStart="FIGURE 5" captionStartId="13.[151,250,1979,2004]" captionTargetBox="[197,1391,174,1960]" captionTargetId="figure@13.[197,1391,174,1960]" captionTargetPageId="13" captionText="FIGURE 5. Comparison of dorsal scalation between the type specimen of H. celebensis (A, ZMB 7393) and an adult specimen of H. microlophus (B, SMF 35996)." figureDoi="http://doi.org/10.5281/zenodo.3694814" httpUri="https://zenodo.org/record/3694814/files/figure.png" pageId="12" pageNumber="287">Figure 5</figureCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF937B54FF2A7C76762EFF6A" blockId="12.[151,1437,314,2034]" lastBlockId="14.[151,1437,153,467]" lastPageId="14" lastPageNumber="289" pageId="12" pageNumber="287">
|
|
<bibRefCitation id="136BC4C2FF937B56FF2A7C7676CFF8DF" author="Bleeker, P." box="[199,384,1863,1890]" pageId="12" pageNumber="287" pagination="81 - 85" refId="ref16574" refString="Bleeker, P. (1860 b) Reptilien van Boni. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 22, 81 - 85." type="journal article" year="1860">Bleeker (1860b)</bibRefCitation>
|
|
reported specimens from “Boni” as
|
|
<taxonomicName id="B0FAC2B0FF937B56FCCB7C797433F8DC" authorityName="Cuvier" authorityYear="1829" box="[806,892,1864,1889]" class="Squamata" family="Agamidae" genus="Istiurus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="287" phylum="Chordata" rank="genus">
|
|
<emphasis id="458E6521FF937B56FCCB7C797433F8DC" box="[806,892,1864,1889]" italics="true" pageId="12" pageNumber="287">Istiurus</emphasis>
|
|
</taxonomicName>
|
|
(=
|
|
<taxonomicName id="B0FAC2B0FF937B56FC4A7C767377F8DC" authorityName="Kaup" authorityYear="1828" box="[935,1080,1863,1889]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="genus">
|
|
<emphasis id="458E6521FF937B56FC4A7C767377F8DC" box="[935,1080,1863,1889]" italics="true" pageId="12" pageNumber="287">Hydrosaurus</emphasis>
|
|
</taxonomicName>
|
|
)
|
|
<taxonomicName id="B0FAC2B0FF937B56FBA67C767398F8DC" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[1099,1239,1863,1889]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="287" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF937B56FBA67C767398F8DC" box="[1099,1239,1863,1889]" italics="true" pageId="12" pageNumber="287">amboinensis</emphasis>
|
|
</taxonomicName>
|
|
. There is a dried specimen at Naturalis (ZMA 12611) that was previously housed in the collection of the Tropeninstituut Amsterdam, and subsequently transferred to the ZMA collection (now at Naturalis). Boni (or Bone State) was a former sultanate on the east coast of the southwestern arm of
|
|
<collectingRegion id="B53E77D1FF937B56FD547C827451F870" box="[697,798,1971,1997]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="287">Sulawesi</collectingRegion>
|
|
near Danau (=Lake) Tempe. The specimen has enlarged conical, near pyramidal scales on either side of the neck and two groups of similar scales on the back. As its dorsal scalation matches that of
|
|
<taxonomicName id="B0FAC2B0FF917B54FE557BAB7528FF0E" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[440,615,153,179]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF917B54FE557BAB7683FF0E" box="[440,460,154,179]" italics="true" pageId="14" pageNumber="289">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF917B54FE317BA87528FF0E" box="[476,615,153,179]" italics="true" pageId="14" pageNumber="289">microlophus</emphasis>
|
|
</taxonomicName>
|
|
and the locality record falls within the range of this species it should be classified as such.
|
|
</paragraph>
|
|
<caption id="2385E9BBFF927B57FF7A7C8A754BF845" ID-DOI="http://doi.org/10.5281/zenodo.3694814" ID-Zenodo-Dep="3694814" httpUri="https://zenodo.org/record/3694814/files/figure.png" pageId="13" pageNumber="288" startId="13.[151,250,1979,2004]" targetBox="[197,1391,174,1960]" targetPageId="13">
|
|
<paragraph id="7745B933FF927B57FF7A7C8A754BF845" blockId="13.[151,1437,1979,2040]" pageId="13" pageNumber="288">
|
|
<emphasis id="458E6521FF927B57FF7A7C8A765DF869" bold="true" box="[151,274,1979,2004]" pageId="13" pageNumber="288">FIGURE 5.</emphasis>
|
|
Comparison of dorsal scalation between the type specimen of
|
|
<taxonomicName id="B0FAC2B0FF927B57FC677C8C735EF869" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[906,1041,1980,2004]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="288" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF927B57FC677C8C74D3F869" box="[906,924,1981,2004]" italics="true" pageId="13" pageNumber="288">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF927B57FC447C8D735EF869" box="[937,1041,1980,2004]" italics="true" pageId="13" pageNumber="288">celebensis</emphasis>
|
|
</taxonomicName>
|
|
(A, ZMB 7393) and an adult specimen of
|
|
<taxonomicName id="B0FAC2B0FF927B57FF5E7CD0761FF845" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[179,336,2016,2040]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="288" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF927B57FF5E7CD0778AF845" box="[179,197,2017,2040]" italics="true" pageId="13" pageNumber="288">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF927B57FF3F7CD1761FF845" box="[210,336,2016,2040]" italics="true" pageId="13" pageNumber="288">microlophus</emphasis>
|
|
</taxonomicName>
|
|
(B, SMF 35996).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="7745B933FF917B54FF2A7BD177AFFE6E" blockId="14.[151,1437,153,467]" pageId="14" pageNumber="289">
|
|
<emphasis id="458E6521FF917B54FF2A7BD176AEFF46" bold="true" box="[199,481,224,251]" pageId="14" pageNumber="289">Intraspecific Variation:</emphasis>
|
|
One variable character is the number of enlarged scales on either side of the neck. The enlarged scales along the neck are sometimes arranged in a single row and sometimes intermixed with slightly smaller ones forming a group (picture collection AGT). The number of enlarged scales varies from three to eleven. The lateral groups of enlarged body scales consist of 10 to 20 scales, with the anterior most group having the highest number of scales. Typically there are three groups of enlarged scales between the fore and hindlimbs. One specimen (picture collection AGT) contains a group of three enlarged scales between the midbody and the onset of the tail group.
|
|
</paragraph>
|
|
<caption id="2385E9BBFF917B54FF7A7EF87551F997" ID-DOI="http://doi.org/10.5281/zenodo.3694816" ID-Zenodo-Dep="3694816" httpUri="https://zenodo.org/record/3694816/files/figure.png" pageId="14" pageNumber="289" startId="14.[151,250,1481,1506]" targetBox="[151,1435,501,1453]" targetPageId="14">
|
|
<paragraph id="7745B933FF917B54FF7A7EF87551F997" blockId="14.[151,1437,1481,1578]" pageId="14" pageNumber="289">
|
|
<emphasis id="458E6521FF917B54FF7A7EF8765CFA5F" bold="true" box="[151,275,1481,1506]" pageId="14" pageNumber="289">FIGURE 6.</emphasis>
|
|
Portraits of male sailfin lizards. A)
|
|
<taxonomicName id="B0FAC2B0FF917B54FD937EFB7453FA5C" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[638,796,1481,1505]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF917B54FD937EFB7453FA5C" box="[638,796,1481,1505]" italics="true" pageId="14" pageNumber="289">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
from Air besar, Ambon B)
|
|
<taxonomicName id="B0FAC2B0FF917B54FBDF7EFB73F6FA5C" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[1074,1209,1481,1505]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF917B54FBDF7EFB73F6FA5C" box="[1074,1209,1481,1505]" italics="true" pageId="14" pageNumber="289">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
from Pattununang; C)
|
|
<taxonomicName id="B0FAC2B0FF917B54FF7A7EDF767CF9B8" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[151,307,1517,1541]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF917B54FF7A7EDF767CF9B8" box="[151,307,1517,1541]" italics="true" pageId="14" pageNumber="289">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
from Bengo Bengo. Please note the oval nostril and absence of a nose crest in
|
|
<taxonomicName id="B0FAC2B0FF917B54FBAF7EDF7390F9B8" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[1090,1247,1517,1541]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF917B54FBAF7EDF7390F9B8" box="[1090,1247,1517,1541]" italics="true" pageId="14" pageNumber="289">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
. Photos by Andrea Glässer-Trobisch & Dietmar Trobisch.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="7745B933FF917B54FF2A7D66750EF89B" blockId="14.[151,1437,1623,2010]" pageId="14" pageNumber="289">
|
|
<emphasis id="458E6521FF917B54FF2A7D6676F3F9CF" bold="true" box="[199,444,1623,1650]" pageId="14" pageNumber="289">Sexual dimorphism:</emphasis>
|
|
Adult males of
|
|
<taxonomicName id="B0FAC2B0FF917B54FD967D687464F9CF" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[635,811,1624,1650]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF917B54FD967D687464F9CF" box="[635,811,1624,1650]" italics="true" pageId="14" pageNumber="289">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
have a considerably higher dorsal sail and a typically more pronounced nasal crest than do females, although we also observed some females with a well-developed nasal crest. In addition, males and females show a clear sexual dimorphic colouration (dichromatism, see above). Thus, in female specimens the groups of enlarged scales are whitish/yellowish coloured on a dark grey to nearly black dorsal ground colour while male specimens typically show a dirty yellow dorsal colouration with the groups of enlarged scales being black or brown in colour.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF917B55FF2A7C1E76C1FE4A" blockId="14.[151,1437,1623,2010]" lastBlockId="15.[151,1437,153,1007]" lastPageId="15" lastPageNumber="290" pageId="14" pageNumber="289">
|
|
<emphasis id="458E6521FF917B54FF2A7C1E7620F8F7" bold="true" box="[199,367,1839,1866]" pageId="14" pageNumber="289">Comparisons:</emphasis>
|
|
<taxonomicName id="B0FAC2B0FF917B54FE977C007564F8F7" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[378,555,1840,1866]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF917B54FE977C007564F8F7" box="[378,555,1840,1866]" italics="true" pageId="14" pageNumber="289">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="B0FAC2B0FF917B54FD277C007433F8F7" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[714,892,1840,1866]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF917B54FD277C007433F8F7" box="[714,892,1840,1866]" italics="true" pageId="14" pageNumber="289">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
by the possession of the following characters: grossly enlarged scales near the onset of the forelimbs; strongly enlarged conical or pyramidal scales in groups forming dorsolateral bands (all of these characters are either not or only slightly developed in
|
|
<taxonomicName id="B0FAC2B0FF917B54FB457C487219F82F" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[1192,1366,1912,1938]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF917B54FB457C4873F3F82F" box="[1192,1212,1913,1938]" italics="true" pageId="14" pageNumber="289">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF917B54FB277C497219F82F" box="[1226,1366,1912,1938]" italics="true" pageId="14" pageNumber="289">amboinensis</emphasis>
|
|
</taxonomicName>
|
|
). Furthermore,
|
|
<taxonomicName id="B0FAC2B0FF917B54FEE77CAC76F8F80B" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[266,439,1948,1974]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF917B54FEE77CAC76F8F80B" box="[266,439,1948,1974]" italics="true" pageId="14" pageNumber="289">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="B0FAC2B0FF917B54FDA67CAC75B6F80B" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[587,761,1948,1974]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF917B54FDA67CAC75B6F80B" box="[587,761,1948,1974]" italics="true" pageId="14" pageNumber="289">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
in colouration (yellowish green in male
|
|
<taxonomicName id="B0FAC2B0FF917B54FB527CAC7222F80B" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[1215,1389,1948,1974]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="289" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF917B54FB527CAC7222F80B" box="[1215,1389,1948,1974]" italics="true" pageId="14" pageNumber="289">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
and green in females with black marbling in both sexes vs. dirty yellow dorsal colouration with the groups of enlarged scales being dark coloured in male
|
|
<taxonomicName id="B0FAC2B0FF907B55FDCE7BAB759FFF0E" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[547,720,153,179]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FDCE7BAB759FFF0E" box="[547,720,153,179]" italics="true" pageId="15" pageNumber="290">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
, plus dark dorsal colouration with the groups of enlarged scales being whitish/yellowish in female
|
|
<taxonomicName id="B0FAC2B0FF907B55FDF27B8F7582FF6A" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[543,717,189,215]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FDF27B8F7582FF6A" box="[543,717,189,215]" italics="true" pageId="15" pageNumber="290">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
together with the top of the head and neck being black and the throat being yellow in colour in both sexes) they also possess a nasal crest (absent in
|
|
<taxonomicName id="B0FAC2B0FF907B55FB857BD37258FF46" baseAuthorityName="Schlosser" baseAuthorityYear="1768" box="[1128,1303,225,251]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="amboinensis">
|
|
<emphasis id="458E6521FF907B55FB857BD37258FF46" box="[1128,1303,225,251]" italics="true" pageId="15" pageNumber="290">H. amboinensis</emphasis>
|
|
</taxonomicName>
|
|
, see
|
|
<figureCitation id="EFC1A5B6FF907B55FABE7BD077FAFE9D" captionStart="FIGURE 6" captionStartId="14.[151,250,1481,1506]" captionTargetBox="[151,1435,501,1453]" captionTargetId="figure@14.[151,1435,501,1453]" captionTargetPageId="14" captionText="FIGURE 6. Portraits of male sailfin lizards. A) H. amboinensis from Air besar, Ambon B) H. celebensis from Pattununang; C) H. microlophus from Bengo Bengo. Please note the oval nostril and absence of a nose crest in H. amboinensis. Photos by Andrea Glässer-Trobisch & Dietmar Trobisch." figureDoi="http://doi.org/10.5281/zenodo.3694816" httpUri="https://zenodo.org/record/3694816/files/figure.png" pageId="15" pageNumber="290">Figure 6a</figureCitation>
|
|
). Most noticeably, the males of
|
|
<taxonomicName id="B0FAC2B0FF907B55FDCD7A377581FEA2" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[544,718,261,287]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FDCD7A377581FEA2" box="[544,718,261,287]" italics="true" pageId="15" pageNumber="290">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
differ in their pattern from the males of
|
|
<taxonomicName id="B0FAC2B0FF907B55FB767A37727EFEA2" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[1179,1329,261,287]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF907B55FB767A37727EFEA2" box="[1179,1329,261,287]" italics="true" pageId="15" pageNumber="290">H. celebensis</emphasis>
|
|
</taxonomicName>
|
|
the latter having complete or near complete black heads, limbs and tails.
|
|
<taxonomicName id="B0FAC2B0FF907B55FC817A1B7354FEFE" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[876,1051,297,323]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FC817A1B7354FEFE" box="[876,1051,297,323]" italics="true" pageId="15" pageNumber="290">H. microlophus</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="B0FAC2B0FF907B55FB5E7A1B726CFEFE" authorityName="Barbour" authorityYear="1911" box="[1203,1315,297,323]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF907B55FB5E7A1B726CFEFE" box="[1203,1315,297,323]" italics="true" pageId="15" pageNumber="290">H. weberi</emphasis>
|
|
</taxonomicName>
|
|
in general colouration and pattern, the latter being grey green.
|
|
<taxonomicName id="B0FAC2B0FF907B55FD017A7F74D3FEDA" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[748,924,333,359]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FD017A7F744FFEDA" box="[748,768,334,359]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FCFC7A7C74D3FEDA" box="[785,924,333,359]" italics="true" pageId="15" pageNumber="290">microlophus</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="B0FAC2B0FF907B55FBD87A7F7383FEDA" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[1077,1228,333,359]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF907B55FBD87A7F7306FEDA" box="[1077,1097,334,359]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FBB47A7C7383FEDA" box="[1113,1228,333,359]" italics="true" pageId="15" pageNumber="290">celebensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="B0FAC2B0FF907B55FAEA7A7F7238FEDA" authorityName="Barbour" authorityYear="1911" box="[1287,1399,333,359]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF907B55FAEA7A7F7254FEDA" box="[1287,1307,334,359]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FAC67A7C7238FEDA" box="[1323,1399,333,359]" italics="true" pageId="15" pageNumber="290">weberi</emphasis>
|
|
</taxonomicName>
|
|
by having groups of enlarged (black in males, white or yellow in females) conical and pyramidal scales on the dorsum. In every case the number of scales in the major groups of enlarged scales is larger than that in
|
|
<taxonomicName id="B0FAC2B0FF907B55FB537AA7721BFE12" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[1214,1364,405,431]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="celebensis">
|
|
<emphasis id="458E6521FF907B55FB537AA7739DFE12" box="[1214,1234,406,431]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FB0C7AA4721BFE12" box="[1249,1364,405,431]" italics="true" pageId="15" pageNumber="290">celebensis</emphasis>
|
|
</taxonomicName>
|
|
. In
|
|
<taxonomicName id="B0FAC2B0FF907B55FA6C7AA7766DFE6E" baseAuthorityName="Bleeker" baseAuthorityYear="1860" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FA6C7AA772DAFE12" box="[1409,1429,406,431]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FF7A7A88766DFE6E" box="[151,290,441,467]" italics="true" pageId="15" pageNumber="290">microlophus</emphasis>
|
|
</taxonomicName>
|
|
the nuchal and dorsal crests are continuous, while they are separated by considerably smaller scales on the neck in
|
|
<taxonomicName id="B0FAC2B0FF907B55FEF47AEF76C8FE4A" authorityName="Barbour" authorityYear="1911" box="[281,391,477,503]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="weberi">
|
|
<emphasis id="458E6521FF907B55FEF47AEF7662FE4A" box="[281,301,478,503]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FED67AEC76C8FE4A" box="[315,391,477,503]" italics="true" pageId="15" pageNumber="290">weberi</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF907B55FF2A793173B3FC3E" pageId="15" pageNumber="290" type="biology_ecology">
|
|
<paragraph id="7745B933FF907B55FF2A7931727CFD3A" blockId="15.[151,1437,153,1007]" pageId="15" pageNumber="290">
|
|
<emphasis id="458E6521FF907B55FF2A79317664FDA6" bold="true" box="[199,299,512,539]" pageId="15" pageNumber="290">Biology:</emphasis>
|
|
<taxonomicName id="B0FAC2B0FF907B55FEDE793376ADFDA6" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[307,482,513,539]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FEDE79337608FDA6" box="[307,327,514,539]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FEBA793076ADFDA6" box="[343,482,513,539]" italics="true" pageId="15" pageNumber="290">microlophus</emphasis>
|
|
</taxonomicName>
|
|
appears to prefer open freshwater habitats mostly surrounded by high trees. We observed this species mainly on stony or rocky ground. In Bantimurung specimens were seen in a river with large boulders and low vegetation on the banks. Remarkably, this site was near the entrance of the Bantimurung National Park that is highly frequented by many visitors. Some specimens even occurred near a busy building site.
|
|
</paragraph>
|
|
<paragraph id="7745B933FF907B55FF2A79A073B3FC3E" blockId="15.[151,1437,153,1007]" pageId="15" pageNumber="290">
|
|
Near Bengo-Bengo six specimens (one male and five females) were observed on a tree. It appeared as if this assemblage had a harem structure with the single male being the dominant individual. According to locals and own observations, individuals living close to human settlements are often seen in trees during the dry season to escape the hunting dogs. In contrast, during the rainy season a more terrestrial behaviour is observed. Individuals or groups of sailfin lizards of different ages are mainly seen on riverbanks and flee into the river when disturbed. We observed several specimens of all age groups living together within a relatively small range especially in the dry season. The semiaquatic habitat was shared with
|
|
<taxonomicName id="B0FAC2B0FF907B55FDDF785B75BAFC3E" baseAuthorityName="Peters" baseAuthorityYear="1872" box="[562,757,874,899]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="togianus">
|
|
<emphasis id="458E6521FF907B55FDDF785B75BAFC3E" box="[562,757,874,899]" italics="true" pageId="15" pageNumber="290">Varanus togianus</emphasis>
|
|
</taxonomicName>
|
|
, a potential predator of juvenile
|
|
<taxonomicName id="B0FAC2B0FF907B55FB89785873BAFC3E" authorityName="Kaup" authorityYear="1828" box="[1124,1269,873,899]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="genus">
|
|
<emphasis id="458E6521FF907B55FB89785873BAFC3E" box="[1124,1269,873,899]" italics="true" pageId="15" pageNumber="290">Hydrosaurus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3FE0EAB8FF907B55FF2A78BD76C2FC52" pageId="15" pageNumber="290" type="distribution">
|
|
<paragraph id="7745B933FF907B55FF2A78BD76C2FC52" blockId="15.[151,1437,153,1007]" pageId="15" pageNumber="290">
|
|
<emphasis id="458E6521FF907B55FF2A78BD762CFC1A" bold="true" box="[199,355,908,935]" pageId="15" pageNumber="290">Distribution:</emphasis>
|
|
<taxonomicName id="B0FAC2B0FF907B55FE8178BF7554FC1A" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[364,539,909,935]" class="Reptilia" family="Agamidae" genus="Hydrosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="290" phylum="Chordata" rank="species" species="microlophus">
|
|
<emphasis id="458E6521FF907B55FE8178BF76CFFC1A" box="[364,384,910,935]" italics="true" pageId="15" pageNumber="290">H</emphasis>
|
|
.
|
|
<emphasis id="458E6521FF907B55FE7D78BC7554FC1A" box="[400,539,909,935]" italics="true" pageId="15" pageNumber="290">microlophus</emphasis>
|
|
</taxonomicName>
|
|
is endemic to
|
|
<collectingRegion id="B53E77D1FF907B55FD2A78BC747FFC1A" box="[711,816,909,935]" country="Indonesia" name="Sulawesi Utara" pageId="15" pageNumber="290">Sulawesi</collectingRegion>
|
|
. It appears to be restricted to Southwest
|
|
<collectingRegion id="B53E77D1FF907B55FAE878BC7222FC1A" box="[1285,1389,909,935]" country="Indonesia" name="Sulawesi Utara" pageId="15" pageNumber="290">Sulawesi</collectingRegion>
|
|
, the bordering southern part of
|
|
<collectingRegion id="B53E77D1FF907B55FE2B788075CAFC76" box="[454,645,945,971]" country="Indonesia" name="Sulawesi Tengah" pageId="15" pageNumber="290">Central Sulawesi</collectingRegion>
|
|
and possibly also inhabits parts of
|
|
<collectingRegion id="B53E77D1FF907B55FBFE788073F8FC76" box="[1043,1207,945,971]" country="Indonesia" name="Sulawesi Barat" pageId="15" pageNumber="290">West Sulawesi</collectingRegion>
|
|
(
|
|
<figureCitation id="EFC1A5B6FF907B55FB2A78807266FC76" box="[1223,1321,945,971]" captionStart="FIGURE 8" captionStartId="18.[151,250,1846,1871]" captionTargetBox="[151,1436,524,1826]" captionTargetId="figure@18.[151,1436,524,1829]" captionTargetPageId="18" captionText="FIGURE 8. Map of Sulawesi showing the distribution of H. microlophus (red) and H. celebensis (grey). Type localities are indicated by stars.. 0 Makassar; 1 Bantimurung; 2 Pattunuang; 3 Bengo-Bengo & Rompegading; 4 Camba; 5 Ujung Lamuru; 6 Belopa; 7 Palopo-Pompengan & Palopo-Siguntu; 8 Toraja; 9 Malele; 10 Poso River, Bay of Tomini; 11 Desa Jelaja; 12 Wasponda; 13 Desa Limbatu; 14 Morowali Nature Reserve; W1 Tempe („at the Minralang river near Tempe“ after Weber1890: 167); W2 Pampanua („at Pampanua on the river Tjinrana“ after Weber 1890: 167)" figureDoi="http://doi.org/10.5281/zenodo.3694822" httpUri="https://zenodo.org/record/3694822/files/figure.png" pageId="15" pageNumber="290">Figure 8</figureCitation>
|
|
). See also the Discussion below.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |