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<document id="FD1407D95696D1E29FFD3B63A42FCBF1" ID-CLB-Dataset="8462" ID-DOI="10.5281/zenodo.4502008" ID-GBIF-Dataset="47a5d23f-46d7-430d-87d9-1179994d7d4f" ID-ISSN="2345-7600" ID-Zenodo-Dep="4502008" ID-ZooBank="0ADEAACE-24DD-4535-9378-9DF50A432A8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612458767264" checkinUser="carolina" docAuthor="Cai, Yixiong &amp; Vidthayanon, Chavalit" docDate="2016" docId="FF4187D7FFA22669FC5937FBFEA8F9AD" docLanguage="en" docName="Raffles.64.117-122.pdf" docOrigin="Raffles Bulletin of Zoology 64" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Macrobrachium spelaeus Cai &amp; Vidthayanon 2016, new species" docType="treatment" docUuid="93B28A24-83DB-468F-94E5-5B91746EB857" docUuidSource="ZooBank" docVersion="7" lastPageNumber="121" masterDocId="0378FFAFFFA2266DFFF5321EFFA8FFF8" masterDocTitle="Macrobrachium spelaeus, a new species of stygobitic freshwater prawn from Thailand (Decapoda: Palaemonidae)" masterLastPageNumber="122" masterPageNumber="117" pageNumber="117" updateTime="1698926722764" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6E5F262D78403C8592A7AEC584E45BF2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="685860CCD6B69FDA3A2634F482D1F470">
<mods:title id="7B81073570EC00F10B4ADBC85C12EEA2">Macrobrachium spelaeus, a new species of stygobitic freshwater prawn from Thailand (Decapoda: Palaemonidae)</mods:title>
</mods:titleInfo>
<mods:name id="E6E89A5A0B30D613934B5CC3D3BC206B" type="personal">
<mods:role id="046F220F85666AC4467F999A8B29AC0B">
<mods:roleTerm id="E56BA2DB15A8E2AF83780793919C8B3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C963D89247EBBA85E43A4F9DEFC6509">Cai, Yixiong</mods:namePart>
</mods:name>
<mods:name id="09A5A7E150F7DF0CD2995E700B9A31AF" type="personal">
<mods:role id="BA0D5C5AD4484B042F441BE81C386C67">
<mods:roleTerm id="19BE14C015C21C96E815076526D916DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9DDAE219C1424B857F1801FAEA049EA8">Vidthayanon, Chavalit</mods:namePart>
</mods:name>
<mods:typeOfResource id="8AC27A381B6CB951E0EBA5932D7FC91D">text</mods:typeOfResource>
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<mods:title id="FDBB07684089134D0F2C9F7A21AA2597">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="9F99C2DF5481411936076BE08000EBC9">
<mods:date id="22C3CC2969CE9C096A8784C93D06AEFB">2016</mods:date>
<mods:detail id="A3447B7A496040E6DD25680D22BED159" type="pubDate">
<mods:number id="B17DE8A4E54D6ACC1B0B65C72AC2B4D2">2016-06-20</mods:number>
</mods:detail>
<mods:detail id="46F396CD5A3138DDC69D757D2BCA482D" type="volume">
<mods:number id="9033B3ADCCC2763CE097C1212BA9D9C5">64</mods:number>
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<mods:start id="BC7F602E7D934E276C519571676B4AA7">117</mods:start>
<mods:end id="8F93732D511BD0E8E9705FE3DDDA64DF">122</mods:end>
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<mods:classification id="00764A9FCCC453ED168B99C07CD30AD8">journal article</mods:classification>
<mods:identifier id="C6D3E2EA11390F44E6507CC0DA8FFBD0" type="CLB-Dataset">8462</mods:identifier>
<mods:identifier id="948DC71758BB1E2C1898DCEDB9055C2A" type="DOI">10.5281/zenodo.4502008</mods:identifier>
<mods:identifier id="EBBECB28CC1D5FF0198409BC0EB9103B" type="GBIF-Dataset">47a5d23f-46d7-430d-87d9-1179994d7d4f</mods:identifier>
<mods:identifier id="4334E10815D760AF2FA04CF8E2D62B49" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E5216919CCED27B5175336A39A191BBE" type="Zenodo-Dep">4502008</mods:identifier>
<mods:identifier id="E321BDA172E0F1A5CD5DE733C733A9AA" type="ZooBank">0ADEAACE-24DD-4535-9378-9DF50A432A8C</mods:identifier>
</mods:mods>
<treatment id="FF4187D7FFA22669FC5937FBFEA8F9AD" ID-DOI="http://doi.org/10.5281/zenodo.4502012" ID-GBIF-Taxon="177210204" ID-Zenodo-Dep="4502012" LSID="urn:lsid:zoobank.org:act:93B28A24-83DB-468F-94E5-5B91746EB857" httpUri="http://treatment.plazi.org/id/FF4187D7FFA22669FC5937FBFEA8F9AD" lastPageId="4" lastPageNumber="121" pageId="0" pageNumber="117">
<subSubSection id="3FF2654AFFA2266DFC5937FBFAF2FA06" box="[940,1370,1509,1534]" pageId="0" pageNumber="117" type="nomenclature">
<paragraph id="775736C1FFA2266DFC5937FBFAF2FA06" blockId="0.[940,1370,1509,1566]" box="[940,1370,1509,1534]" pageId="0" pageNumber="117">
<heading id="2C1F81ADFFA2266DFC5937FBFAF2FA06" bold="true" box="[940,1370,1509,1534]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="117" reason="2">
<emphasis id="459CEAD3FFA2266DFC5937FBFAF2FA06" bold="true" box="[940,1370,1509,1534]" pageId="0" pageNumber="117">
<taxonomicName id="B0E84D42FFA2266DFC5937FBFB6DFA05" ID-CoL="5d6a9faf-b39f-4c12-9bf9-4f3346df05d8" authority="Cai &amp; Vidthayanon, 2016" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[940,1221,1509,1533]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
<emphasis id="459CEAD3FFA2266DFC5937FBFB6DFA05" bold="true" box="[940,1221,1509,1533]" italics="true" pageId="0" pageNumber="117">Macrobrachium spelaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5EAF57A8FFA2266DFB2137F8FAF2FA06" box="[1236,1370,1510,1534]" pageId="0" pageNumber="117" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3FF2654AFFA2266DFBB23418FB16F9E6" box="[1095,1214,1542,1566]" pageId="0" pageNumber="117" type="description">
<paragraph id="775736C1FFA2266DFBB23418FB16F9E6" blockId="0.[940,1370,1509,1566]" box="[1095,1214,1542,1566]" pageId="0" pageNumber="117">
(
<figureCitation id="EFD32A44FFA2266DFBBA3418FB1DF9E6" box="[1103,1205,1542,1566]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[113,148,2006,2028]" captionStartId-1="3.[113,148,2064,2086]" captionStartId-2="4.[136,171,1072,1094]" captionTargetBox-0="[113,1451,200,1991]" captionTargetBox-1="[114,1453,207,2051]" captionTargetBox-2="[136,1474,189,1056]" captionTargetId-0="figure-10@1.[94,1460,176,1994]" captionTargetId-1="figure-10@3.[100,1464,187,2057]" captionTargetId-2="figure-22@4.[136,1474,189,1056]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." captionText-1="Fig. 2. Macrobrachium spelaeus, new species. A, first pereiopods; B, C, second pereiopods; D, E, third pereiopods, Scale bars = 2 mm [A, C]; 4 mm [B]; 1.5 mm [D]; 0.5 mm [E] (holotype: male, cl 13.7 mm, NIFI SH 00176)." captionText-2="Fig. 3. Colour photograph of Macrobrachium spelaeus, new species (paratype, male, cl 11.7mm, OUMNH.ZC.2016-01-022)" figureDoi-0="http://doi.org/10.5281/zenodo.4502010" figureDoi-1="http://doi.org/10.5281/zenodo.4502014" figureDoi-2="http://doi.org/10.5281/zenodo.4502016" httpUri-0="https://zenodo.org/record/4502010/files/figure.png" httpUri-1="https://zenodo.org/record/4502014/files/figure.png" httpUri-2="https://zenodo.org/record/4502016/files/figure.png" pageId="0" pageNumber="117">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3FF2654AFFA2266DFCB63458FC00F8C6" pageId="0" pageNumber="117" type="materials_examined">
<paragraph id="775736C1FFA2266DFCB63458FC00F8C6" blockId="0.[834,1475,1606,1854]" pageId="0" pageNumber="117">
<emphasis id="459CEAD3FFA2266DFCB63458FB89F9A6" bold="true" box="[835,1057,1606,1630]" pageId="0" pageNumber="117">Material examined.</emphasis>
<materialsCitation id="C7803C9CFFA2266DFBD23458FA6BF926" ID-GBIF-Occurrence="3029797302" collectingDate="2002-08-28" collectionCode="NIFI" collectorName="C. Vidthayanon." country="Thailand" county="Pitsanulok Province" elevation="100" latitude="16.8379" location="Tham Phra Wangdaeng" longLatPrecision="53" longitude="100.877" municipality="Thung Salaeng Luang National Park" pageId="0" pageNumber="117" specimenCode="SH 00176" specimenCount="100" typeStatus="holotype">
<typeStatus id="A8538863FFA2266DFBD23458FB26F9A6" box="[1063,1166,1606,1630]" pageId="0" pageNumber="117" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="B0109B24FFA2266DFB043458FAFAF9A5" box="[1265,1362,1606,1630]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="0" pageNumber="117" unit="mm" value="13.7">13.7 mm</quantity>
, (
<collectionCode id="11F9AE04FFA2266DFA913458FA32F9A6" box="[1380,1434,1606,1630]" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="0" pageNumber="117">NIFI</collectionCode>
<specimenCode id="274E9EBAFFA2266DFA6A3458FC22F986" collectionCode="NIFI" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="0" pageNumber="117">SH 00176</specimenCode>
),
<location id="7237601AFFA2266DFC6F3478FB3EF986" LSID="urn:lsid:plazi:treatment:FF4187D7FFA22669FC5937FBFEA8F9AD:7237601AFFA2266DFC6F3478FB3EF986" box="[922,1174,1638,1662]" country="Thailand" county="Pitsanulok Province" latitude="16.8379" longLatPrecision="53" longitude="100.877" municipality="Thung Salaeng Luang National Park" name="Tham Phra Wangdaeng" pageId="0" pageNumber="117">Tham Phra Wangdaeng</location>
,
<location id="7237601AFFA2266DFB6A3478FA9FF986" LSID="urn:lsid:plazi:treatment:FF4187D7FFA22669FC5937FBFEA8F9AD:7237601AFFA2266DFB6A3478FA9FF986" box="[1183,1335,1638,1662]" country="Thailand" county="Pitsanulok Province" latitude="16.8379" longLatPrecision="53" longitude="100.877" municipality="Thung Salaeng Luang National Park" name="Baan Chompu" pageId="0" pageNumber="117">Baan Chompu</location>
southwest of
<collectingMunicipality id="9733ACBBFFA2266DFCB73498FB7DF966" box="[834,1237,1670,1694]" pageId="0" pageNumber="117">Thung Salaeng Luang National Park</collectingMunicipality>
,
<collectingCounty id="9E364E4DFFA2266DFB143498FA15F966" box="[1249,1469,1670,1694]" pageId="0" pageNumber="117">Pitsanulok Province</collectingCounty>
, lower north
<collectingCountry id="0FFF7651FFA2266DFC2334B8FB96F946" box="[982,1086,1702,1726]" name="Thailand" pageId="0" pageNumber="117">Thailand</collectingCountry>
,
<specimenCount id="61EEFD48FFA2266DFBBB34B8FB08F945" box="[1102,1184,1702,1726]" pageId="0" pageNumber="117" type="generic">
<quantity id="B0109B24FFA2266DFBBB34B8FB34F945" box="[1102,1180,1702,1726]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="117" unit="m" value="100.0">
<elevation id="FCC5D1F2FFA2266DFBBB34B8FB34F945" box="[1102,1180,1702,1726]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="117" unit="m" value="100.0">100 m</elevation>
</quantity>
.
</specimenCount>
from the cave entrance:
<geoCoordinate id="12DC5006FFA2266DFCB634D8FC1DF926" box="[835,949,1734,1758]" degrees="16.8379" direction="north" orientation="latitude" pageId="0" pageNumber="117" precision="5" value="16.8379">16.8379°N</geoCoordinate>
<geoCoordinate id="12DC5006FFA2266DFC4E34D8FB85F926" box="[955,1069,1734,1758]" degrees="100.877" direction="east" orientation="longitude" pageId="0" pageNumber="117" precision="55" value="100.877">100.877°E</geoCoordinate>
,
<date id="03561001FFA2266DFBC034D8FB74F926" box="[1077,1244,1734,1758]" pageId="0" pageNumber="117" value="2002-08-28">
<collectingDate id="1312E9E9FFA2266DFBC034D8FB74F926" box="[1077,1244,1734,1758]" pageId="0" pageNumber="117" value="2002-08-28">28 August 2002</collectingDate>
</date>
, coll.
<collectorName id="DA1D5317FFA2266DFAE234D8FA6BF926" box="[1303,1475,1734,1758]" pageId="0" pageNumber="117">C. Vidthayanon.</collectorName>
</materialsCitation>
<materialsCitation id="C7803C9CFFA2266DFCB734F8FA15F906" ID-GBIF-Occurrence="3029797301" box="[834,1469,1766,1790]" collectionCode="OUMNH, ZC" country="Thailand" pageId="0" pageNumber="117" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="A8538863FFA2266DFCB734F8FC06F906" box="[834,942,1766,1790]" pageId="0" pageNumber="117" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="61EEFD48FFA2266DFC4D34F8FBABF906" box="[952,1027,1766,1790]" pageId="0" pageNumber="117" type="male">1 male</specimenCount>
, cl
<quantity id="B0109B24FFA2266DFBDF34F8FB2AF906" box="[1066,1154,1766,1790]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="0" pageNumber="117" unit="mm" value="11.7">11.7mm</quantity>
(
<collectionCode id="11F9AE04FFA2266DFB6734F8FB57F906" box="[1170,1279,1766,1790]" pageId="0" pageNumber="117">OUMNH</collectionCode>
.
<collectionCode id="11F9AE04FFA2266DFAF734F8FA8EF906" box="[1282,1318,1766,1790]" country="Austria" name="Zoological Collection, University of Vienna" pageId="0" pageNumber="117" type="University or college">ZC</collectionCode>
.2016-01-022)
</materialsCitation>
,
<materialsCitation id="C7803C9CFFA2266DFCB13518FC0CF8C6" ID-GBIF-Occurrence="3029797304" collectingDate="2002-08-28" collectionCode="ZRC" collectorName="C. Vidthayanon." country="Thailand" county="Pitsanulok Province" elevation="100" latitude="16.8379" location="Tham Phra Wangdaeng" longLatPrecision="53" longitude="100.877" municipality="Thung Salaeng Luang National Park" pageId="0" pageNumber="117" specimenCode="ZRC 2016.0049" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="61EEFD48FFA2266DFCB13518FC01F8E6" box="[836,937,1798,1822]" pageId="0" pageNumber="117" type="female">1 female</specimenCount>
, cl
<quantity id="B0109B24FFA2266DFC203518FB87F8E6" box="[981,1071,1798,1822]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="0" pageNumber="117" unit="mm" value="11.9">11.9mm</quantity>
(
<specimenCode id="274E9EBAFFA2266DFBB53518FB51F8E6" box="[1088,1273,1798,1822]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="0" pageNumber="117">ZRC 2016.0049</specimenCode>
), same locality as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<footnote id="14F32ACFFFA2266DFF723576FF56F80D" pageId="0" pageNumber="117">
<paragraph id="775736C1FFA2266DFF723576FE96F85F" blockId="0.[135,776,1896,2037]" pageId="0" pageNumber="117">
<superScript id="809D9B89FFA2266DFF723576FF24F88A" attach="right" box="[135,140,1896,1906]" fontSize="4" pageId="0" pageNumber="117">1</superScript>
National Biodiversity Centre, National Parks Board,
<collectingCountry id="0FFF7651FFA2266DFDC23577FD2FF882" box="[567,647,1897,1914]" name="Singapore" pageId="0" pageNumber="117">Singapore</collectingCountry>
, 1 Cluny Road,
<collectingCountry id="0FFF7651FFA2266DFF723561FF7EF868" box="[135,214,1919,1936]" name="Singapore" pageId="0" pageNumber="117">Singapore</collectingCountry>
259569,
<collectingCountry id="0FFF7651FFA2266DFED03561FE75F868" box="[293,477,1919,1936]" name="Singapore" pageId="0" pageNumber="117">Republic of Singapore</collectingCountry>
; Email: cai_yixiong@nparks.gov.sg (*corresponding author)
</paragraph>
<subSubSection id="3FF2654AFFA2266FFCB63578FA84FD0D" lastPageId="2" lastPageNumber="119" pageId="0" pageNumber="117" type="description">
<paragraph id="775736C1FFA2266FFCB63578FD4BFEED" blockId="0.[835,1476,1894,2110]" lastBlockId="2.[135,774,189,277]" lastPageId="2" lastPageNumber="119" pageId="0" pageNumber="117">
<emphasis id="459CEAD3FFA2266DFCB63578FC66F886" bold="true" box="[835,974,1894,1918]" pageId="0" pageNumber="117">Description.</emphasis>
Rostrum (
<figureCitation id="EFD32A44FFA2266DFBB43578FB1EF886" box="[1089,1206,1894,1918]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="0" pageNumber="117">Fig. 1A, B</figureCitation>
) straight, reaching distal end of scaphocerite, rostral formula: 4-5+6-7/2-3. Teeth more widely spaced on anterior than postorbital region. Antennular peduncle about 0.4 times as long as carapace. Carapace smooth, inferior orbital margin moderately produced. Antennal spine sharp (
<figureCitation id="EFD32A44FFA2266DFB5E3A18FAA8F7E6" box="[1195,1280,2054,2078]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="0" pageNumber="117">Fig. 1D</figureCitation>
), situated slightly lower than inferior orbital angle, reaching slightly beyond carapace margin. Hepatic spine smaller, situated behind and substantially below antennal spine. Branchiostegal suture running from base of hepatic spine to carapace margin.
</paragraph>
<paragraph id="775736C1FFA2266DFF7235A8FF56F80D" blockId="0.[135,776,1896,2037]" pageId="0" pageNumber="117">
<superScript id="809D9B89FFA2266DFF7235A8FF24F838" attach="right" box="[135,140,1974,1984]" fontSize="4" pageId="0" pageNumber="117">2</superScript>
Environment Division, Mekong River Commission Secretariat, Fa Ngum Road, Ban Sithan Noua,
<collectingRegion id="B52CF823FFA2266DFF0435D3FE92F826" box="[241,314,1997,2014]" country="Laos" name="Viangchan" pageId="0" pageNumber="117">Vientiane</collectingRegion>
01000, Lao People Democratic Republic; Email: chavalit@ mrcmekong.org
</paragraph>
<pageTitle id="3777EEA6FFA2266DFF723A08FDCBF7B9" pageId="0" pageNumber="117">
<paragraph id="775736C1FFA2266DFF723A08FE17F7D1" blockId="0.[135,611,2070,2114]" box="[135,447,2070,2090]" pageId="0" pageNumber="117">
© National University of
<collectingCountry id="0FFF7651FFA2266DFE9D3A08FE17F7D1" box="[360,447,2070,2089]" name="Singapore" pageId="0" pageNumber="117">Singapore</collectingCountry>
</paragraph>
<paragraph id="775736C1FFA2266DFF723A31FDCBF7B9" blockId="0.[135,611,2070,2114]" box="[135,611,2094,2114]" pageId="0" pageNumber="117">
ISSN
<docID-ISSN id="6DC55190FFA2266DFF493A30FEB1F7B9" box="[188,281,2094,2113]" pageId="0" pageNumber="117">
<docIdISSN id="EFE4E057FFA2266DFF493A30FEB1F7B9" box="[188,281,2094,2113]" pageId="0" pageNumber="117">2345-7600</docIdISSN>
</docID-ISSN>
(electronic) | ISSN 0217-2445 (print)
</paragraph>
</pageTitle>
<caption id="23976649FFA3266CFF8435C8FE31F7BB" ID-DOI="http://doi.org/10.5281/zenodo.4502010" ID-Zenodo-Dep="4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="1" pageNumber="118" startId="1.[113,148,2006,2028]" targetBox="[113,1451,200,1991]" targetPageId="1">
<paragraph id="775736C1FFA3266CFF8435C8FE31F7BB" blockId="1.[113,1452,2006,2115]" pageId="1" pageNumber="118">
Fig. 1.
<taxonomicName id="B0E84D42FFA3266CFF4F35C8FE1AF813" authority="Cai &amp; Vidthayanon, 2016" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[186,434,2006,2027]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
<emphasis id="459CEAD3FFA3266CFF4F35C8FE1AF813" box="[186,434,2006,2027]" italics="true" pageId="1" pageNumber="118">Macrobrachium spelaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5EAF57A8FFA3266CFE3535C9FD91F814" box="[448,569,2006,2028]" pageId="1" pageNumber="118" rank="species">new species</taxonomicNameLabel>
. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1
<superScript id="809D9B89FFA3266CFDFE35EDFD8AF7F0" attach="none" box="[523,546,2035,2056]" fontSize="5" pageId="1" pageNumber="118">st–</superScript>
3
<superScript id="809D9B89FFA3266CFDD935ECFD90F806" attach="left" box="[556,568,2034,2046]" fontSize="5" pageId="1" pageNumber="118">rd</superScript>
abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176).
</paragraph>
</caption>
<paragraph id="775736C1FFA0266FFF723323FDCDFB8D" blockId="2.[133,776,317,1141]" pageId="2" pageNumber="119">
Fourth thoracic sternite (
<figureCitation id="EFD32A44FFA0266FFE6D3323FE41FEAD" box="[408,489,317,341]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1F</figureCitation>
) without median process. Eighth thoracic sternum (
<figureCitation id="EFD32A44FFA0266FFE553343FE44FE8D" box="[416,492,349,373]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig.1G</figureCitation>
) with narrowly separated anterolateral lobes, without median process. Abdomen smooth (
<figureCitation id="EFD32A44FFA0266FFF783383FF79FE4D" box="[141,209,413,437]" captionStart="Fig" captionStartId="4.[136,171,1072,1094]" captionTargetBox="[136,1474,189,1056]" captionTargetId="figure-22@4.[136,1474,189,1056]" captionTargetPageId="4" captionText="Fig. 3. Colour photograph of Macrobrachium spelaeus, new species (paratype, male, cl 11.7mm, OUMNH.ZC.2016-01-022)" figureDoi="http://doi.org/10.5281/zenodo.4502016" httpUri="https://zenodo.org/record/4502016/files/figure.png" pageId="2" pageNumber="119">Fig. 3</figureCitation>
), glabrous, first to third pleurites broadly rounded, fourth and fifth pleurites feebly produced posteriorly, fourth pleurite sub-triangular, fifth pleurite sub-rectangular, sixth abdominal somite 1.5 times as long as fifth, with posteroventral angle feebly produced, subacute. Telson (
<figureCitation id="EFD32A44FFA0266FFD2E3003FF35FDAD" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1J</figureCitation>
) moderate, stout, 3.0 times as long as wide, 1.6 times as long as sixth abdominal segment, with 2 pairs of dorsal spines, ending in a median projection, lateral spines of distal margin slightly larger than dorsal spines, intermediate spines well developed, with long plumose setae. First 3 abdominal sternites (
<figureCitation id="EFD32A44FFA0266FFF1A30C3FEEAFD0D" box="[239,322,733,757]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1H</figureCitation>
) with transverse ridge and a median tooth each, with first one very prominent, much more developed than the other two. Second one triangular in form. Third one much smaller, pointed. Inter-uropodal sclerite armed with well-developed preanal carina (
<figureCitation id="EFD32A44FFA0266FFE183143FD9EFC8D" box="[493,566,861,885]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1I</figureCitation>
).Ocular stalk and cone reduced, 0.12 times of carapace length, facets present, pigment visible, anterior end only reaching middle of basal segment of antennular peduncle. Stylocerite pointed, reaching 0.8 times of second segment. Antenna with stout basicerite and prominent distoventral tooth. Carpocerite reaching to about 0.35 times of scaphocerite length. Scaphocerite (
<figureCitation id="EFD32A44FFA0266FFD2E3603FF00FBAD" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1C</figureCitation>
) slender, 0.6 times as long as carapace, about 3.0 times as long as wide, with straight outer margin.
</paragraph>
<paragraph id="775736C1FFA0266FFF723683FEB2FAAD" blockId="2.[134,776,1181,1365]" pageId="2" pageNumber="119">
Epistome (
<figureCitation id="EFD32A44FFA0266FFF033683FEEAFB4D" box="[246,322,1181,1205]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1E</figureCitation>
) bilobed by a shallow triangular depression. Mouth parts typical of the genus. Third maxilliped with robust endopod, ultimate segment reaching to anterior end of antennal peduncle; ultimate segment slightly shorter than penultimate segment; exopod short, with numerous plumose setae distally.
</paragraph>
<paragraph id="775736C1FFA0266FFF723763FB7BFECD" blockId="2.[134,776,1405,2101]" lastBlockId="2.[834,1475,189,309]" pageId="2" pageNumber="119">
First pereiopods (
<figureCitation id="EFD32A44FFA0266FFEB93763FE0CFA6D" box="[332,420,1405,1429]" captionStart="Fig" captionStartId="3.[113,148,2064,2086]" captionTargetBox="[114,1453,207,2051]" captionTargetId="figure-10@3.[100,1464,187,2057]" captionTargetPageId="3" captionText="Fig. 2. Macrobrachium spelaeus, new species. A, first pereiopods; B, C, second pereiopods; D, E, third pereiopods, Scale bars = 2 mm [A, C]; 4 mm [B]; 1.5 mm [D]; 0.5 mm [E] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502014" httpUri="https://zenodo.org/record/4502014/files/figure.png" pageId="2" pageNumber="119">Fig. 2A</figureCitation>
) very slender, reaching beyond scaphocerite by entire chela and 0.2 carpus length, equal in length, similar in form, carpus 1.8 times as long as chela, chela with fingers slightly shorter than palm. Second pereiopods (
<figureCitation id="EFD32A44FFA0266FFF7A37E3FF55F9ED" box="[143,253,1533,1557]" captionStart="Fig" captionStartId="3.[113,148,2064,2086]" captionTargetBox="[114,1453,207,2051]" captionTargetId="figure-10@3.[100,1464,187,2057]" captionTargetPageId="3" captionText="Fig. 2. Macrobrachium spelaeus, new species. A, first pereiopods; B, C, second pereiopods; D, E, third pereiopods, Scale bars = 2 mm [A, C]; 4 mm [B]; 1.5 mm [D]; 0.5 mm [E] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502014" httpUri="https://zenodo.org/record/4502014/files/figure.png" pageId="2" pageNumber="119">Fig. 2B, C</figureCitation>
) not sexually dimorphic; both cylindrical, similar in form, unequal in length. Both covered with tufts of velvety setae on fingers and distal half of palms. Major one with smooth or with indistinct spinules, subequal to body length, reaching beyond distal end of scaphocerite by both entire carpus and chela; merus as long as palm, both segments distinctly longer than palm but shorter than fingers; capus 0.65 times as long as palm; palm slightly inflated, fingers with no gap when closed, with a row of about 15 small teeth on each side of cutting edge. Third pereiopods (
<figureCitation id="EFD32A44FFA0266FFD7E3503FD55F8CD" box="[651,765,1821,1845]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1D, E</figureCitation>
) slender, reaching beyond scaphocerite by entire dactylus, propodus 10 times as long as broad, 3.4 times as long as dactylus; dactylus 4.5 times as long as wide, terminating in an unguis. Fourth pereiopods slender, slightly longer than third pereiopods, similar in form. Fifth pereiopods most slender, longest, reaching beyond scaphocerite by entire dactylus. Endopod of male first pleopod about 0.45 times as long as exopod, weakly broadened distally, slightly curved mesially. Appendix masculina of male second pleopod longer than appendix interna, with spiniform setae on dorsal surface. Appendix interna of male second pleopod slender, reaching to 0.6 length of appendix masculina.
</paragraph>
<paragraph id="775736C1FFA0266FFCB63343FBE4FE6D" blockId="2.[834,1475,349,405]" pageId="2" pageNumber="119">
Uropodal diaeresis (
<figureCitation id="EFD32A44FFA0266FFBE23343FBCFFE8D" box="[1047,1127,349,373]" captionStart="Fig" captionStartId="1.[113,148,2006,2028]" captionTargetBox="[113,1451,200,1991]" captionTargetId="figure-10@1.[94,1460,176,1994]" captionTargetPageId="1" captionText="Fig. 1. Macrobrachium spelaeus, new species. A, B, cephalothorax; C, scaphocerite; D, antennal spine; E, epistome; F, fourth thoracic sternite; G, eighth thoracic sternum; H, 1st–3rd abdominal sterna with transverse ridge and median tooth; I, preanal carina; J, telson; K, uropodal diaeresis. Scale bars = 4 mm [A–C]; 1.5 mm [D]; 0.8 mm [E, H, I]; 1.2 mm [G, J]; 1.0 mm [F]; 0.5 mm [K] (holotype: male, cl 13.7 mm, NIFI SH 00176)." figureDoi="http://doi.org/10.5281/zenodo.4502010" httpUri="https://zenodo.org/record/4502010/files/figure.png" pageId="2" pageNumber="119">Fig. 1K</figureCitation>
) with a movable spine, distinctly shorter than outer angle.
</paragraph>
<paragraph id="775736C1FFA0266FFCB633A3FA03FDCC" blockId="2.[835,1474,445,565]" pageId="2" pageNumber="119">
<emphasis id="459CEAD3FFA0266FFCB633A3FBBCFE2D" bold="true" box="[835,1044,445,469]" pageId="2" pageNumber="119">Colouration (live).</emphasis>
Yellowish pale cephalothorax with pale yellow and brown visceral mass, rostrum translucent, eyes black, abdomen opaque with brownish hue dorsally. Hairs on chelae pale brown, antennas and appendages opaque.
</paragraph>
<paragraph id="775736C1FFA0266FFCB63043FA84FD0D" blockId="2.[835,1476,605,757]" pageId="2" pageNumber="119">
<emphasis id="459CEAD3FFA0266FFCB63043FC0EFD8D" bold="true" box="[835,934,605,629]" pageId="2" pageNumber="119">Habitat.</emphasis>
<taxonomicName id="B0E84D42FFA0266FFC443043FB79FD8D" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[945,1233,605,629]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="459CEAD3FFA0266FFC443043FB79FD8D" box="[945,1233,605,629]" italics="true" pageId="2" pageNumber="119">Macrobrachium spelaeus</emphasis>
</taxonomicName>
is known only from shallow (
<quantity id="B0109B24FFA0266FFC5C3063FBADFD6C" box="[937,1029,637,661]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="2" pageNumber="119" unit="m" value="0.65" valueMax="1.0" valueMin="0.3">0.3–1 m</quantity>
depth) subterranean streams in the
<typeStatus id="A8538863FFA0266FFA613063FA6AFD6D" box="[1428,1474,637,661]" pageId="2" pageNumber="119">type</typeStatus>
locality, over
<quantity id="B0109B24FFA0266FFC2F3083FBB7FD4C" box="[986,1055,669,693]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="119" unit="m" value="100.0">100 m</quantity>
from the cave entrance, at Tham Phra Wang Daeng of the Tham Phra Karst in Klong Chompu area of Thung Salaeng Luang National Park.
</paragraph>
</subSubSection>
</footnote>
<subSubSection id="3FF2654AFFA0266FFCB63103FB6BFCAD" pageId="2" pageNumber="119" type="etymology">
<paragraph id="775736C1FFA0266FFCB63103FB6BFCAD" blockId="2.[835,1474,797,853]" pageId="2" pageNumber="119">
<emphasis id="459CEAD3FFA0266FFCB63103FC6DFCCC" bold="true" box="[835,965,797,821]" pageId="2" pageNumber="119">Etymology.</emphasis>
The new species is named
<taxonomicName id="B0E84D42FFA0266FFB023103FAFBFCCD" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[1271,1363,797,821]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="459CEAD3FFA0266FFB023103FAFBFCCD" box="[1271,1363,797,821]" italics="true" pageId="2" pageNumber="119">spelaeus</emphasis>
</taxonomicName>
(L., cavedwelling), after its stygobitic habit.
</paragraph>
</subSubSection>
<subSubSection id="3FF2654AFFA02669FCB63163FEA8F9AD" lastPageId="4" lastPageNumber="121" pageId="2" pageNumber="119" type="discussion">
<paragraph id="775736C1FFA02669FCB63163FD85FACD" blockId="2.[834,1476,893,2101]" lastBlockId="4.[136,775,1149,1333]" lastPageId="4" lastPageNumber="121" pageId="2" pageNumber="119">
<emphasis id="459CEAD3FFA0266FFCB63163FC1AFC6D" bold="true" box="[835,946,893,917]" pageId="2" pageNumber="119">Remarks.</emphasis>
<taxonomicName id="B0E84D42FFA0266FFC4D3163FB13FC6D" authority="Cai &amp; Vidthayanon, 2016" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[952,1211,893,917]" class="Malacostraca" family="Palaemonidae" genus="Mcrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
<emphasis id="459CEAD3FFA0266FFC4D3163FB13FC6D" box="[952,1211,893,917]" italics="true" pageId="2" pageNumber="119">Mcrobrachium spelaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5EAF57A8FFA0266FFB3D3163FAE4FC6D" box="[1224,1356,893,917]" pageId="2" pageNumber="119" rank="species">new species</taxonomicNameLabel>
, represents the first stygobitic
<taxonomicName id="B0E84D42FFA0266FFBF73183FB05FC4D" authorityName="Bate" authorityYear="1868" box="[1026,1197,925,949]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="459CEAD3FFA0266FFBF73183FB05FC4D" box="[1026,1197,925,949]" italics="true" pageId="2" pageNumber="119">Macrobrachium</emphasis>
</taxonomicName>
species found in
<collectingCountry id="0FFF7651FFA0266FFAAB3183FA17FC4D" box="[1374,1471,925,949]" name="Thailand" pageId="2" pageNumber="119">Thailand</collectingCountry>
. With the existence of the tufts of velvety setae on the dactylus of second pereiopods, short carpus in second pereiopods, relatively short rostrum which does not exceed the end of the scaphocerite,
<taxonomicName id="B0E84D42FFA0266FFBFC3603FA8BFBCD" authority="Cai &amp; Vidthayanon, 2016" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[1033,1315,1053,1077]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
<emphasis id="459CEAD3FFA0266FFBFC3603FA8BFBCD" box="[1033,1315,1053,1077]" italics="true" pageId="2" pageNumber="119">Macrobrachium spelaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5EAF57A8FFA0266FFAC03603FA15FBCD" box="[1333,1469,1053,1077]" pageId="2" pageNumber="119" rank="species">new species</taxonomicNameLabel>
, should be referred to the
<taxonomicName id="B0E84D42FFA0266FFBA43623FAC6FBAD" baseAuthorityName="De Man" baseAuthorityYear="1879" box="[1105,1390,1085,1109]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="pilimanus">
<emphasis id="459CEAD3FFA0266FFBA43623FAC6FBAD" box="[1105,1390,1085,1109]" italics="true" pageId="2" pageNumber="119">Macrobrachium pilimanus</emphasis>
</taxonomicName>
species group (cf.
<bibRefCitation id="13794B30FFA0266FFC4E3643FBC4FB8D" author="Cai Y &amp; Naiyanetr P &amp; Ng PKL" box="[955,1132,1117,1141]" pageId="2" pageNumber="119" pagination="581 - 649" refId="ref2654" refString="Cai Y, Naiyanetr P &amp; Ng PKL (2004) The freshwater prawns of the genus Macrobrachium Bate, 1868, of Thailand (Crustacea: Decapoda: Palaemonidae). Journal of Natural History, 38: 581 - 649." type="journal article" year="2004">Cai et al., 2004</bibRefCitation>
). To date, there are 15 species recognised in the
<taxonomicName id="B0E84D42FFA0266FFBF63663FA88FB6D" baseAuthorityName="De Man" baseAuthorityYear="1879" box="[1027,1312,1149,1173]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="pilimanus">
<emphasis id="459CEAD3FFA0266FFBF63663FA88FB6D" box="[1027,1312,1149,1173]" italics="true" pageId="2" pageNumber="119">Macrobrachium pilimanus</emphasis>
</taxonomicName>
group, namely
<taxonomicName id="B0E84D42FFA0266FFCB13680FB26FB4D" authority="(De Man, 1879)" baseAuthorityName="De Man" baseAuthorityYear="1879" box="[836,1166,1181,1205]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="pilimanus">
<emphasis id="459CEAD3FFA0266FFCB13680FC7BFB4D" box="[836,979,1181,1205]" italics="true" pageId="2" pageNumber="119">M. pilimanus</emphasis>
(
<bibRefCitation id="13794B30FFA0266FFC163683FB2FFB4D" author="De Man JG" box="[995,1159,1181,1205]" pageId="2" pageNumber="119" pagination="165 - 184" refId="ref2967" refString="De Man JG (1879) On some species of the genus Palaemon Fabr. with descriptions of two new forms. Notes from the Leyden Museum, 1 (41): 165 - 184." type="journal article" year="1879">De Man, 1879</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFB693680FC2CFB2D" authority="(De Man, 1892)" baseAuthorityName="De Man" baseAuthorityYear="1892" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="leptodactylus">
<emphasis id="459CEAD3FFA0266FFB693680FAF8FB4D" box="[1180,1360,1181,1205]" italics="true" pageId="2" pageNumber="119">M. leptodactylus</emphasis>
(
<bibRefCitation id="13794B30FFA0266FFA943683FCD4FB2D" author="De Man JG" pageId="2" pageNumber="119" pagination="265 - 527" refId="ref3004" refString="De Man JG (1892) Decapoden des IndischenArchipels. In: Weber M, Zoologische Ergebnisseeiner Reise in Niederlandische Ost- Indien, 2: 265 - 527, pls. 15 - 29." type="journal article" year="1892">De Man, 1892</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFC7A36A0FB49FB2D" authority="(Tiwari, 1952)" baseAuthorityName="Tiwari" baseAuthorityYear="1952" box="[911,1249,1213,1237]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="hirsutimanus">
<emphasis id="459CEAD3FFA0266FFC7A36A0FB95FB2D" box="[911,1085,1213,1237]" italics="true" pageId="2" pageNumber="119">M. hirsutimanus</emphasis>
(
<bibRefCitation id="13794B30FFA0266FFBBE36A3FB71FB2D" author="Tiwari KK" box="[1099,1241,1213,1237]" pageId="2" pageNumber="119" pagination="27 - 32" refId="ref3342" refString="Tiwari KK (1952) Diagnosis of new species and subspecies of the genus Palaemon Fabricius (Crustacea: Decapoda). Annals and Magazine of Natural History, (12) 5: 27 - 32." type="journal article" year="1952">Tiwari, 1952</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFB1E36A0FBF3FB0D" authority="Dang &amp; Nguyen, 1972" authorityName="Dang &amp; Nguyen" authorityYear="1972" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="dienbienphuense">
<emphasis id="459CEAD3FFA0266FFB1E36A0FA69FB2D" box="[1259,1473,1213,1237]" italics="true" pageId="2" pageNumber="119">M. dienbienphuense</emphasis>
<bibRefCitation id="13794B30FFA0266FFCB636C3FBF3FB0D" author="Dang NT &amp; Nguyen HY" box="[835,1115,1245,1269]" pageId="2" pageNumber="119" pagination="1 - 6" refId="ref2895" refString="Dang NT &amp; Nguyen HY (1972) Dan lieu ve tom nuoc ngot giong Macrobrachium Bate (Palaemonidae) o cac thuy vuc vung nui mien bac Viet Nam (Faunistic study of freshwater prawn genus Macrobrachium Bate from mountainous region of North Vietnam). Tap san Sinh Vat-Dai Hoc, 10 (1 - 4): 1 - 6. [In Vietnamese with French summary]." type="journal article" year="1972">Dang &amp; Nguyen, 1972</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFB9E36C0FA3BFB0D" authority="Holthuis, 1984" authorityName="Holthuis" authorityYear="1984" box="[1131,1427,1245,1269]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFB9E36C0FB78FB0D" box="[1131,1232,1246,1269]" italics="true" pageId="2" pageNumber="119">M. poeti</emphasis>
<bibRefCitation id="13794B30FFA0266FFB2836C3FA3BFB0D" author="Holthuis LB" box="[1245,1427,1245,1269]" pageId="2" pageNumber="119" pagination="141 - 148" refId="ref3083" refString="Holthuis LB (1984) Freshwater prawns (Crustacea Decapoda: Natantia) from subterranean waters of the Gunung Sewu area, Central Java, Indonesia. Zoologische Mededelingen, Leiden, 58 (9): 141 - 148." type="journal article" year="1984">Holthuis, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFA5136C0FBE9FAED" authority="Dai, 1984" authorityName="Dai" authorityYear="1984" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="eriocheirum">
<emphasis id="459CEAD3FFA0266FFA5136C0FC6FFAED" italics="true" pageId="2" pageNumber="119">M. eriocheirum</emphasis>
<bibRefCitation id="13794B30FFA0266FFC2436E3FBE9FAED" author="Dai AY" box="[977,1089,1277,1301]" pageId="2" pageNumber="119" pagination="244 - 252" refId="ref2852" refString="Dai AY (1984) A preliminary study on the freshwater prawn genus Macrobrachium of China (Decapoda: Caridea). Acta Zootaxanomica Sinica, 9 (3): 244 - 252. [In Chinese with English summary]." type="journal article" year="1984">Dai, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFBA536E0FA6AFAED" authority="Chong &amp; Khoo, 1987" authorityName="Chong &amp; Khoo" authorityYear="1987" box="[1104,1474,1277,1301]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="ahkowi">
<emphasis id="459CEAD3FFA0266FFBA536E0FB6DFAED" box="[1104,1221,1277,1301]" italics="true" pageId="2" pageNumber="119">M. ahkowi</emphasis>
Chong &amp; Khoo, 1987
</taxonomicName>
[=
<taxonomicName id="B0E84D42FFA0266FFCAE3700FB66FACD" authority="Chong &amp; Khoo, 1987" authorityName="Chong &amp; Khoo" authorityYear="1987" box="[859,1230,1309,1333]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="459CEAD3FFA0266FFCAE3700FC73FACD" box="[859,987,1309,1333]" italics="true" pageId="2" pageNumber="119">M. johnsoni</emphasis>
Chong &amp; Khoo, 1987
</taxonomicName>
],
<taxonomicName id="B0E84D42FFA0266FFB2A3700FA16FACD" authority="Chong, 1989" authorityName="Chong" authorityYear="1989" box="[1247,1470,1309,1333]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="gua">
<emphasis id="459CEAD3FFA0266FFB2A3700FA83FACD" box="[1247,1323,1310,1333]" italics="true" pageId="2" pageNumber="119">M. gua</emphasis>
<bibRefCitation id="13794B30FFA0266FFAC73703FA16FACD" author="Chong SSC" box="[1330,1470,1309,1333]" pageId="2" pageNumber="119" pagination="31 - 38" refId="ref2697" refString="Chong SSC (1989) A new species of freshwater prawn, Macrobrachium gua sp. nov. (Decapoda, Caridea, Palaemonidae) from Sabah, East Malaysia, Borneo. Crustaceana, 56 (1): 31 - 38." type="journal article" year="1989">Chong, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFCB63720FBE7FAAD" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[835,1103,1341,1365]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="forcipatum">
<emphasis id="459CEAD3FFA0266FFCB63720FC76FAAD" box="[835,990,1341,1365]" italics="true" pageId="2" pageNumber="119">M. forcipatum</emphasis>
<bibRefCitation id="13794B30FFA0266FFC123723FBE7FAAD" author="Ng PKL" box="[999,1103,1341,1365]" pageId="2" pageNumber="119" pagination="249 - 257" refId="ref3162" refString="Ng PKL (1995) Freshwater decapod crustaceans (Potamidae, Palaemonidae) of Temengor Forest Reserve, Hulu Perak, Malaysia. Malayan Naturalist Journal, 48: 249 - 257." type="journal article" year="1995">Ng, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFBA93720FA16FAAD" authority="Ou &amp; Yeo, 1995" authorityName="Ou &amp; Yeo" authorityYear="1995" box="[1116,1470,1341,1365]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="platycheles">
<emphasis id="459CEAD3FFA0266FFBA93720FB52FAAD" box="[1116,1274,1341,1365]" italics="true" pageId="2" pageNumber="119">M. platycheles</emphasis>
<bibRefCitation id="13794B30FFA0266FFAF63723FA16FAAD" author="Ou ACT &amp; Yeo DCJ" box="[1283,1470,1341,1365]" pageId="2" pageNumber="119" pagination="299 - 308" refId="ref3238" refString="Ou ACT &amp; Yeo DCJ (1995) A new species of freshwater prawn, Macrobrachium platycheles (Decapoda: Caridea: Palaemonidae) from Singapore and Peninsular Malaysia. Raffles Bulletin of Zoology, 43 (2): 299 - 308." type="journal article" year="1995">Ou &amp; Yeo, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFCB63740FB2EFA8D" authority="Cai &amp; Dai, 1999" authorityName="Cai &amp; Dai" authorityYear="1999" box="[835,1158,1373,1397]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="pilosum">
<emphasis id="459CEAD3FFA0266FFCB63740FC16FA8D" box="[835,958,1373,1397]" italics="true" pageId="2" pageNumber="119">M. pilosum</emphasis>
<bibRefCitation id="13794B30FFA0266FFC3C3743FB2EFA8D" author="Cai Y &amp; Dai AY" box="[969,1158,1373,1397]" pageId="2" pageNumber="119" pagination="211 - 241" refId="ref2618" refString="Cai Y &amp; Dai AY (1999) Freshwater shrimps (Crustacea: Decapoda: Caridea) from the Xishuangbanna region of Yunnan Province, southern China. Hydrobiologia, 400: 211 - 241." type="journal article" year="1999">Cai &amp; Dai, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFB613740FCD3FA6D" authority="Cai &amp; Dai, 1999" authorityName="Cai &amp; Dai" authorityYear="1999" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="amplimanus">
<emphasis id="459CEAD3FFA0266FFB613740FA95FA8D" box="[1172,1341,1373,1397]" italics="true" pageId="2" pageNumber="119">M. amplimanus</emphasis>
<bibRefCitation id="13794B30FFA0266FFAB23743FCD3FA6D" author="Cai Y &amp; Dai AY" pageId="2" pageNumber="119" pagination="211 - 241" refId="ref2618" refString="Cai Y &amp; Dai AY (1999) Freshwater shrimps (Crustacea: Decapoda: Caridea) from the Xishuangbanna region of Yunnan Province, southern China. Hydrobiologia, 400: 211 - 241." type="journal article" year="1999">Cai &amp; Dai, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFC733760FB62FA6D" authority="Naiyanetr, 2001" authorityName="Naiyanetr" authorityYear="2001" box="[902,1226,1405,1429]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="sirindhorn">
<emphasis id="459CEAD3FFA0266FFC733760FBB2FA6D" box="[902,1050,1405,1429]" italics="true" pageId="2" pageNumber="119">M. sirindhorn</emphasis>
<bibRefCitation id="13794B30FFA0266FFBEA3763FB62FA6D" author="Naiyanetr P" box="[1055,1226,1405,1429]" pageId="2" pageNumber="119" pagination="609 - 616" refId="ref3124" refString="Naiyanetr P (2001) Macrobrachium sirindhorn n. sp., a new freshwater prawn from northern Thailand (Decapoda, Caridea, Palaemonidae). Crustaceana, 74 (7): 609 - 616." type="journal article" year="2001">Naiyanetr, 2001</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B0E84D42FFA0266FFB2D3760FC4DFA4D" authority="Wowor &amp; Short, 2007" authorityName="Wowor &amp; Short" authorityYear="2007" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="kelianense">
<emphasis id="459CEAD3FFA0266FFB2D3760FAC4FA6D" box="[1240,1388,1405,1429]" italics="true" pageId="2" pageNumber="119">M. kelianense</emphasis>
<bibRefCitation id="13794B30FFA0266FFA873763FC4DFA4D" author="Wowor D &amp; Short JW" pageId="2" pageNumber="119" pagination="77 - 87" refId="ref3484" refString="Wowor D &amp; Short JW (2007) The new freshwater prawns of the genus Macrobrachium Bate, 1868 (Crustacea: Decapoda: Palaemonidae) from the Kelian River, East Kalimantan, Indonesia. Raffles Bulletin of Zoology, 55 (1): 77 - 87." type="journal article" year="2007">Wowor &amp; Short, 2007</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="B0E84D42FFA0266FFBD63780FAF4FA4D" authority="Wowor, 2010" authorityName="Wowor" authorityYear="2010" box="[1059,1372,1437,1461]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="empulipke">
<emphasis id="459CEAD3FFA0266FFBD63780FB10FA4D" box="[1059,1208,1437,1461]" italics="true" pageId="2" pageNumber="119">M. empulipke</emphasis>
<bibRefCitation id="13794B30FFA0266FFB373783FAF4FA4D" author="Wowor D" box="[1218,1372,1437,1461]" pageId="2" pageNumber="119" pagination="715 - 726" refId="ref3429" refString="Wowor D (2010) Macrobrachium empulipke, a new freshwater prawn species (Decapoda, Palaemonidae) from Indonesia. In: Fransen CHJM, De Grave S &amp; Ng PKL (eds.) Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume, Crustaceana Monographs, 14: 715 - 726." type="journal article" year="2010">Wowor, 2010</bibRefCitation>
</taxonomicName>
. With its reduced eyes,
<taxonomicName id="B0E84D42FFA0266FFC2037A0FBFAFA2D" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[981,1106,1469,1493]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="459CEAD3FFA0266FFC2037A0FBFAFA2D" box="[981,1106,1469,1493]" italics="true" pageId="2" pageNumber="119">M. spelaeus</emphasis>
</taxonomicName>
could be easily separated by all the other members of the
<taxonomicName id="B0E84D42FFA0266FFBC237C0FB6DFA0D" baseAuthorityName="De Man" baseAuthorityYear="1879" box="[1079,1221,1501,1525]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="pilimanus">
<emphasis id="459CEAD3FFA0266FFBC237C0FB6DFA0D" box="[1079,1221,1501,1525]" italics="true" pageId="2" pageNumber="119">M. pilimanus</emphasis>
</taxonomicName>
group except
<taxonomicName id="B0E84D42FFA0266FFA9737C0FA13FA0D" authorityName="Holthuis" authorityYear="1984" box="[1378,1467,1502,1525]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFA9737C0FA13FA0D" box="[1378,1467,1502,1525]" italics="true" pageId="2" pageNumber="119">M. poeti</emphasis>
</taxonomicName>
.
<taxonomicName id="B0E84D42FFA0266FFCB637E3FB86F9ED" authorityName="Holthuis" authorityYear="1984" box="[835,1070,1533,1557]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFCB637E3FB86F9ED" box="[835,1070,1533,1557]" italics="true" pageId="2" pageNumber="119">Macrobrachium poeti</emphasis>
</taxonomicName>
was described from several caves in Gunung Sewu,
<collectingRegion id="B52CF823FFA0266FFC063403FB8DF9CC" box="[1011,1061,1565,1588]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="119">Java</collectingRegion>
,
<collectingCountry id="0FFF7651FFA0266FFBC13403FB08F9CD" box="[1076,1184,1565,1589]" name="Indonesia" pageId="2" pageNumber="119">Indonesia</collectingCountry>
.
<taxonomicName id="B0E84D42FFA0266FFB593403FA6AF9CD" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[1196,1474,1565,1589]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="459CEAD3FFA0266FFB593403FA6AF9CD" box="[1196,1474,1565,1589]" italics="true" pageId="2" pageNumber="119">Macrobrachium spelaeus</emphasis>
</taxonomicName>
could be distinguished from
<taxonomicName id="B0E84D42FFA0266FFB773420FB76F9AD" authorityName="Holthuis" authorityYear="1984" box="[1154,1246,1598,1621]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFB773420FB76F9AD" box="[1154,1246,1598,1621]" italics="true" pageId="2" pageNumber="119">M. poeti</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="13794B30FFA0266FFAE23423FA13F9AD" author="Holthuis LB" box="[1303,1467,1597,1621]" pageId="2" pageNumber="119" pagination="141 - 148" refId="ref3083" refString="Holthuis LB (1984) Freshwater prawns (Crustacea Decapoda: Natantia) from subterranean waters of the Gunung Sewu area, Central Java, Indonesia. Zoologische Mededelingen, Leiden, 58 (9): 141 - 148." type="journal article" year="1984">Holthuis, 1984</bibRefCitation>
) by having more teeth on the lower margin of the rostrum (2 or 3 vs.
<quantity id="B0109B24FFA0266FFC353463FC43F96D" box="[960,1003,1661,1685]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="119" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="B0E84D42FFA0266FFC013460FBE7F96D" authorityName="Holthuis" authorityYear="1984" box="[1012,1103,1662,1685]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFC013460FBE7F96D" box="[1012,1103,1662,1685]" italics="true" pageId="2" pageNumber="119">M. poeti</emphasis>
</taxonomicName>
); the shorter merus of the second pereiopods (as long as palm vs. distinctly longer than palm in
<taxonomicName id="B0E84D42FFA0266FFCB734A0FC32F92D" authorityName="Holthuis" authorityYear="1984" box="[834,922,1726,1749]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFCB734A0FC32F92D" box="[834,922,1726,1749]" italics="true" pageId="2" pageNumber="119">M. poeti</emphasis>
</taxonomicName>
); the larger number of teeth on the cutting edges of the fingers (15 vs. several in
<taxonomicName id="B0E84D42FFA0266FFB8334C0FB66F90D" authorityName="Holthuis" authorityYear="1984" box="[1142,1230,1758,1781]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFB8334C0FB66F90D" box="[1142,1230,1758,1781]" italics="true" pageId="2" pageNumber="119">M. poeti</emphasis>
</taxonomicName>
), the slender propodus in third pereiopods (3.4 times as long as dactylus vs.3 times in
<taxonomicName id="B0E84D42FFA0266FFC953500FC11F8CD" authorityName="Holthuis" authorityYear="1984" box="[864,953,1822,1845]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="poeti">
<emphasis id="459CEAD3FFA0266FFC953500FC11F8CD" box="[864,953,1822,1845]" italics="true" pageId="2" pageNumber="119">M. poeti</emphasis>
</taxonomicName>
). Compared to epigean species of the group,
<taxonomicName id="B0E84D42FFA0266FFA533500FC09F8AD" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="459CEAD3FFA0266FFA533500FC09F8AD" italics="true" pageId="2" pageNumber="119">M. spelaeus</emphasis>
</taxonomicName>
morphologically resembles
<taxonomicName id="B0E84D42FFA0266FFB163520FB94F88D" authority="Dang &amp; Nguyen, 1972" authorityName="Dang &amp; Nguyen" authorityYear="1972" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="dienbienphuense">
<emphasis id="459CEAD3FFA0266FFB163520FA6AF8AD" box="[1251,1474,1853,1877]" italics="true" pageId="2" pageNumber="119">M. dienbienphuense</emphasis>
<bibRefCitation id="13794B30FFA0266FFCB63543FB94F88D" author="Dang NT &amp; Nguyen HY" box="[835,1084,1885,1909]" pageId="2" pageNumber="119" pagination="1 - 6" refId="ref2895" refString="Dang NT &amp; Nguyen HY (1972) Dan lieu ve tom nuoc ngot giong Macrobrachium Bate (Palaemonidae) o cac thuy vuc vung nui mien bac Viet Nam (Faunistic study of freshwater prawn genus Macrobrachium Bate from mountainous region of North Vietnam). Tap san Sinh Vat-Dai Hoc, 10 (1 - 4): 1 - 6. [In Vietnamese with French summary]." type="journal article" year="1972">Dang &amp; Nguyen, 1972</bibRefCitation>
</taxonomicName>
, a species originally described from northern
<collectingCountry id="0FFF7651FFA0266FFC5A3563FBBAF86D" box="[943,1042,1917,1941]" name="Vietnam" pageId="2" pageNumber="119">Vietnam</collectingCountry>
, but also reported from
<collectingCountry id="0FFF7651FFA0266FFAC43563FADEF86D" box="[1329,1398,1917,1941]" name="China" pageId="2" pageNumber="119">China</collectingCountry>
,
<collectingCountry id="0FFF7651FFA0266FFA733563FA16F86C" box="[1414,1470,1917,1940]" name="Laos" pageId="2" pageNumber="119">Laos</collectingCountry>
, and
<collectingCountry id="0FFF7651FFA0266FFC833583FC70F84D" box="[886,984,1949,1973]" name="Thailand" pageId="2" pageNumber="119">Thailand</collectingCountry>
(
<bibRefCitation id="13794B30FFA0266FFC1F3583FB0AF84D" author="Cai Y &amp; Naiyanetr P &amp; Ng PKL" box="[1002,1186,1949,1973]" pageId="2" pageNumber="119" pagination="581 - 649" refId="ref2654" refString="Cai Y, Naiyanetr P &amp; Ng PKL (2004) The freshwater prawns of the genus Macrobrachium Bate, 1868, of Thailand (Crustacea: Decapoda: Palaemonidae). Journal of Natural History, 38: 581 - 649." type="journal article" year="2004">Cai et al., 2004</bibRefCitation>
;
<bibRefCitation id="13794B30FFA0266FFB473583FA1DF84D" author="Hanamura Y &amp; Imai H &amp; Lasasimma O &amp; Souliyamath P &amp; Sayaka I" box="[1202,1461,1949,1973]" pageId="2" pageNumber="119" pagination="1 - 37" refId="ref3039" refString="Hanamura Y, Imai H, Lasasimma O, Souliyamath P &amp; Sayaka I (2011) Freshwater prawns of the genus Macrobrachium Bate, 1868 (Crustacea, Decapoda, Palaemonidae) from Laos. Zootaxa, 3025: 1 - 37." type="journal article" year="2011">Hanamura et al., 2011</bibRefCitation>
), especially when taking into account of the form of second pereiopods. Besides the eyes,
<taxonomicName id="B0E84D42FFA0266FFB8E35C0FB50F80D" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[1147,1272,2013,2037]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="459CEAD3FFA0266FFB8E35C0FB50F80D" box="[1147,1272,2013,2037]" italics="true" pageId="2" pageNumber="119">M. spelaeus</emphasis>
</taxonomicName>
could be separated from
<taxonomicName id="B0E84D42FFA0266FFC7435E0FBF2F7ED" authorityName="Dang &amp; Nguyen" authorityYear="1972" box="[897,1114,2045,2069]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="dienbienphuense">
<emphasis id="459CEAD3FFA0266FFC7435E0FBF2F7ED" box="[897,1114,2045,2069]" italics="true" pageId="2" pageNumber="119">M. dienbienphuense</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="13794B30FFA0266FFB6435E3FA38F7ED" author="Dang NT &amp; Nguyen HY" box="[1169,1424,2045,2069]" pageId="2" pageNumber="119" pagination="1 - 6" refId="ref2895" refString="Dang NT &amp; Nguyen HY (1972) Dan lieu ve tom nuoc ngot giong Macrobrachium Bate (Palaemonidae) o cac thuy vuc vung nui mien bac Viet Nam (Faunistic study of freshwater prawn genus Macrobrachium Bate from mountainous region of North Vietnam). Tap san Sinh Vat-Dai Hoc, 10 (1 - 4): 1 - 6. [In Vietnamese with French summary]." type="journal article" year="1972">Dang &amp; Nguyen, 1972</bibRefCitation>
;
<bibRefCitation id="13794B30FFA0266FFA6835E3FC12F7CD" author="Cai Y &amp; Naiyanetr P &amp; Ng PKL" pageId="2" pageNumber="119" pagination="581 - 649" refId="ref2654" refString="Cai Y, Naiyanetr P &amp; Ng PKL (2004) The freshwater prawns of the genus Macrobrachium Bate, 1868, of Thailand (Crustacea: Decapoda: Palaemonidae). Journal of Natural History, 38: 581 - 649." type="journal article" year="2004">Cai et al. 2004</bibRefCitation>
,
<bibRefCitation id="13794B30FFA0266FFC323A03FB1EF7CD" author="Hanamura Y &amp; Imai H &amp; Lasasimma O &amp; Souliyamath P &amp; Sayaka I" box="[967,1206,2077,2101]" pageId="2" pageNumber="119" pagination="1 - 37" refId="ref3039" refString="Hanamura Y, Imai H, Lasasimma O, Souliyamath P &amp; Sayaka I (2011) Freshwater prawns of the genus Macrobrachium Bate, 1868 (Crustacea, Decapoda, Palaemonidae) from Laos. Zootaxa, 3025: 1 - 37." type="journal article" year="2011">Hanamura et al. 2011</bibRefCitation>
) by the form of rostrum (straight vs. convex); shorter major second pereiopods in male specimens (as long as body length vs. distinctly longer in
<taxonomicName id="B0E84D42FFA62669FF5D36A0FE21FB2D" authorityName="Dang &amp; Nguyen" authorityYear="1972" box="[168,393,1213,1237]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="dienbienphuense">
<emphasis id="459CEAD3FFA62669FF5D36A0FE21FB2D" box="[168,393,1213,1237]" italics="true" pageId="4" pageNumber="121">M. dienbienphuense</emphasis>
</taxonomicName>
) less setae/pubescence in second pereiopods (see fig. 3 vs. fig.
<quantity id="B0109B24FFA62669FE3A36C3FE53FB0D" box="[463,507,1245,1269]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="121" unit="in" value="3.0">3 in</quantity>
<bibRefCitation id="13794B30FFA62669FDF636C3FD52FB0D" author="Dang NT &amp; Nguyen HY" box="[515,762,1245,1269]" pageId="4" pageNumber="121" pagination="1 - 6" refId="ref2895" refString="Dang NT &amp; Nguyen HY (1972) Dan lieu ve tom nuoc ngot giong Macrobrachium Bate (Palaemonidae) o cac thuy vuc vung nui mien bac Viet Nam (Faunistic study of freshwater prawn genus Macrobrachium Bate from mountainous region of North Vietnam). Tap san Sinh Vat-Dai Hoc, 10 (1 - 4): 1 - 6. [In Vietnamese with French summary]." type="journal article" year="1972">Dang &amp; Nguyen 1972</bibRefCitation>
); fewer number of teeth on the cutting edges of the fingers (15 vs.
<quantity id="B0109B24FFA62669FF2E3703FE94FACD" box="[219,316,1309,1333]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" metricValueMax="8.128" metricValueMin="4.572" pageId="4" pageNumber="121" unit="in" value="25.0" valueMax="32.0" valueMin="18.0">18–32 in</quantity>
<taxonomicName id="B0E84D42FFA62669FEB33700FDB5FACD" authorityName="Dang &amp; Nguyen" authorityYear="1972" box="[326,541,1309,1333]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="dienbienphuense">
<emphasis id="459CEAD3FFA62669FEB33700FDB5FACD" box="[326,541,1309,1333]" italics="true" pageId="4" pageNumber="121">M. dienbienphuense</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="23976649FFA1266EFF843A0EFC5FF7BB" ID-DOI="http://doi.org/10.5281/zenodo.4502014" ID-Zenodo-Dep="4502014" httpUri="https://zenodo.org/record/4502014/files/figure.png" pageId="3" pageNumber="120" startId="3.[113,148,2064,2086]" targetBox="[114,1453,207,2051]" targetPageId="3">
<paragraph id="775736C1FFA1266EFF843A0EFC5FF7BB" blockId="3.[113,1451,2064,2115]" pageId="3" pageNumber="120">
Fig. 2.
<taxonomicName id="B0E84D42FFA1266EFF4C3A0EFE18F7DD" authority="Cai &amp; Vidthayanon, 2016" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[185,432,2064,2085]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="120" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
<emphasis id="459CEAD3FFA1266EFF4C3A0EFE18F7DD" box="[185,432,2064,2085]" italics="true" pageId="3" pageNumber="120">Macrobrachium spelaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5EAF57A8FFA1266EFE4B3A0EFD9EF7DE" box="[446,566,2064,2086]" pageId="3" pageNumber="120" rank="species">new species</taxonomicNameLabel>
. A, first pereiopods; B, C, second pereiopods; D, E, third pereiopods, Scale bars = 2 mm [A, C]; 4 mm [B]; 1.5 mm [D]; 0.5 mm [E] (holotype: male, cl 13.7 mm, NIFI SH 00176).
</paragraph>
</caption>
<caption id="23976649FFA62669FF7D362EFAF9FBBE" ID-DOI="http://doi.org/10.5281/zenodo.4502016" ID-Zenodo-Dep="4502016" box="[136,1361,1072,1094]" httpUri="https://zenodo.org/record/4502016/files/figure.png" pageId="4" pageNumber="121" startId="4.[136,171,1072,1094]" targetBox="[136,1474,189,1056]" targetPageId="4">
<paragraph id="775736C1FFA62669FF7D362EFAF9FBBE" blockId="4.[136,1361,1072,1094]" box="[136,1361,1072,1094]" pageId="4" pageNumber="121">
Fig. 3. Colour photograph of
<taxonomicName id="B0E84D42FFA62669FE59362EFD09FBBD" authority="Cai &amp; Vidthayanon, 2016" authorityName="Cai &amp; Vidthayanon" authorityYear="2016" box="[428,673,1072,1093]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
<emphasis id="459CEAD3FFA62669FE59362EFD09FBBD" box="[428,673,1072,1093]" italics="true" pageId="4" pageNumber="121">Macrobrachium spelaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5EAF57A8FFA62669FD5A362EFC8CFBBE" box="[687,804,1072,1094]" pageId="4" pageNumber="121" rank="species">new species</taxonomicNameLabel>
(paratype, male, cl 11.7mm, OUMNH.ZC.2016-01-022)
</paragraph>
</caption>
<paragraph id="775736C1FFA62669FF7D3743FEA8F9AD" blockId="4.[134,776,1373,1621]" pageId="4" pageNumber="121">
<emphasis id="459CEAD3FFA62669FF7D3743FD96FA8D" bold="true" box="[136,574,1373,1397]" pageId="4" pageNumber="121">Comparative specimens examined.</emphasis>
<taxonomicName id="B0E84D42FFA62669FDB93743FE95FA6D" authorityName="Dang &amp; Nguyen" authorityYear="1972" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="dienbienphuense">
<emphasis id="459CEAD3FFA62669FDB93743FE95FA6D" italics="true" pageId="4" pageNumber="121">Macrobrachium dienbienphuense</emphasis>
</taxonomicName>
:
<materialsCitation id="C7803C9CFFA62669FEBA3763FEFEFA0D" ID-GBIF-Occurrence="3029797305" collectingDate="1998-06-20" collectionCode="ZRC" collectorName="Y. Cai" country="Thailand" location="Chai Badan" municipality="Central" pageId="4" pageNumber="121" specimenCode="ZRC 2000.2693" specimenCount="10" specimenCount-male="10" stateProvince="Lop Buri">
<specimenCount id="61EEFD48FFA62669FEBA3763FE1DFA6D" box="[335,437,1405,1429]" pageId="4" pageNumber="121" type="male">10 males</specimenCount>
, cl 9.0–11.0 mm, 5 ovigerous females, cl
<quantity id="B0109B24FFA62669FF083783FED4FA4C" box="[253,380,1437,1461]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.3" metricValueMin="6.7" pageId="4" pageNumber="121" unit="mm" value="7.5" valueMax="8.3" valueMin="6.7">6.7–8.3 mm</quantity>
, eggs 1.3 × 1.0 mm (
<specimenCode id="274E9EBAFFA62669FDBB3783FD54FA4D" box="[590,764,1437,1461]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="121">ZRC 2000.2693</specimenCode>
),
<collectingMunicipality id="9733ACBBFFA62669FF7337A3FF7DFA2D" box="[134,213,1469,1493]" pageId="4" pageNumber="121">Central</collectingMunicipality>
<collectingCountry id="0FFF7651FFA62669FF2837A3FE97FA2D" box="[221,319,1469,1493]" name="Thailand" pageId="4" pageNumber="121">Thailand</collectingCountry>
,
<collectingRegion id="B52CF823FFA62669FEBF37A3FDB5FA2D" box="[330,541,1469,1493]" country="Thailand" name="Lop Buri" pageId="4" pageNumber="121">Lop Buri Province</collectingRegion>
,
<location id="7237601AFFA62669FDDD37A3FD01FA2D" LSID="urn:lsid:plazi:treatment:FF4187D7FFA22669FC5937FBFEA8F9AD:7237601AFFA62669FDDD37A3FD01FA2D" box="[552,681,1469,1493]" country="Thailand" municipality="Central" name="Chai Badan" pageId="4" pageNumber="121" stateProvince="Lop Buri">Chai Badan</location>
, coll.
<collectorName id="DA1D5317FFA62669FD1E37A3FF19FA0D" pageId="4" pageNumber="121">Y. Cai</collectorName>
,
<date id="03561001FFA62669FF4A37C3FEFEFA0D" box="[191,342,1501,1525]" pageId="4" pageNumber="121" value="1998-06-20">
<collectingDate id="1312E9E9FFA62669FF4A37C3FEFEFA0D" box="[191,342,1501,1525]" pageId="4" pageNumber="121" value="1998-06-20">20 June 1998</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C7803C9CFFA62669FE9037C3FF54F9AD" ID-GBIF-Occurrence="3029797303" collectingDate="1998-06-16" collectorName="Y. Cai" country="Thailand" location="Thailand" pageId="4" pageNumber="121" specimenCount="9" specimenCount-male="9">
<specimenCount id="61EEFD48FFA62669FE9037C3FE15FA0D" box="[357,445,1501,1525]" pageId="4" pageNumber="121" type="male">9 males</specimenCount>
, cl
<quantity id="B0109B24FFA62669FE1C37C3FD21FA0C" box="[489,649,1501,1525]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.17" metricValueMin="1.02" pageId="4" pageNumber="121" unit="mm" value="10.95" valueMax="11.7" valueMin="10.2">10.2–11.7 mm</quantity>
, North-east
<collectingCountry id="0FFF7651FFA62669FF7D37E3FF42F9ED" box="[136,234,1533,1557]" name="Thailand" pageId="4" pageNumber="121">Thailand</collectingCountry>
, Phibun, Mangsahan, Ubon Rachathani Mae Nam Moon, Mangsahan afternoon market, coll.
<collectorName id="DA1D5317FFA62669FDAC3403FD09F9CD" box="[601,673,1565,1589]" pageId="4" pageNumber="121">Y. Cai</collectorName>
et al.,
<date id="03561001FFA62669FD1E3403FF54F9AD" pageId="4" pageNumber="121" value="1998-06-16">
<collectingDate id="1312E9E9FFA62669FD1E3403FF54F9AD" pageId="4" pageNumber="121" value="1998-06-16">16 June 1998</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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