191 lines
25 KiB
XML
191 lines
25 KiB
XML
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<mods:title id="6BBE67B910DA6D294FCCAFAE4618E701">A new species of Lasiosmylus from the Early Cretaceous, China clarifies its genus-group placement in Ithonidae (Neuroptera)</mods:title>
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<mods:namePart id="444B998B1AA8B13786A71AD5AA48DC9C">Zheng, Bingyu</mods:namePart>
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<mods:namePart id="86B5CFD5A93068B931D4876CB050C7A8">Ren, Dong</mods:namePart>
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<mods:namePart id="0808D5F8216722F1B9E9416B4E18CF58">Wang, Yongjie</mods:namePart>
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<mods:date id="BAB36DA1C7A84B2438FB935E36BEC511">2016</mods:date>
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<treatment id="FB3B1723AD6DE3CDABDABF92882C97C0" ID-GBIF-Taxon="127859785" LSID="urn:lsid:plazi:treatment:FB3B1723AD6DE3CDABDABF92882C97C0" httpUri="http://treatment.plazi.org/id/FB3B1723AD6DE3CDABDABF92882C97C0" lastPageId="5" lastPageNumber="46" pageId="2" pageNumber="43">
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<subSubSection id="CD8EE504932D5D60B9D205F7024D0141" pageId="2" pageNumber="43" type="nomenclature">
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<paragraph id="07460AA7F93D79382CE9E63B93506E0E" pageId="2" pageNumber="43">
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Genus
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<taxonomicName id="301CC9BF49780B17DCA8B8E3C691E316" ID-CoL="62XF8" authority="Ren & Guo, 1996" authorityName="Ren & Guo" authorityYear="1996" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">Lasiosmylus Ren & Guo, 1996</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="46110950D7A1DA5F09AF2F379F54A33E" pageId="2" pageNumber="43" type="type species">
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<paragraph id="8F184638068F63C12413F854DD2018C6" pageId="2" pageNumber="43">Type species.</paragraph>
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<paragraph id="63786FC63C90E51C6AC16BB1D7FDF0C5" pageId="2" pageNumber="43">
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<taxonomicName id="532007932534744A9FC45EBE324E23E4" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
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Ren & Guo, 1996.
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</paragraph>
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</subSubSection>
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<subSubSection id="C686BAFEE54AD0A7624FC5D6506D2146" pageId="2" pageNumber="43" type="species included">
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<paragraph id="7C287AC89D0FD6B155085DBFB9CB63F0" pageId="2" pageNumber="43">Species included.</paragraph>
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<paragraph id="CF76292F193A06C95C0F9BE5187EFC44" pageId="2" pageNumber="43">
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<taxonomicName id="A8C7578D79C626AA739DC04065118FDE" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
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Ren & Guo, 1996,
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<taxonomicName id="11174D51EDC633A3A78BB12F872C7E37" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus longus" order="Neuroptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="longus">Lasiosmylus longus</taxonomicName>
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sp. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="E0B1D28AB27304E38247DAF19158B744" pageId="2" pageNumber="43" type="amended diagnosis">
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<paragraph id="1EB4987233DEE28D21806C807397215F" pageId="2" pageNumber="43">Amended diagnosis.</paragraph>
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<paragraph id="4B6C1AEFBD067CC37B161CF350AAFCD1" pageId="2" pageNumber="43">Body stout (ca. 11-17 mm long), covered with dense setae; head hypognathous, protruding from pronotum partly; antenna filiform (ca. 2-5 mm, incompletely preserved); compound eye large, ocelli absent; thorax robust, long setae concentrated on pronotum. Forewing ca. 12-23 mm long, 5-8 mm wide, membranous area with many fuscous spots; humeral plate distinct; dense setae along the veins, especially on the wing margin; trichosors and nygmata undetectable; costal space dilated basally and narrowed distally; humeral veinlet recurrent, with several simple branches; costal cross-veins simple, moderately curved distally in the apical half of the costal space; Sc and R1 separate distally, entering the margin before the wing apex; one or two sc-r1 crossveins; R1 with four to eleven pectinate branches distally; the origin of Rs distant from the wing base, with seven to thirteen branches regularly arranged; relatively few crossveins present in radial area; MA simple, dichotomously branched terminally; MP first fork distant from wing base. Hind wing ca. 11-18 mm long, 4-8 mm wide, partly preserved, venation similar to forewing except for the following characters: costal space narrow, only slightly expand in proximal portion.</paragraph>
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</subSubSection>
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<subSubSection id="80B1F759918AD08093FDDEADF21DFA0D" lastPageId="5" lastPageNumber="46" pageId="2" pageNumber="43" type="remarks">
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<paragraph id="F108C36A91AAF0A0755D576E05715311" pageId="2" pageNumber="43">Remarks.</paragraph>
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<paragraph id="D4B4BD401A439B0D21475904945EBAEA" pageId="2" pageNumber="43">
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<taxonomicName id="F154AA01149DB3E4607F2B32536C7DB2" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">Lasiosmylus</taxonomicName>
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shows a superficial similarity with osmylids, sharing plesiomorphic features such as the fork of MP in forewing usually between the separation of MA and first Rs branch, sometimes opposite the separation of MA; wings not falcate, with few crossveins (
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<bibRefCitation id="BE777B0E974C14A307AB0A0265E7BCC9" author="Ren, D" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="7" pageNumber="48" pagination="461 - 479" title="On the new fossil genera and species of Neuroptera (Insecta) from the Late Jurassic of northeast China." volume="21" year="1996">Ren and Guo 1996</bibRefCitation>
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). However, all these characters do not well support the assignment of
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<taxonomicName id="B9B752C6E0036F3741D8E1E37851B8BB" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">Lasiosmylus</taxonomicName>
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to
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<taxonomicName id="925EA1679082F7D723971DC0A4C70F90" lsidName="" pageId="2" pageNumber="43" rank="subfamily" subfamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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, or
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<taxonomicName id="55CF1DA35FB498E96106838639B226A8" family="Osmylidae" lsidName="" pageId="2" pageNumber="43" rank="family">Osmylidae</taxonomicName>
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in general because they also frequently occur in other families (e.g.,
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<taxonomicName id="8A17CDB26A92E22EBEB3A0097C0EE786" family="Ithonidae" lsidName="" pageId="2" pageNumber="43" rank="family">Ithonidae</taxonomicName>
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,
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<taxonomicName id="C9C4994DDAB4570ADEC33CD77174F3B9" family="Berothidae" lsidName="" pageId="2" pageNumber="43" rank="family">Berothidae</taxonomicName>
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, some
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<taxonomicName id="BA1A8B7AC630C2F20467514721365C7D" family="Mantispidae" lsidName="" pageId="2" pageNumber="43" rank="family">Mantispidae</taxonomicName>
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). The subsequent transfer to
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<taxonomicName id="A653C153046C0AD11C5C407D6A7A776B" family="Ithonidae" lsidName="" pageId="2" pageNumber="43" rank="family">Ithonidae</taxonomicName>
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by
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<bibRefCitation id="3FD585D27801FB758A65E479EF535022" author="Makarkin, VN" journalOrPublisher="Cretaceous Research" pageId="7" pageNumber="48" pagination="57 - 68" title="A comparative overview of the neuropteran assemblage of the Early Cretaceous Yixian Formation (China), with description of a new genus of Psychopsidae (Insecta: Neuroptera)." url="10.1016/j.cretres.2011.11.013" volume="35" year="2012">Makarkin et al. (2012</bibRefCitation>
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,
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<bibRefCitation id="54A9B7D6020C8C69FEBA5B26F6F2A7C2" author="Makarkin, VN" journalOrPublisher="Zootaxa" pageId="7" pageNumber="48" pagination="385 - 393" title="First record of the family Ithonidae (Neuroptera) from Baltic amber." url="10.11646/zootaxa.3796.2.10" volume="3796" year="2014">2014</bibRefCitation>
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) seems reasonable; moreover, recently it was classified further as belonging to the polystoechotid genus-group by
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<bibRefCitation id="036F1A9A4D8E94FB3BBFE61E94DA36EE" author="Zheng, BY" journalOrPublisher="PLoS ONE" pageId="7" pageNumber="48" title="Acta Palaeontologica Polonica" url="10.4202/app.00259.2016" year="2016">Zheng et al. (2016)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="6B553F50C44E1934C7142302D9BE27B2" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
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Herein, nine new-collected specimens are examined in this study. All these specimens are placed in
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<taxonomicName id="C38FADC3AF9553347EA2E23D184AEC76" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">Lasiosmylus</taxonomicName>
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based on the following characters: numerous dispersed spots on the forewing, simple costal crossveins, two subcostal crossveins, Rs less than ten branches (about six to eight branches), MP distant from the wing base and beyond MA fork, MP1 and MP2 simple, one mp1-mp2 crossvein, CuA dichotomously branched distally (in particular, obs. CNU-NEU-LB2015001P/C and CNU-NEU-LB2015002, see Figs 1, 2; and
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<bibRefCitation id="D780FCA06F54F3141EB4DBA1391AEA63" author="Ren, D" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="7" pageNumber="48" pagination="461 - 479" title="On the new fossil genera and species of Neuroptera (Insecta) from the Late Jurassic of northeast China." volume="21" year="1996">Ren and Guo 1996</bibRefCitation>
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: fig. 5, fig. 10, pl. 3, fig. 11, pl. 2). Noticeably, during checking the specimens, we found some variable characters that are distinctly different from the type specimen, e.g., humeral veinlet and separated Sc and R1. A recurrent humeral veinlet is considered as a synapomorphy for
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<taxonomicName id="D6AAA76E043765558084DECBB7807AD8" family="Ithonidae" lsidName="" pageId="2" pageNumber="43" rank="family">Ithonidae</taxonomicName>
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<pageBreakToken id="F061F5FC783EEB81F662DB51AF12B567" pageId="3" pageNumber="44" start="start">(</pageBreakToken>
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<bibRefCitation id="8EA31AB736C31CB2A622966B6B72C221" author="Yang, Q" journalOrPublisher="PLoS ONE" pageId="7" pageNumber="48" title="A remarkable new family of Jurassic insects (Neuroptera) with primitive wing venation and its phylogenetic position in Neuropterida." url="10.1371/journal.pone.0044762" volume="7" year="2012">Yang et al. 2012</bibRefCitation>
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, Makarkin et al. 2013,
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<bibRefCitation id="75AECD19DCEAB798E6655EDE94B5EDFA" author="Zheng, BY" journalOrPublisher="PLoS ONE" pageId="7" pageNumber="48" title="Acta Palaeontologica Polonica" url="10.4202/app.00259.2016" year="2016">Zheng et al. 2016</bibRefCitation>
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). However, this character is absent in the line drawing of
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<taxonomicName id="27BC5FE878482CB45E509395A7F85050" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
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(
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<bibRefCitation id="63D29AF6E179CAC3C8F2A4C149C40490" author="Ren, D" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="7" pageNumber="48" pagination="461 - 479" title="On the new fossil genera and species of Neuroptera (Insecta) from the Late Jurassic of northeast China." volume="21" year="1996">Ren and Guo 1996</bibRefCitation>
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: fig. 5), although some trace of recurrent humeral veinlet can be detected in the photograph of
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<taxonomicName id="00259D42F452A38778191FC2D1D0E4B8" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">Lasiosmylus</taxonomicName>
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(
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<bibRefCitation id="EE9D9B20D117828D5D117E6539405851" author="Ren, D" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="7" pageNumber="48" pagination="461 - 479" title="On the new fossil genera and species of Neuroptera (Insecta) from the Late Jurassic of northeast China." volume="21" year="1996">Ren and Guo 1996</bibRefCitation>
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: fig. 11, pl. 4). Regretfully, the holotype of
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<taxonomicName id="988F6E4F340F0777D7D69A4B10171815" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
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was not available for examination during this study (possibly lost). However, it is reasonable to assume now that the recurrent humeral veinlet occurs in
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<taxonomicName id="8D7A828E4634D76F0331F71D04DCBFAE" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
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according to these new specimens.
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</paragraph>
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<caption id="08C085DF87D7958FED236838413B13FD" pageId="3" pageNumber="44">
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<paragraph id="E9F182C6FBE6A39E79886EAA655BE7AD" pageId="3" pageNumber="44">
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Figure 1. New materials of
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<taxonomicName id="2B45AF5F7AF9D4C8B1EF0CB1EEAE096A" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
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: CNU-NEU-LB2015001P/C, CNU-NEU-LB2015002. A composite photographs of habitus of part and counterpart (CNU-NEU-LB2015001P/C) hp, humeral plate (CNU-NEU-LB2015001C) B line drawing (CNU-NEU-LB2015001C) C habitus photograph, hp, humeral plate (CNU-NEU-LB2015002) D line drawing (CNU-NEU-LB2015002). Scale bars: 5 mm (
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<normalizedToken id="BA1CF4418C62C58967BA5FD2CEE6A1A5" originalValue="A–D">A-D</normalizedToken>
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).
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</paragraph>
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</caption>
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<caption id="596BE511CAE077A9114DA2AD6A2B2862" pageId="3" pageNumber="44">
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<paragraph id="E58838307462B33042D93206BC098781" pageId="3" pageNumber="44">
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Figure 2. New materials of
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<taxonomicName id="F49BAF5D914F720D1657ACCF7B957C0F" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
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. Line drawings of CNU-NEU-LB2015001P/C, A left forewing B right forewing C right hind wing D left hind wing. Line drawings of CNU-NEU-LB2015002 E left forewing F right forewing G left hind wing H right hind wing. Scale bars: 5 mm (
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<normalizedToken id="40FED2F24864E0CA6EF01A952628BCA9" originalValue="A–H">A-H</normalizedToken>
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).
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</paragraph>
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</caption>
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<paragraph id="2D726F2DBC7A71376A079A714B572D94" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44">
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In addition, the distally separated Sc and R1 were regarded as a synapomorphic character of
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<taxonomicName id="5E7A21C184C62A0D17CF949A024EB189" family="Ithonidae" lsidName="" pageId="3" pageNumber="44" rank="family">Ithonidae</taxonomicName>
|
||
(
|
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<bibRefCitation id="644E5EF3172226848E57D2DB0DAF100D" author="Zheng, BY" journalOrPublisher="PLoS ONE" pageId="7" pageNumber="48" title="Acta Palaeontologica Polonica" url="10.4202/app.00259.2016" year="2016">Zheng et al. 2016</bibRefCitation>
|
||
). In the original illustration of
|
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<taxonomicName id="135569A719AA4387D905CC12EBC501EF" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
|
||
, Sc and R1 were drawn with fused termination. Unfortunately, the photograph of
|
||
<taxonomicName id="8D908ECF273EE729F820BA45943572AF" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
|
||
is too obscure for us to discern the condition of Sc and R1 (
|
||
<bibRefCitation id="B1AA59A351F09D0BA1814D9D3FB061E1" author="Ren, D" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="7" pageNumber="48" pagination="461 - 479" title="On the new fossil genera and species of Neuroptera (Insecta) from the Late Jurassic of northeast China." volume="21" year="1996">Ren and Guo 1996</bibRefCitation>
|
||
: fig. 10,
|
||
<pageBreakToken id="5406A3C74CC675D34ADE5724B5302FE1" pageId="4" pageNumber="45" start="start">pl</pageBreakToken>
|
||
. 3, fig. 11, pl. 2). In extant members of the polystoechotid genus-group Sc and R1 are closely approximated but are actually not fused, e.g.,
|
||
<taxonomicName id="C7278A4AC536D7AE8A8793CE26848F7D" class="Insecta" family="Ithonidae" genus="Fontecilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fontecilla" order="Neuroptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">Fontecilla</taxonomicName>
|
||
<normalizedToken id="900B9211FA73D89131275E51031BC341" originalValue="Navás">Navas</normalizedToken>
|
||
, 1931,
|
||
<taxonomicName id="2C92288E6F46B1B8104BEBB0DFACAD34" class="Insecta" family="Ithonidae" genus="Platystoechotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platystoechotes" order="Neuroptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">Platystoechotes</taxonomicName>
|
||
Carpenter, 1940,
|
||
<taxonomicName id="385272D58A63EAA213C4EA39B2B9D5D7" class="Insecta" family="Ithonidae" genus="Polystoechotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polystoechotes" order="Neuroptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">Polystoechotes</taxonomicName>
|
||
Burmeister, 1839 (see
|
||
<bibRefCitation id="DEDC8AB090EF4AA2DFF050A42C3ADD35" author="Winterton, SL" journalOrPublisher="Annals of the Entomological Society of America" pageId="7" pageNumber="48" pagination="511 - 522" title="Phylogeny of moth lacewings and giant lacewings (Neuroptera: Ithonidae, Polystoechotidae) using DNA sequence data, morphology and fossils." url="10.1603/AN10026" volume="103" year="2010">Winterton and Makarkin 2010</bibRefCitation>
|
||
). While this character was not fully investigated in the fossil lineages, most fossil polystoechotid genera were illustrated with the fused Sc and R1.
|
||
</paragraph>
|
||
<paragraph id="3DCD58931E2C0C1E9D422AFAB58CB844" pageId="4" pageNumber="45">
|
||
During the examination of the new materials, it is clear that all specimens assigned to
|
||
<taxonomicName id="9C6F961FC8C84E95D0C84CFBFA56CF47" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">Lasiosmylus</taxonomicName>
|
||
(Figs 1, 2) show a separate Sc and R1. Furthermore, nine specimens (CNU-NEU-LB2015001P/C, CNU-NEU-LB2015002, CNU-NEU-LB2016001P/C, CNU-NEU-LB2016002, CNU-NEU-LB2016003, CNU-NEU-LB2016004, CNU-NEU-LB2016005, CNU-NEU-LB2016006, CNUNEU-LB2016007) exhibit the typically venation with
|
||
<taxonomicName id="C3CB4DE1839E10E6FF41C79A1E674C56" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
|
||
with exception for the incompatible conditions of Sc and R1. These nine specimens are considered to be
|
||
<taxonomicName id="1914E11F53B7212FD30AAE6927D6A737" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="02F5E79FBBB7D8DDB00AF44CD95FB49E" pageId="5" pageNumber="46">
|
||
<pageBreakToken id="8E9D9B0820E15401B23B9514BC333471" pageId="5" pageNumber="46" start="start">It</pageBreakToken>
|
||
is concluded here that the genus
|
||
<taxonomicName id="AA904AEE04371C112D297C6C42B528B2" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="genus">Lasiosmylus</taxonomicName>
|
||
most commonly has the separated Sc and R1 that is consistent with other moth lacewings. The exception of Sc and R1 in the holotype of
|
||
<taxonomicName id="4CC4D17063DA6327586D1FA6EEB901C6" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus newi" order="Neuroptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="newi">Lasiosmylus newi</taxonomicName>
|
||
possibly represents a particularly individual variation, inaccuracy in line drawing or obscurity in the specimen. Based on this we consider
|
||
<taxonomicName id="F1084D74A09FFF466A4B7AD94FAA0858" class="Insecta" family="Ithonidae" genus="Lasiosmylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasiosmylus" order="Neuroptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="genus">Lasiosmylus</taxonomicName>
|
||
is unquestionably assigned to the ithonid genus-group by the following combination of characters: robust and hairy body, retracted head under pronotum, costal space dilated basally and narrowed disproportionately distally, separated Sc and R1 reaching the anterior margin straightly before the wing apex, MP first fork distant from the wing base and beyond the divergence of MA.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |