268 lines
35 KiB
XML
268 lines
35 KiB
XML
<document id="19D7BC83CE7B4DEBD52F75B481C602CA" ID-DOI="10.11646/zootaxa.5297.3.8" ID-ISSN="1175-5326" ID-ZooBank="EC0AD75A-AB80-45CD-906F-39DFD6BCE6CC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685960276401" checkinUser="plazi" docAuthor="Alekseev, Vitalii I. & Nabozhenko, Maxim V." docDate="2023" docId="F21687DACB37771DFF4E3527FA0EFBBF" docLanguage="en" docName="zootaxa.5297.3.8.pdf" docOrigin="Zootaxa 5297 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5297.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zopheromimus auriborussiensis Alekseev & Nabozhenko 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="442" masterDocId="0E2FFFA2CB34771AFFD9372CFF92FF8A" masterDocTitle="News from Baltic amber: a new tribe of extinct Paleogene ironclad beetles (Coleoptera: Zopheridae: Zopherinae)" masterLastPageNumber="445" masterPageNumber="435" pageNumber="438" updateTime="1685960335237" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
|
||
<mods:mods id="434B29B3DDF8461C20CD700DB2982575" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="810DE6CF123505E61E020CFFDF519309">
|
||
<mods:title id="F77CD4D17B1E8EA2100692956F64CC7B">News from Baltic amber: a new tribe of extinct Paleogene ironclad beetles (Coleoptera: Zopheridae: Zopherinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="BD454626F05C5B989B339DC4E7C892A5" type="personal">
|
||
<mods:role id="94D10E2212DB4F44BEC97B5AAEB42FA8">
|
||
<mods:roleTerm id="9078EC7675673100C2F9CEC0E653CCED">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2B954A4441C021CEF46FED016804AE1D">Alekseev, Vitalii I.</mods:namePart>
|
||
<mods:affiliation id="D2C289CC39CBCC473ACD1180D7D11AA8">Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia. & Immanuel Kant Baltic Federal University, A. Nevskii str. 14, Kaliningrad 236016, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="0DD99CCC86C2D8D613BEF9413E4198BD" type="personal">
|
||
<mods:role id="7E7D310862805CCC40A380B7949ADDC5">
|
||
<mods:roleTerm id="AE51859849528FF581009775364481B5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="381DE6FD0CEA11BD08203C4337E813EA">Nabozhenko, Maxim V.</mods:namePart>
|
||
<mods:affiliation id="2F7AE305D3576C08F89F3B5D3B6BA946">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. & Dagestan State University, M. Gadzhiev str. 43 a, Makhachkala, Republic of Dagestan, 367000, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="CAD9810AC7274F4914D417EF75EA5CCC">text</mods:typeOfResource>
|
||
<mods:relatedItem id="3BF82E21000E388BFF3EE113435C84F3" type="host">
|
||
<mods:titleInfo id="530B9826F5BECF62DA7AB1DDD1729165">
|
||
<mods:title id="014065D5A0C94F08D45D433821E69DF6">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="0F5C955B47B5BB162C3D45ADE24CC8F1">
|
||
<mods:date id="65894DFBD89E75989081304F5E0D4414">2023</mods:date>
|
||
<mods:detail id="11B51CC19A0D09794401D1581D37E8D0" type="pubDate">
|
||
<mods:number id="D1F8E76AEFE9A17757D934529931D1BA">2023-06-02</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="48F05BDDEB99B1276AA960B8CE881C2B" type="volume">
|
||
<mods:number id="2A1445F960C4CAB2CA6D0B7859647E7C">5297</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="9C4F0082424679588A6B94CA3864D079" type="issue">
|
||
<mods:number id="C98329771D513DE11AD3932F47D9FCD7">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="11B6DD1FA2BA77863422BBA305AE13D3" unit="page">
|
||
<mods:start id="A103FE8E50F918FB745E2E48347D6054">435</mods:start>
|
||
<mods:end id="37A87F7E7B6086A8E0A90E17828C721D">445</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="CBAC2CA71D82E03899E0D15D6FB808FC">
|
||
<mods:url id="4C0D3F0523D645067730E8174B96AE02">http://dx.doi.org/10.11646/zootaxa.5297.3.8</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="3572D5B715105813E5DE2C674B43911E">journal article</mods:classification>
|
||
<mods:identifier id="D8B8875751BB659861A110C064AAF2C5" type="DOI">10.11646/zootaxa.5297.3.8</mods:identifier>
|
||
<mods:identifier id="2BF36D047208114F155AA683D339FA67" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="BEAF6478A3E76202B801671FE85F5BE6" type="ZooBank">EC0AD75A-AB80-45CD-906F-39DFD6BCE6CC</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="F21687DACB37771DFF4E3527FA0EFBBF" LSID="urn:lsid:plazi:treatment:F21687DACB37771DFF4E3527FA0EFBBF" httpUri="http://treatment.plazi.org/id/F21687DACB37771DFF4E3527FA0EFBBF" lastPageId="7" lastPageNumber="442" pageId="3" pageNumber="438">
|
||
<subSubSection id="32A56547CB377719FF4E3527FD11FDAF" box="[151,643,523,550]" pageId="3" pageNumber="438" type="nomenclature">
|
||
<paragraph id="7A0036CCCB377719FF4E3527FD11FDAF" blockId="3.[151,643,523,585]" box="[151,643,523,550]" pageId="3" pageNumber="438">
|
||
<heading id="214881A0CB377719FF4E3527FD11FDAF" bold="true" box="[151,643,523,550]" fontSize="11" level="1" pageId="3" pageNumber="438" reason="1">
|
||
<emphasis id="48CBEADECB377719FF4E3527FD11FDAF" bold="true" box="[151,643,523,550]" pageId="3" pageNumber="438">
|
||
<taxonomicName id="BDBF4D4FCB377719FF4E3527FD8CFDAC" authority="Alekseev & Nabozhenko, 2023" authorityName="Alekseev & Nabozhenko" authorityYear="2023" box="[151,542,523,550]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="438" phylum="Arthropoda" rank="species" species="auriborussiensis" status="sp. nov.">
|
||
<emphasis id="48CBEADECB377719FF4E3527FD8CFDAC" bold="true" box="[151,542,523,550]" italics="true" pageId="3" pageNumber="438">Zopheromimus auriborussiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="53F857A5CB377719FDFC3527FD11FDAF" box="[549,643,523,549]" pageId="3" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="32A56547CB377719FF4E3503FE99FDC3" box="[151,267,559,585]" pageId="3" pageNumber="438" type="description">
|
||
<paragraph id="7A0036CCCB377719FF4E3503FE99FDC3" blockId="3.[151,643,523,585]" box="[151,267,559,585]" pageId="3" pageNumber="438">
|
||
(
|
||
<figureCitation id="E2842A49CB377719FF463503FE90FDC3" box="[159,258,559,585]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–4" captionStart-2="FIGURES 5–7" captionStartId-0="4.[151,264,1935,1959]" captionStartId-1="5.[151,264,1891,1915]" captionStartId-2="6.[151,264,1943,1967]" captionTargetBox-0="[151,1436,554,1909]" captionTargetBox-1="[177,1411,181,1855]" captionTargetBox-2="[158,1430,181,1919]" captionTargetId-0="figure-179@4.[151,1436,554,1909]" captionTargetId-1="figure-17@5.[177,1411,181,1867]" captionTargetId-2="figure-25@6.[158,1430,181,1919]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1–2. Zopheromimus auriborussiensis sp. nov., holotype, No KAM 8325/2: 1—habitus, dorso-lateral view; 2— habitus, ventro-lateral view. Scale bar = 1 mm." captionText-1="FIGURES 3–4. Zopheromimus auriborussiensis sp. nov., holotype, No KAM 8325/2, forebody, ventro-lateral view: 3— microphotograph; 4—schematic drawing. Abbreviations: A2—antennomere 2 (as long as antennomere 3); E—eye (lateral, hemispherical); PH—prothoracic hypomeron (not grooved); PC—procoxal cavity (externally closed); AP—abdominal procoxal process (triangular)." captionText-2="FIGURES 5–7. Representatives of the tribe Zopheromimini trib. nov. (microphotographs): 5—head of Zopheromimus auriborussiensis; 6—abdomen of Z. auriborussiensis; 7—abdomen of Yantaroxenos colydioides. Abbreviations: mem— membranes between non-connate ventrites; pag—preapical groove on abdominal ventrite 5." pageId="3" pageNumber="438">Figs 1–6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="32A56547CB377719FF4E355CFEADFCB1" pageId="3" pageNumber="438" type="materials_examined">
|
||
<paragraph id="7A0036CCCB377719FF4E355CFC31FD5B" blockId="3.[151,1437,624,2034]" pageId="3" pageNumber="438">
|
||
<emphasis id="48CBEADECB377719FF4E355CFED5FD00" bold="true" box="[151,327,624,650]" pageId="3" pageNumber="438">Type material.</emphasis>
|
||
<typeStatus id="A504886ECB377719FE89355CFE2FFD00" box="[336,445,624,650]" pageId="3" pageNumber="438">Holotype</typeStatus>
|
||
: No. KAM 8325/2 [KRAM] (ex. coll. Konstantin Andruschenko), adult, sex unknown. Complete beetle inclusion in a transparent, yellow, polished amber piece with approximate dimensions 46×16×
|
||
<quantity id="BD479B29CB377719FA4935BFFF54FD5B" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="438" unit="mm" value="8.0">8 mm</quantity>
|
||
; preserved without supplementary fixation. Syninclusion: absent.
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB377719FF1E35F6FC6CFD7E" blockId="3.[151,1437,624,2034]" box="[199,1022,730,757]" pageId="3" pageNumber="438">
|
||
<emphasis id="48CBEADECB377719FF1E35F6FEC4FD7E" bold="true" box="[199,342,730,756]" pageId="3" pageNumber="438">
|
||
<typeStatus id="A504886ECB377719FF1E35F6FE90FD7E" box="[199,258,730,756]" pageId="3" pageNumber="438">Type</typeStatus>
|
||
strata.
|
||
</emphasis>
|
||
Baltic amber from Eocene amber-bearing Blue Earth layers.
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB377719FF1E35D2FEADFCB1" blockId="3.[151,1437,624,2034]" pageId="3" pageNumber="438">
|
||
<materialsCitation id="CAD73C91CB377719FF1E35D2FEA9FCB1" country="Russia" location="Yantarny" municipality="Sambian" pageId="3" pageNumber="438" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
|
||
<emphasis id="48CBEADECB377719FF1E35D2FEF8FC9D" bold="true" box="[199,362,765,792]" pageId="3" pageNumber="438">
|
||
<typeStatus id="A504886ECB377719FF1E35D2FE90FC92" box="[199,258,766,792]" pageId="3" pageNumber="438">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="7F606017CB377719FEAC35D2FE48FC92" LSID="urn:lsid:plazi:treatment:F21687DACB37771DFF4E3527FA0EFBBF:7F606017CB377719FEAC35D2FE48FC92" box="[373,474,766,792]" country="Russia" municipality="Sambian" name="Yantarny" pageId="3" pageNumber="438" stateProvince="Kaliningrad">Yantarny</location>
|
||
settlement (formerly Palmnicken),
|
||
<collectingMunicipality id="9A64ACB6CB377719FCA535D2FC4CFC92" box="[892,990,766,792]" pageId="3" pageNumber="438">Sambian</collectingMunicipality>
|
||
(Samland) peninsula, the
|
||
<collectingRegion id="B87BF82ECB377719FACF35D2FF73FCB1" country="Russia" name="Kaliningrad" pageId="3" pageNumber="438">Kaliningrad region</collectingRegion>
|
||
,
|
||
<collectingCountry id="02A8765CCB377719FF34340DFEA9FCB1" box="[237,315,801,827]" name="Russia" pageId="3" pageNumber="438">Russia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="32A56547CB377719FF1E3468FCF6FC2C" pageId="3" pageNumber="438" type="etymology">
|
||
<paragraph id="7A0036CCCB377719FF1E3468FCF6FC2C" blockId="3.[151,1437,624,2034]" pageId="3" pageNumber="438">
|
||
<emphasis id="48CBEADECB377719FF1E3468FE37FCD4" bold="true" box="[199,421,836,862]" pageId="3" pageNumber="438">Derivatio nominis.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="BDBF4D4FCB377719FD473469FCC5FCD5" authorityName="Alekseev & Nabozhenko" authorityYear="2023" box="[670,855,837,863]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="438" phylum="Arthropoda" rank="species" species="auriborussiensis">
|
||
<emphasis id="48CBEADECB377719FD473469FCC5FCD5" box="[670,855,837,863]" italics="true" pageId="3" pageNumber="438">auriborussiensis</emphasis>
|
||
</taxonomicName>
|
||
” used as an adjective, is a compound word derived from the Medieval Latin
|
||
<emphasis id="48CBEADECB377719FE763445FD11FC08" box="[431,643,872,898]" italics="true" pageId="3" pageNumber="438">aurum borussiense</emphasis>
|
||
(the gold of Prussia) and referring to the German metaphoric names of Baltic amber (“das Gold Ostpreussens” or “das deutsche Gold”).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="32A56547CB37771DFF1E3483FB13FD47" lastPageId="7" lastPageNumber="442" pageId="3" pageNumber="438" type="description">
|
||
<paragraph id="7A0036CCCB377719FF1E3483FC58FBF1" blockId="3.[151,1437,624,2034]" pageId="3" pageNumber="438">
|
||
<emphasis id="48CBEADECB377719FF1E3483FECBFC43" bold="true" box="[199,345,943,969]" pageId="3" pageNumber="438">Description.</emphasis>
|
||
Measurements: body length about
|
||
<quantity id="BD479B29CB377719FD3F3483FCADFC40" box="[742,831,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="438" unit="mm" value="5.3">5.3 mm</quantity>
|
||
; maximum width across both elytra
|
||
<quantity id="BD479B29CB377719FB003483FAA1FC40" box="[1241,1331,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="438" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; pronotal length
|
||
<quantity id="BD479B29CB377719FF3D34FEFEA9FC67" box="[228,315,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="438" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, pronotum maximum width
|
||
<quantity id="BD479B29CB377719FDA134FEFD4DFC67" box="[632,735,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="438" unit="mm" value="1.15">1.15 mm</quantity>
|
||
; elytral length about
|
||
<quantity id="BD479B29CB377719FC1034FEFBB3FC67" box="[969,1057,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="438" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, elytron maximum width
|
||
<quantity id="BD479B29CB377719FA9934FFFA0BFC67" box="[1344,1433,979,1005]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="438" unit="mm" value="0.7">0.7 mm</quantity>
|
||
. Body elongate, subcylindrical, moderately convex dorsally. Integument uniformly black. Body vestiture consisting of (1) short, recumbent, scale-like, elongate-oval light setae, and (2) curved, semirecumbent, bristle-like dark setae. Dense light oval scales forming patterns on the dorsum (median stripe on pronotum and diagonally striped pattern on elytra) and almost entirely covering head and ventral side of the body.
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB377719FF1E33A8FC1CF98A" blockId="3.[151,1437,624,2034]" pageId="3" pageNumber="438">
|
||
<emphasis id="48CBEADECB377719FF1E33A8FE96FB14" box="[199,260,1156,1182]" italics="true" pageId="3" pageNumber="438">Head</emphasis>
|
||
. Prognathous, almost as long as wide, finely granulate, covered with oval scales and semirecumbent bristles. Anterior margin of epistoma widely rounded. Genae narrower than eyes. Frons flat, with elevated lateral sides near antennal insertions (frontal ridges) and near inner eye margins. Compound eyes lateral, protruding, hemispherical, without interfacetal setae, widely separated, with entire and straight anterior margin, finely faceted. Antennal grooves absent. Labrum with concavity on anterior margin. Terminal maxillary palpomere oval, twice as long as preceding palpomere, apex truncate; terminal labial palpomere elongate, spindle-shaped. Submentum with small round tubercle. Antennal insertions concealed under a lateral expansion of the frons and not visible in dorsal view. Antenna 11-segmented, almost moniliform, with weak club, reaching posteriorly to anterior one-third of pronotum. Club 2-segmented, setose, relatively short and stout. Scape nearly spherical. Pedicel subcylindrical, slightly longer than wide, longer than scape.Antennomeres 3–8subquadrate, as long as wide, subequal.Antennomeres 9–10 slightly transverse. Antennomere 11 rounded, as long as wide.
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB377719FF1E3126FA9EF897" blockId="3.[151,1437,624,2034]" pageId="3" pageNumber="438">
|
||
<emphasis id="48CBEADECB377719FF1E3126FE85F9AE" box="[199,279,1546,1572]" italics="true" pageId="3" pageNumber="438">Thorax</emphasis>
|
||
. Pronotum weakly elongate, about 1.3× as long as wide, with maximum width at midlength, pronotal base distinctly narrower than elytral base; pronotal disc slightly convex, with well-developed elongate median impression, consisting of three parts (small rounded anterior area, large oval central area, and triangular posterior area). Lateral pronotal sides weakly serrate, rounded, with moderately protruded and convex anterior part at middle. Lateral margins of anterior impressed part with bean-shaped callosities. Pronotum weakly bisinuate at the base, with weakly rounded middle lobe, not margined posteriorly; evenly arcuate anteriorly. Anterior angles obtuse; posterior angles widely rounded. Pronotal surface uniformly and finely granulated; each granule bearing a single curved semirecumbent bristle-like dark seta. Median impression covered with elongate-oval, recumbent, scale-like light setae.
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB377719FF1E300AFDF3F87B" blockId="3.[151,1437,624,2034]" pageId="3" pageNumber="438">Prothoracic notosternal suture well-visible, complete. Prosternum and hypomeron densely covered with oval scale-like setae, hiding the punctation. Hypomeron impressed basally, with raised posterior edge. Procoxae separated by elongate intercoxal prosternal process (as wide as procoxal diameter), distinctly widened apically and forming posterolateral projections extending to posterior margin of procoxae. Prosternal process with two fine parallel lines. Procoxal cavity posteriorly closed by projection of hypomeron and posterolateral projection of intercoxal prosternal process in the ratio 50:50 of its diameter.</paragraph>
|
||
<paragraph id="7A0036CCCB30771EFF1E37BBFC62FEE8" blockId="4.[151,1437,150,533]" pageId="4" pageNumber="439">Elytra elongate (about 2.5× as long as wide), subparallel, slightly convex, punctate-striate, with weakly carinate interstriae and straight (not rounded) apices. Elytral punctures round, arranged in 9 or 10 longitudinal rows, mostly hidden by setation. Scutellary striole not apparent. Interstriae granulate, densely covered by short setae. Setae mostly flat, oval, scale-like. Dense light oval scales making a pattern in the form of diagonal and semicircular stripes. Scutellar shield transverse, oval. Metathoracic wings are not apparent on the specimen, possibly present. Epipleura narrowed posteriorly, extending posteriorly to about the middle of abdomen.</paragraph>
|
||
<paragraph id="7A0036CCCB30771EFF1E3647FC92FD9E" blockId="4.[151,1437,150,533]" pageId="4" pageNumber="439">Mesoventrite punctate, metanepisternum granulate, metaventrite densely covered with oval scale-like setation and sparse dark hair-like setae. Mesocoxal cavities laterally closed by meso- and metaventrites. Mesanepisternum triangular; mesepimeron narrow, with one row of punctures; metanepisternum narrow and long; with one row of small granules. Mesocoxae narrowly separated by 1/4 mesocoxal diameter. Metaventrite with convex disc; discrimen, distinct, as long as one-half of metaventrite.</paragraph>
|
||
<caption id="2EC06644CB30771EFF4E30A3FDE0F840" pageId="4" pageNumber="439" startId="4.[151,264,1935,1959]" targetBox="[151,1436,554,1909]" targetPageId="4" targetType="figure">
|
||
<paragraph id="7A0036CCCB30771EFF4E30A3FDE0F840" blockId="4.[151,1436,1935,1995]" pageId="4" pageNumber="439">
|
||
<emphasis id="48CBEADECB30771EFF4E30A3FEACF82D" bold="true" box="[151,318,1935,1959]" pageId="4" pageNumber="439">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="BDBF4D4FCB30771EFE9130A3FD02F82D" authority="Alekseev & Nabozhenko, 2023" authorityName="Alekseev & Nabozhenko" authorityYear="2023" box="[328,656,1935,1959]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="439" phylum="Arthropoda" rank="species" species="auriborussiensis" status="sp. nov.">
|
||
<emphasis id="48CBEADECB30771EFE9130A3FD02F82D" box="[328,656,1935,1959]" italics="true" pageId="4" pageNumber="439">Zopheromimus auriborussiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="48CBEADECB30771EFD4330A3FD7FF82D" bold="true" box="[666,749,1935,1959]" pageId="4" pageNumber="439">
|
||
<taxonomicNameLabel id="53F857A5CB30771EFD4330A3FD7FF82D" box="[666,749,1935,1959]" pageId="4" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, No KAM 8325/2: 1—habitus, dorso-lateral view; 2— habitus, ventro-lateral view. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2EC06644CB31771FFF4E304FFEF6F86D" pageId="5" pageNumber="440" startId="5.[151,264,1891,1915]" targetBox="[177,1411,181,1855]" targetPageId="5" targetType="figure">
|
||
<paragraph id="7A0036CCCB31771FFF4E304FFEF6F86D" blockId="5.[151,1437,1891,2023]" pageId="5" pageNumber="440">
|
||
<emphasis id="48CBEADECB31771FFF4E304FFEADF8F1" bold="true" box="[151,319,1891,1915]" pageId="5" pageNumber="440">FIGURES 3–4.</emphasis>
|
||
<taxonomicName id="BDBF4D4FCB31771FFE93304FFD01F8F1" authority="Alekseev & Nabozhenko, 2023" authorityName="Alekseev & Nabozhenko" authorityYear="2023" box="[330,659,1891,1915]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="auriborussiensis" status="sp. nov.">
|
||
<emphasis id="48CBEADECB31771FFE93304FFD01F8F1" box="[330,659,1891,1915]" italics="true" pageId="5" pageNumber="440">Zopheromimus auriborussiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="48CBEADECB31771FFD47304FFD61F8F1" bold="true" box="[670,755,1891,1915]" pageId="5" pageNumber="440">
|
||
<taxonomicNameLabel id="53F857A5CB31771FFD47304FFD61F8F1" box="[670,755,1891,1915]" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, No KAM 8325/2, forebody, ventro-lateral view: 3— microphotograph; 4—schematic drawing. Abbreviations: A2—antennomere 2 (as long as antennomere 3); E—eye (lateral, hemispherical); PH—prothoracic hypomeron (not grooved); PC—procoxal cavity (externally closed); AP—abdominal procoxal process (triangular).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2EC06644CB32771CFF4E30BBFBA9F87D" pageId="6" pageNumber="441" startId="6.[151,264,1943,1967]" targetBox="[158,1430,181,1919]" targetPageId="6" targetType="figure">
|
||
<paragraph id="7A0036CCCB32771CFF4E30BBFBA9F87D" blockId="6.[151,1436,1943,2040]" pageId="6" pageNumber="441">
|
||
<emphasis id="48CBEADECB32771CFF4E30BBFED3F83A" bold="true" box="[151,321,1943,1968]" pageId="6" pageNumber="441">FIGURES 5–7.</emphasis>
|
||
Representatives of the tribe
|
||
<taxonomicName id="BDBF4D4FCB32771CFD5A30BBFCB0F825" authority="Alekseev & Nabozhenko, 2023" authorityName="Alekseev & Nabozhenko" authorityYear="2023" box="[643,802,1943,1967]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="441" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Zopheromimini">Zopheromimini</taxonomicName>
|
||
<taxonomicNameLabel id="53F857A5CB32771CFCF730BBFC05F825" box="[814,919,1943,1968]" pageId="6" pageNumber="441" rank="tribe">
|
||
<emphasis id="48CBEADECB32771CFCF730BBFC03F83A" bold="true" box="[814,913,1943,1968]" pageId="6" pageNumber="441">trib. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(microphotographs): 5—head of
|
||
<taxonomicName id="BDBF4D4FCB32771CFADC30BBFEADF859" authorityName="Alekseev & Nabozhenko" authorityYear="2023" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="441" phylum="Arthropoda" rank="species" species="auriborussiensis">
|
||
<emphasis id="48CBEADECB32771CFADC30BBFEADF859" italics="true" pageId="6" pageNumber="441">Zopheromimus auriborussiensis</emphasis>
|
||
</taxonomicName>
|
||
; 6—abdomen of
|
||
<taxonomicName id="BDBF4D4FCB32771CFDDD3090FD5CF859" authorityName="Alekseev & Nabozhenko" authorityYear="2023" box="[516,718,1979,2003]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="441" phylum="Arthropoda" rank="species" species="auriborussiensis">
|
||
<emphasis id="48CBEADECB32771CFDDD3090FD5CF859" box="[516,718,1979,2003]" italics="true" pageId="6" pageNumber="441">Z. auriborussiensis</emphasis>
|
||
</taxonomicName>
|
||
; 7—abdomen of
|
||
<taxonomicName id="BDBF4D4FCB32771CFC4A3090FB09F859" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[915,1179,1979,2003]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="441" phylum="Arthropoda" rank="species" species="colydioides">
|
||
<emphasis id="48CBEADECB32771CFC4A3090FB09F859" box="[915,1179,1979,2003]" italics="true" pageId="6" pageNumber="441">Yantaroxenos colydioides</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: mem— membranes between non-connate ventrites; pag—preapical groove on abdominal ventrite 5.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7A0036CCCB33771DFF1E37BBFB1DFECB" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="442">
|
||
<emphasis id="48CBEADECB33771DFF1E37BBFF6EFF3A" box="[199,252,151,176]" italics="true" pageId="7" pageNumber="442">Legs</emphasis>
|
||
. Relatively similar in shape, setose; femora and tibiae covered with dense scale-like setae. Pro- and mesocoxae hemispherical and convex, metacoxae strongly transverse. Trochanters oblique, visible on all legs. Femora slightly longer than tibiae, flattened. Tibiae straight. Tarsi about as long as tibiae. All tarsomeres narrow, simple (non-lobed). Tarsal formula 5-5-4. Relative length ratios of metatarsomeres 1–4 equal to 9:5:7:18. Tarsal claws slender, simple, without basal tooth or expansion, about 0.3× as long as tarsomere 4.
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB33771DFF1E3667FD30FE7F" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="442">
|
||
<emphasis id="48CBEADECB33771DFF1E3667FEA3FEEF" box="[199,305,331,357]" italics="true" pageId="7" pageNumber="442">Abdomen</emphasis>
|
||
. With five visible, freely articulated abdominal ventrites; sutures between ventrites deep and complete. Ventrites are punctured and densely covered with oval scale-like setae and sparse dark hair-like setae. Intercoxal process of abdominal ventrite 1 triangular, acute. Relative length ratios of abdominal ventrites 1–5 equal to 48:30:25:20:22 (measured medially). Abdominal ventrite 5 with two shallow, transverse preapical grooves. Apex of ventrite 5 with dense semierect pubescence.
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB33771DFF1E36D3FB13FD47" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="442">
|
||
<emphasis id="48CBEADECB33771DFF1E36D3FE9AFD93" bold="true" box="[199,264,511,537]" pageId="7" pageNumber="442">Note:</emphasis>
|
||
The number of antennomeres in the club was determined by using patches of dense and fine pubescence (interpreted as sensory areas) as indicators of the distal surface of a club antennomere. The apical antennomeres are not fused and all antennomeres look rather uniformly articulated, therefore the form of antennae is described as “almost moniliform.” However antennomeres 9–10 are weakly expanded in comparison to more proximal antennomeres. Only antennomeres 10 and 11 bear a finely pubescent apical surface (discernible in frontal view). Therefore, it was decided that the specimen possesses a weak 2-segmented antennal club.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="32A56547CB33771DFF1E35FBFB28FD7B" box="[199,1210,727,753]" pageId="7" pageNumber="442" type="discussion">
|
||
<paragraph id="7A0036CCCB33771DFF1E35FBFB28FD7B" blockId="7.[151,1437,151,1077]" box="[199,1210,727,753]" pageId="7" pageNumber="442">
|
||
<emphasis id="48CBEADECB33771DFF1E35FBFEA0FD7B" bold="true" box="[199,306,727,753]" pageId="7" pageNumber="442">Remark:</emphasis>
|
||
The genitalia are not exposed, making definite sexual determination impossible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="32A56547CB33771DFF1E35D6FA0EFBBF" pageId="7" pageNumber="442" type="diagnosis">
|
||
<paragraph id="7A0036CCCB33771DFF1E35D6FD52FC0B" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="442">
|
||
<emphasis id="48CBEADECB33771DFF1E35D6FE42FC9E" bold="true" box="[199,464,762,788]" pageId="7" pageNumber="442">Differential diagnosis.</emphasis>
|
||
The new extinct species from Baltic amber can be easily distinguished from extinct
|
||
<taxonomicName id="BDBF4D4FCB33771DFF4E3433FE85FCB3" authorityName="Solier" authorityYear="1834" box="[151,279,799,825]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="family">Zopheridae</taxonomicName>
|
||
from Paleogene ambers by the combination of subfamilial and tribal characters, but most obviously in 5-5-4 tarsal formula (in contrast to the 4-4-4 tarsal formula in all
|
||
<taxonomicName id="BDBF4D4FCB33771DFCBE346FFC70FCD7" authorityName="Erichson" authorityYear="1842" box="[871,994,835,861]" class="Insecta" family="Colydiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="family">Colydiinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="BDBF4D4FCB33771DFBCE346FFB33FCD6" box="[1047,1185,835,860]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="48CBEADECB33771DFBCE346FFB33FCD6" box="[1047,1185,835,860]" italics="true" pageId="7" pageNumber="442">Pycnomerus</emphasis>
|
||
</taxonomicName>
|
||
, or 4-5-
|
||
<quantity id="BD479B29CB33771DFB20346EFAB7FCD7" box="[1273,1317,834,861]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="442" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="BDBF4D4FCB33771DFAF2346FFEC9FC0B" authority="Alekseev" authorityName="Alekseev" class="Insecta" family="Colydiinae" genus="Damzenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="groehni">
|
||
<emphasis id="48CBEADECB33771DFAF2346FFF7FFC0B" italics="true" pageId="7" pageNumber="442">Damzenia groehni</emphasis>
|
||
Alekseev
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="1E2E4B3DCB33771DFEA4344BFD26FC0B" author="Alekseev, V. I. & Alekseev, P. I." box="[381,692,871,897]" pageId="7" pageNumber="442" pagination="1 - 10" refId="ref4951" refString="Alekseev, V. I. & Alekseev, P. I. (2019) New fossil cylindrical bark beetle (Zopheridae: Colydiinae: Gempylodini) from Eocene Baltic amber: an abnormal or intermediate form within Tenebrionoidea. Palaeontologia Electronica, 22.2.43 A, 1 - 10. https: // doi. org / 10.26879 / 963" type="book chapter" year="2019">Alekseev & Alekseev 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7A0036CCCB33771DFF1E34A7FA0EFBBF" blockId="7.[151,1437,151,1077]" pageId="7" pageNumber="442">
|
||
In many respects, this new genus resembles the Baltic amber species
|
||
<taxonomicName id="BDBF4D4FCB33771DFC3834A7FE2BFC43" authority="Nabozhenko, Kirejtshuk et Merkl, 2016" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="colydioides">
|
||
<emphasis id="48CBEADECB33771DFC3834A7FA92FC2F" box="[993,1280,907,933]" italics="true" pageId="7" pageNumber="442">Yantaroxenos colydioides</emphasis>
|
||
<bibRefCitation id="1E2E4B3DCB33771DFAD134A7FE2BFC43" author="Nabozhenko, M. & Kirejtshuk, A. & Merkl, O." pageId="7" pageNumber="442" pagination="563 - 566" refId="ref6010" refString="Nabozhenko, M., Kirejtshuk, A. & Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">Nabozhenko, Kirejtshuk et Merkl, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
, originally assigned to the tenebrionid tribe
|
||
<taxonomicName id="BDBF4D4FCB33771DFC443483FAB2FC43" authority="(Nabozhenko et al. 2016)" baseAuthorityName="Nabozhenko" baseAuthorityYear="2016" box="[925,1312,943,969]" class="Insecta" family="Lagriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="tribe" tribe="Belopini">
|
||
Belopini (
|
||
<bibRefCitation id="1E2E4B3DCB33771DFBD23483FA8AFC43" author="Nabozhenko, M. & Kirejtshuk, A. & Merkl, O." box="[1035,1304,943,969]" pageId="7" pageNumber="442" pagination="563 - 566" refId="ref6010" refString="Nabozhenko, M., Kirejtshuk, A. & Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">
|
||
Nabozhenko
|
||
<emphasis id="48CBEADECB33771DFB793483FB45FC43" box="[1184,1239,943,969]" italics="true" pageId="7" pageNumber="442">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but differs in having protruding, hemispherical eyes (eyes longitudinally oval in
|
||
<taxonomicName id="BDBF4D4FCB33771DFC1B34FFFBCAFC66" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[962,1112,979,1004]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="48CBEADECB33771DFC1B34FFFBCAFC66" box="[962,1112,979,1004]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
|
||
</taxonomicName>
|
||
), a different pronotal shape (strongly convex with protruding anterior half that covers the base of the head in
|
||
<taxonomicName id="BDBF4D4FCB33771DFBC134DBFB3CFB9A" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[1048,1198,1015,1040]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="48CBEADECB33771DFBC134DBFB3CFB9A" box="[1048,1198,1015,1040]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
|
||
</taxonomicName>
|
||
), shorter body length (
|
||
<quantity id="BD479B29CB33771DFF463337FF64FBBF" box="[159,246,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="7" pageNumber="442" unit="mm" value="8.2">8.2 mm</quantity>
|
||
in
|
||
<taxonomicName id="BDBF4D4FCB33771DFEC13337FE3CFBBE" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[280,430,1051,1076]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="48CBEADECB33771DFEC13337FE3CFBBE" box="[280,430,1051,1076]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
|
||
</taxonomicName>
|
||
), and 11-segmented antennae (antennae “with 10 visible antennomeres” in
|
||
<taxonomicName id="BDBF4D4FCB33771DFB2F3337FA1EFBBE" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[1270,1420,1051,1076]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="48CBEADECB33771DFB2F3337FA1EFBBE" box="[1270,1420,1051,1076]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |