172 lines
21 KiB
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172 lines
21 KiB
XML
<document id="DE70AB8289AD992E5B8F1AAEBBBA53DF" ID-DOI="10.11646/zootaxa.4039.4.1" ID-GBIF-Dataset="ca91ab86-49b9-4d90-ac6f-2051ca9d2f85" ID-ISSN="1175-5326" ID-Zenodo-Dep="232034" ID-ZooBank="4B12AA52-6A3F-44B9-BBEC-EBABF0E22DF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461332656152" checkinUser="plazi" docAuthor="Blake, James A. & Göransson, Peter" docDate="2015" docId="EE2B2061FFC60276FF2F0ADCFC4AF8E8" docLanguage="en" docName="zt04039p515.pdf" docOrigin="Zootaxa 4039 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tharyx Webster & Benedict 1887" docType="treatment" docVersion="8" lastPageNumber="502" masterDocId="12125819FFC70277FFB8095EFFE8FF8E" masterDocTitle="Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)" masterLastPageNumber="515" masterPageNumber="501" pageNumber="502" updateTime="1698618770507" updateUser="plazi">
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<mods:title id="F6BFB0743B27A6E3149FB4D2A4F67933">Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)</mods:title>
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<mods:namePart id="8C079D19E97EF5F66C17790DA84E126B">Blake, James A.</mods:namePart>
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<mods:namePart id="38AB7C1078B777E2F6B95DF9BD08E3B3">Göransson, Peter</mods:namePart>
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<mods:typeOfResource id="A6A2CA15CBE521B923DB2D2C4D99FB55">text</mods:typeOfResource>
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<mods:title id="6BDB163F25DF19040667CA7FF0ACDF91">Zootaxa</mods:title>
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<mods:date id="7B5C16FA3AFFB84C1F3A29A33133ED6A">2015</mods:date>
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<mods:number id="73F42DD1D1EF21AEF90DCD5614BC7B55">4039</mods:number>
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<mods:number id="47EB243654436E5A0A764ECF9E73F604">4</mods:number>
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<mods:identifier id="C49267140973F438FE378F0F68CC141D" type="DOI">10.11646/zootaxa.4039.4.1</mods:identifier>
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<treatment id="EE2B2061FFC60276FF2F0ADCFC4AF8E8" ID-DOI="http://doi.org/10.5281/zenodo.5658075" ID-GBIF-Taxon="119644305" ID-Zenodo-Dep="5658075" LSID="urn:lsid:plazi:treatment:EE2B2061FFC60276FF2F0ADCFC4AF8E8" httpUri="http://treatment.plazi.org/id/EE2B2061FFC60276FF2F0ADCFC4AF8E8" lastPageNumber="502" pageId="1" pageNumber="502">
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<subSubSection id="2E98C2FCFFC60276FF2F0ADCFD7BFC12" box="[151,659,898,924]" pageId="1" pageNumber="502" type="nomenclature">
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<paragraph id="663D9177FFC60276FF2F0ADCFD7BFC12" blockId="1.[151,659,898,924]" box="[151,659,898,924]" pageId="1" pageNumber="502">
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<heading id="3D75261BFFC60276FF2F0ADCFD7BFC12" bold="true" box="[151,659,898,924]" fontSize="11" level="1" pageId="1" pageNumber="502" reason="1">
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<emphasis id="54F64D65FFC60276FF2F0ADCFD7BFC12" bold="true" box="[151,659,898,924]" pageId="1" pageNumber="502">
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Genus
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<taxonomicName id="A182EAF4FFC60276FF560ADCFD7BFC12" ID-CoL="84V7M" authority="Webster & Benedict, 1887" authorityName="Webster & Benedict" authorityYear="1887" box="[238,659,898,924]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FF560ADCFEAEFC12" bold="true" box="[238,326,898,924]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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<bibRefCitation id="0213EC86FFC60276FEF50ADCFD7BFC12" author="Webster" box="[333,659,898,924]" pageId="1" pageNumber="502" refString="Webster, H. E & Benedict, J. E. (1887) The Annelida Chaetopoda from Eastport, Maine. Reports of the U. S. Fish Commission for 1885, 707 - 755, pls. 1 - 8." type="journal article" year="1887">Webster & Benedict, 1887</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="2E98C2FCFFC60276FF2F0A97FA7BFC6C" box="[151,1427,969,994]" pageId="1" pageNumber="502" type="reference_group">
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<paragraph id="663D9177FFC60276FF2F0A97FA7BFC6C" blockId="1.[151,1437,969,1894]" box="[151,1427,969,994]" pageId="1" pageNumber="502">
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<emphasis id="54F64D65FFC60276FF2F0A97FEC5FC6C" bold="true" box="[151,301,969,994]" pageId="1" pageNumber="502">Type-species</emphasis>
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<taxonomicName id="A182EAF4FFC60276FE8C0A97FCEEFC6C" ID-CoL="564LP" authority="Webster & Benedict, 1887" authorityName="Webster & Benedict" authorityYear="1887" box="[308,774,969,994]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="species" species="acutus">
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<emphasis id="54F64D65FFC60276FE8C0A97FE3CFC6F" box="[308,468,969,994]" italics="true" pageId="1" pageNumber="502">Tharyx acutus</emphasis>
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<bibRefCitation id="0213EC86FFC60276FE630A97FCEEFC6C" author="Webster" box="[475,774,969,994]" pageId="1" pageNumber="502" refString="Webster, H. E & Benedict, J. E. (1887) The Annelida Chaetopoda from Eastport, Maine. Reports of the U. S. Fish Commission for 1885, 707 - 755, pls. 1 - 8." type="journal article" year="1887">Webster & Benedict, 1887</bibRefCitation>
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</taxonomicName>
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, designated by
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<bibRefCitation id="0213EC86FFC60276FC0B0A94FB81FC6C" author="Hartman" box="[947,1129,969,994]" pageId="1" pageNumber="502" refString="Hartman, O. (1959) Catalogue of the Polychaetous Annelids of the World. Allan Hancock Foundation Occasional Paper, 23, 1 - 628." type="journal article" year="1959">Hartman (1959)</bibRefCitation>
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. Revised by
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<bibRefCitation id="0213EC86FFC60276FB410A97FA67FC6C" author="Blake" box="[1273,1423,969,994]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. & Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="2E98C2FCFFC60276FF7F0AB3FF24FA8C" pageId="1" pageNumber="502" type="diagnosis">
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<paragraph id="663D9177FFC60276FF7F0AB3FF24FA8C" blockId="1.[151,1437,969,1894]" pageId="1" pageNumber="502">
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<emphasis id="54F64D65FFC60276FF7F0AB3FEA9FB88" bold="true" box="[199,321,1005,1030]" pageId="1" pageNumber="502">Diagnosis.</emphasis>
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Prostomium conical; peristomium elongate, with pair of grooved dorsal tentacles arising on posterior margin anterior to setiger 1; first pair of branchiae typically arising immediately posterior to dorsal tentacles either on posterior margin of peristomium or rarely on anterior margin of setiger 1; abdominal segments sometimes beadlike. Noto- and neuropodia located close to one another, not widely separated. Setae including simple capillaries in anterior and middle setigers, acicular spines in posterior setigers with irregular notched tips, sometimes appearing weakly bidentate, with pair of stunted or rounded knobs but never with distinct, sharply pointed teeth; spines present either in both posterior noto- and neuropodia or only in neuropodia. Pygidium with terminal anus and small ventral lobe or disk.
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</paragraph>
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</subSubSection>
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<subSubSection id="2E98C2FCFFC60276FF7F0C53FC4AF8E8" pageId="1" pageNumber="502" type="discussion">
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<paragraph id="663D9177FFC60276FF7F0C53FD9AF9E4" blockId="1.[151,1437,969,1894]" pageId="1" pageNumber="502">
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<emphasis id="54F64D65FFC60276FF7F0C53FEDCFAA8" bold="true" box="[199,308,1293,1318]" pageId="1" pageNumber="502">Remarks</emphasis>
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.
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<bibRefCitation id="0213EC86FFC60276FEFB0C53FE3EFAA8" author="Blake" box="[323,470,1293,1318]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. & Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
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determined that the
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<typeStatus id="B9392FD5FFC60276FD790C50FD19FAA8" box="[705,753,1294,1318]" pageId="1" pageNumber="502">type</typeStatus>
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species of the genus
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<emphasis id="54F64D65FFC60276FC5C0C53FB40FAAB" box="[996,1192,1293,1318]" italics="true" pageId="1" pageNumber="502">
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<taxonomicName id="A182EAF4FFC60276FC5C0C53FBDEFAA8" box="[996,1078,1293,1318]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">Tharyx</taxonomicName>
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,
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<taxonomicName id="A182EAF4FFC60276FBFB0C50FB40FAAB" authority="Webster & Benedict, 1887" authorityName="Webster & Benedict" authorityYear="1887" box="[1091,1192,1294,1317]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="species" species="acutus">T. acutus</taxonomicName>
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</emphasis>
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<bibRefCitation id="0213EC86FFC60276FB080C53FF3AFAC4" author="Webster" pageId="1" pageNumber="502" refString="Webster, H. E & Benedict, J. E. (1887) The Annelida Chaetopoda from Eastport, Maine. Reports of the U. S. Fish Commission for 1885, 707 - 755, pls. 1 - 8." type="journal article" year="1887">Webster & Benedict, 1887</bibRefCitation>
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, had knob-tipped hooks or spines in addition to capillaries. This discovery differed from the traditional concept of
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<taxonomicName id="A182EAF4FFC60276FEA10C0BFE80FAE0" box="[281,360,1365,1390]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FEA10C0BFE80FAE0" box="[281,360,1365,1390]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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, which had been based upon specimens having all capillaries (
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<bibRefCitation id="0213EC86FFC60276FBFE0C08FB18FAE0" author="Hartman" box="[1094,1264,1365,1390]" pageId="1" pageNumber="502" refString="Hartman, O. (1961) Polychaetous annelids from California. Allan Hancock Pacific Expeditions, 22, 1 - 226." type="journal article" year="1961">Hartman 1961</bibRefCitation>
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).
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<bibRefCitation id="0213EC86FFC60276FABE0C0BFA74FAE0" author="Blake" box="[1286,1436,1365,1390]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. & Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
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restricted species of
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<taxonomicName id="A182EAF4FFC60276FE3E0C27FE3EFA1C" box="[390,470,1401,1426]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FE3E0C27FE3EFA1C" box="[390,470,1401,1426]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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to those having blunt or knob-tipped spines and moved species having simple and denticulated capillaries to the genera
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<taxonomicName id="A182EAF4FFC60276FD800CC3FD3BFA38" box="[568,723,1437,1462]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FD800CC3FD3BFA38" box="[568,723,1437,1462]" italics="true" pageId="1" pageNumber="502">Aphelochaeta</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="A182EAF4FFC60276FCB00CC3FC7FFA38" box="[776,919,1437,1462]" class="Polychaeta" family="Cirratulidae" genus="Monticellina" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FCB00CC3FC7FFA38" box="[776,919,1437,1462]" italics="true" pageId="1" pageNumber="502">Monticellina</emphasis>
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</taxonomicName>
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, respectively. The genus
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<taxonomicName id="A182EAF4FFC60276FB0B0CC3FAEBFA38" box="[1203,1283,1437,1462]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FB0B0CC3FAEBFA38" box="[1203,1283,1437,1462]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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appears most closely related to
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<taxonomicName id="A182EAF4FFC60276FEE40C9FFE0BFA54" box="[348,483,1473,1498]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FEE40C9FFE0BFA54" box="[348,483,1473,1498]" italics="true" pageId="1" pageNumber="502">Caulleriella</emphasis>
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</taxonomicName>
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in that both genera have species with modified spines that are more or less bidentate. In addition to the nature of the posterior spines,
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<taxonomicName id="A182EAF4FFC60276FD700CBBFCA7FA70" box="[712,847,1509,1534]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FD700CBBFCA7FA70" box="[712,847,1509,1534]" italics="true" pageId="1" pageNumber="502">Caulleriella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="A182EAF4FFC60276FC330CBBFC32FA70" box="[907,986,1509,1534]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FC330CBBFC32FA70" box="[907,986,1509,1534]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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species are easily distinguished by the position of the noto- and neuropodia. In
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<taxonomicName id="A182EAF4FFC60276FDD90F57FD00F9AC" box="[609,744,1545,1570]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FDD90F57FD00F9AC" box="[609,744,1545,1570]" italics="true" pageId="1" pageNumber="502">Caulleriella</emphasis>
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</taxonomicName>
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, the setal fascicles are widely separated from one another, so much so, that in cross section of some species, they appear at four corners of the body. In
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<taxonomicName id="A182EAF4FFC60276FB090F73FAE8F9C8" box="[1201,1280,1581,1606]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FB090F73FAE8F9C8" box="[1201,1280,1581,1606]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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, on the other hand, the setal fascicles are close together.
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</paragraph>
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<paragraph id="663D9177FFC60276FF7F0F2BFC4AF8E8" blockId="1.[151,1437,969,1894]" pageId="1" pageNumber="502">
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<bibRefCitation id="0213EC86FFC60276FF7F0F2BFEB3F900" author="Blake" box="[199,347,1653,1678]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. & Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
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originally included five species in the genus
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<taxonomicName id="A182EAF4FFC60276FCE60F2BFC45F900" box="[862,941,1653,1678]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FCE60F2BFC45F900" box="[862,941,1653,1678]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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, one of which,
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<taxonomicName id="A182EAF4FFC60276FBE70F28FB10F900" box="[1119,1272,1653,1678]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="species" species="killariensis">
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<emphasis id="54F64D65FFC60276FBE70F28FB10F900" box="[1119,1272,1653,1678]" italics="true" pageId="1" pageNumber="502">T. killariensis</emphasis>
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</taxonomicName>
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is redescribed in the present paper. Four additional species were added by
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<bibRefCitation id="0213EC86FFC60276FCF30FC7FC32F93C" author="Blake" box="[843,986,1689,1714]" pageId="1" pageNumber="502" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. & Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">Blake (1996</bibRefCitation>
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,
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<bibRefCitation id="0213EC86FFC60276FC5F0FC7FBC9F93C" author="Blake" box="[999,1057,1689,1714]" pageId="1" pageNumber="502" refString="Blake, J. A. (2015) New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitzbergen in the Norwegian Arctic. Zootaxa, 3919 (3), 501 - 552. http: // dx. doi. org / 10.11646 / zootaxa. 3919.3.5" type="journal article" year="2015">2015</bibRefCitation>
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) and
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<bibRefCitation id="0213EC86FFC60276FBDC0FC7FF0CF958" author="Magalhaes" pageId="1" pageNumber="502" refString="Magalhaes, W. F. & Bailey-Brock, J. H. (2013) Bitentaculate Cirratulidae (Annelida: Polychaeta) from the northwestern Pacific Islands with a description of nine new species. Zootaxa, 3630 (1), 80 - 116. http: // dx. doi. org / 10.11646 / zootaxa. 3630.1.3." type="journal article" year="2013">Magalhães & Bailey-Brock (2013)</bibRefCitation>
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. Two new species discovered in shallow-water sediments in
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<collectingCountry id="1E95D1E7FFC60276FC210FE3FC1AF958" box="[921,1010,1725,1750]" name="Sweden" pageId="1" pageNumber="502">Sweden</collectingCountry>
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are here described, bringing the total number of species in the genus to eleven. In addition, another eight species of
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<taxonomicName id="A182EAF4FFC60276FBBE0FBFFBBEF974" box="[1030,1110,1761,1786]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FBBE0FBFFBBEF974" box="[1030,1110,1761,1786]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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are known to the first author from North
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<collectingCountry id="1E95D1E7FFC60276FEA50E5BFE95F890" box="[285,381,1797,1822]" name="United States of America" pageId="1" pageNumber="502">America</collectingCountry>
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and
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<collectingCountry id="1E95D1E7FFC60276FE0D0E5BFDC5F890" box="[437,557,1797,1822]" name="Antarctica" pageId="1" pageNumber="502">Antarctica</collectingCountry>
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. With the number of species thus nearly tripled over the past 24 years, there is now a larger suite of characters available to define the genus than was available in the past. The morphological characters that best define the known species of
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<taxonomicName id="A182EAF4FFC60276FD0C0E13FCECF8E8" box="[692,772,1869,1894]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
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<emphasis id="54F64D65FFC60276FD0C0E13FCECF8E8" box="[692,772,1869,1894]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
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</taxonomicName>
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are reviewed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |