treatments-xml/data/ED/6F/FF/ED6FFF21D020A402FF66FB5DB452AA61.xml

<document id="D938FF190E2584C967677D50D51719BA" ID-DOI="10.5281/zenodo.12766004" ID-ISSN="1989-6581" ID-Zenodo-Dep="12766004" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721246549113" checkinUser="felipe" docAuthor="Fa ̇ ndez, Eduardo I. &amp; Rider, David A." docDate="2016" docId="ED6FFF21D020A402FF66FB5DB452AA61" docLanguage="es" docName="ArqEntomol.15.383-389.pdf" docOrigin="Arquivos Entomolóxicos 15" docStyle="DocumentStyle:DF23C849EA72F80F6DF49FA5382909D5.2:ArqEntomol.2008-.journal_article.1column" docStyleId="DF23C849EA72F80F6DF49FA5382909D5" docStyleName="ArqEntomol.2008-.journal_article.1column" docStyleVersion="2" docTitle="Acledra (Acledra) albocostata subsp. lueri ndez &amp; Rider, 2016, n. ssp." docType="treatment" docVersion="2" lastPageNumber="386" masterDocId="11568759D022A401FFD0FFF9B010AE58" masterDocTitle="Notas sobre el complejo de Acledra (Acledra) albocostata (Spinola, 1852) (Hemiptera: Heteroptera: Pentatomidae: Pentatominae)." masterLastPageNumber="389" masterPageNumber="383" pageNumber="385" updateTime="1721253630782" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3359A0F1F9B1047344DCE59B4B014DFE">Notas sobre el complejo de Acledra (Acledra) albocostata (Spinola, 1852) (Hemiptera: Heteroptera: Pentatomidae: Pentatominae).</mods:title>
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<mods:namePart id="4A27A7A234D4FAEF2559D195464BCB5C">Fa ̇ ndez, Eduardo I.</mods:namePart>
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<mods:namePart id="8828402D5D2D51C5F821729ACEEA60E4">Rider, David A.</mods:namePart>
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<mods:date id="27FB34C39C3A4BDC253F92DF65AB24C3">2016</mods:date>
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<paragraph id="65794E37D020A403FF66FB5DB3EFAAE5" blockId="2.[182,1023,1188,1213]" box="[182,1023,1188,1213]" pageId="2" pageNumber="385">
<heading id="3E31F95BD020A403FF66FB5DB3EFAAE5" bold="true" box="[182,1023,1188,1213]" fontSize="10" level="2" pageId="2" pageNumber="385" reason="6">
<emphasis id="57B29225D020A403FF66FB5DB3EFAAE5" bold="true" box="[182,1023,1188,1213]" pageId="2" pageNumber="385">
<taxonomicName id="A2C635B4D020A403FF66FB5DB26CAAE5" authorityName="Fȧndez &amp; Rider" authorityYear="2016" box="[182,636,1188,1213]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="subSpecies" species="albocostata" status="subsp. nov." subGenus="Acledra" subSpecies="lueri">
<emphasis id="57B29225D020A403FF66FB5DB26CAAE5" bold="true" box="[182,636,1188,1213]" italics="true" pageId="2" pageNumber="385">Acledra (Acledra) albocostata lueri</emphasis>
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<taxonomicNameLabel id="4C812F5ED020A403FD55FB5DB2C1AAE5" box="[645,721,1188,1213]" pageId="2" pageNumber="385" rank="subSpecies">n. ssp.</taxonomicNameLabel>
(
<figureCitation id="FDFD52B2D020A403FD30FB5DB365AAE5" box="[736,885,1188,1213]" captionStart="Figs" captionStartId="5.[168,215,1887,1907]" captionTargetBox="[148,1445,612,1846]" captionTargetId="figure-114@5.[148,1445,610,1847]" captionTargetPageId="5" captionText="Figs. 1-9.- Acledra (Acledra) albocostata. 1.- A. (A.) albocostata albocostata, habitus. 2.- A. (A.) albocostata carinata, habitus. 3.- A. (A.) albocostata lueri, habitus. 4.- A. (A.) albocostata albocostata, pigóforo en vista dorsal. 5.- A. (A.) albocostata carinata, pigóforo en vista dorsal. 6.- A. (A.) albocostata lueri, pigóforo en vista dorsal. 7.- A. (A.) albocostata albocostata, pigóforo en vista ventral. 8.- A. (A.) albocostata carinata, pigóforo en vista ventral. 9.- A. (A.) albocostata lueri, pigóforo en vista ventral." figureDoi="http://doi.org/10.5281/zenodo.12766006" httpUri="https://zenodo.org/record/12766006/files/figure.png" pageId="2" pageNumber="385">Figs. 3, 6, 9</figureCitation>
, 
<figureCitation id="FDFD52B2D020A403FC52FB5DB3E7AAE5" box="[898,1015,1188,1213]" captionStart="Figs" captionStartId="6.[198,245,1751,1771]" captionTargetBox="[198,1398,219,1701]" captionTargetId="figure-18@6.[198,1408,219,1701]" captionTargetPageId="6" captionText="Figs. 10-14.- Acledra (Acledra) albocostata. 10.- A. (A.) albocostata albocostata, pronoto. 11.- A. (A.) albocostata carinata, pronoto. 12.- A. (A.) albocostata lueri, pronoto. 13.- A. (A.) albocostata lueri, terminalia femenina. 14.- Distribucion de A. (A) albocostata: área azul, A. (A.) albocostata albocostata; círculos rojos, A. (A.) albocostata carinata; cuadrados verdes, A. (A.) albocostata lueri." figureDoi="http://doi.org/10.5281/zenodo.12766008" httpUri="https://zenodo.org/record/12766008/files/figure.png" pageId="2" pageNumber="385">12, 13, 14</figureCitation>
)
</emphasis>
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</paragraph>
<paragraph id="65794E37D020A403FF3EFB21B4C8AAAA" blockId="2.[182,1444,1240,2047]" box="[238,1240,1240,1267]" pageId="2" pageNumber="385">
<emphasis id="57B29225D020A403FF3EFB21B4C8AAAA" bold="true" box="[238,1240,1240,1267]" pageId="2" pageNumber="385">
Descripción: Cuerpo alargado, ocreáceo con puntuación densa y concolora (
<figureCitation id="FDFD52B2D020A403FB52FB20B4DAAAAA" box="[1154,1226,1241,1266]" captionStart="Figs" captionStartId="5.[168,215,1887,1907]" captionTargetBox="[148,1445,612,1846]" captionTargetId="figure-114@5.[148,1445,610,1847]" captionTargetPageId="5" captionText="Figs. 1-9.- Acledra (Acledra) albocostata. 1.- A. (A.) albocostata albocostata, habitus. 2.- A. (A.) albocostata carinata, habitus. 3.- A. (A.) albocostata lueri, habitus. 4.- A. (A.) albocostata albocostata, pigóforo en vista dorsal. 5.- A. (A.) albocostata carinata, pigóforo en vista dorsal. 6.- A. (A.) albocostata lueri, pigóforo en vista dorsal. 7.- A. (A.) albocostata albocostata, pigóforo en vista ventral. 8.- A. (A.) albocostata carinata, pigóforo en vista ventral. 9.- A. (A.) albocostata lueri, pigóforo en vista ventral." figureDoi="http://doi.org/10.5281/zenodo.12766006" httpUri="https://zenodo.org/record/12766006/files/figure.png" pageId="2" pageNumber="385">Fig. 3</figureCitation>
).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2DDC1DBCD020A402FF3EFB07B483AC69" lastPageId="3" lastPageNumber="386" pageId="2" pageNumber="385" type="description">
<paragraph id="65794E37D020A403FF3EFB07B476ABDE" blockId="2.[182,1444,1240,2047]" pageId="2" pageNumber="385">
<emphasis id="57B29225D020A403FF3EFB07B476ABDE" bold="true" pageId="2" pageNumber="385">Cabeza: más larga que ancha; ojos prominentes; ocelos ubicados detrás de los ojos, no contiguos con el pronoto; paraclípeos gruesos, levemente pasando el anteclípeo, no contiguos; tubérculo antenífero con pequeño diente lateral, antenas unicolores (ocre a rojizo), primer antenómero no sobrepasa el final de la cabeza; rostrum alargado, alcanzando las metacoxas.</emphasis>
</paragraph>
<paragraph id="65794E37D020A403FF3EFA6BB134A88D" blockId="2.[182,1444,1240,2047]" pageId="2" pageNumber="385">
<emphasis id="57B29225D020A403FF3EFA6BB134A88D" bold="true" pageId="2" pageNumber="385">
Tórax: Pronoto densamente puntuado (
<figureCitation id="FDFD52B2D020A403FD00FA6BB338ABF3" box="[720,808,1426,1451]" captionStart="Figs" captionStartId="6.[198,245,1751,1771]" captionTargetBox="[198,1398,219,1701]" captionTargetId="figure-18@6.[198,1408,219,1701]" captionTargetPageId="6" captionText="Figs. 10-14.- Acledra (Acledra) albocostata. 10.- A. (A.) albocostata albocostata, pronoto. 11.- A. (A.) albocostata carinata, pronoto. 12.- A. (A.) albocostata lueri, pronoto. 13.- A. (A.) albocostata lueri, terminalia femenina. 14.- Distribucion de A. (A) albocostata: área azul, A. (A.) albocostata albocostata; círculos rojos, A. (A.) albocostata carinata; cuadrados verdes, A. (A.) albocostata lueri." figureDoi="http://doi.org/10.5281/zenodo.12766008" httpUri="https://zenodo.org/record/12766008/files/figure.png" pageId="2" pageNumber="385">Fig. 13</figureCitation>
), sin parches o líneas impuntuadas, lados cóncavos, ángulos anterolaterales crenulados, ángulos posterolaterales redondeados, cicatrices inmaculadas; escutelo triangular (isósceles), ápice redondeado, y una gruesa carena marfileña a lo largo de la línea media, ángulos basales con dos manchas marfileñas gruesas y puntuadas; hemiélitros sobrepasando el final del abdomen, 
<emphasis id="57B29225D020A403FE72F9DEB1E5A818" bold="true" box="[418,501,1575,1600]" italics="true" pageId="2" pageNumber="385">corium</emphasis>
con una carena marfileña a lo largo de la vena radial seguido de un pequeño parche impuntuado entre la vena radial y el borde del 
<emphasis id="57B29225D020A403FC88F9B5B3BAA83D" bold="true" box="[856,938,1612,1637]" italics="true" pageId="2" pageNumber="385">corium</emphasis>
, membrana ahumada con ṁltiples venas logitudinales; peritrema ostiolar corto (alcanzando 1/3 de la metapleura), evaporatoria rugosa pasando levemente la mitad de la metapleura; patas cubiertas de pilosidad color caramelo, tibias sulcadas.
</emphasis>
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<emphasis id="57B29225D020A402FF3EF918B234AF3C" bold="true" lastPageId="3" lastPageNumber="386" pageId="2" pageNumber="385">
Abdomen: base del abdomen sin espina o tubérculo, peritrema espiracular color caramelo, conexiva aserrada. Genitalia femenina: espermateca alargada y oblonga, bastón esclerotizado grueso con la base ensanchada, ápice agudo precedido de una dilatación circular, dilatación del ducto espermatecal escasamente inflada, rodeando el bastón esclerotizado, reborde proximal de la bomba ovalado y plano, reborde distal de la bomba subcónico, bomba espermatecal ovalada; primeras gonocoxas fuertemente convexas posteriormente (
<figureCitation id="FDFD52B2D020A403FCE1F862B399A9EC" box="[817,905,1947,1972]" captionStart="Figs" captionStartId="6.[198,245,1751,1771]" captionTargetBox="[198,1398,219,1701]" captionTargetId="figure-18@6.[198,1408,219,1701]" captionTargetPageId="6" captionText="Figs. 10-14.- Acledra (Acledra) albocostata. 10.- A. (A.) albocostata albocostata, pronoto. 11.- A. (A.) albocostata carinata, pronoto. 12.- A. (A.) albocostata lueri, pronoto. 13.- A. (A.) albocostata lueri, terminalia femenina. 14.- Distribucion de A. (A) albocostata: área azul, A. (A.) albocostata albocostata; círculos rojos, A. (A.) albocostata carinata; cuadrados verdes, A. (A.) albocostata lueri." figureDoi="http://doi.org/10.5281/zenodo.12766008" httpUri="https://zenodo.org/record/12766008/files/figure.png" pageId="2" pageNumber="385">Fig. 13</figureCitation>
), sección caudal de cada gonocoxito recta; segundas gonocoxas ovaladas y poco conspicuas; paraterguitos 8 subromboidales con el ápice romo; paraterguitos 9 subtriangulares, ápice romo y lados exteriores con una leve concavidad. Genitalia masculina: Pigóforo subrectangular (
<figureCitation id="FDFD52B2D021A402FDA4FF25B2FAAEAD" box="[628,746,220,245]" captionStart="Figs" captionStartId="5.[168,215,1887,1907]" captionTargetBox="[148,1445,612,1846]" captionTargetId="figure-114@5.[148,1445,610,1847]" captionTargetPageId="5" captionText="Figs. 1-9.- Acledra (Acledra) albocostata. 1.- A. (A.) albocostata albocostata, habitus. 2.- A. (A.) albocostata carinata, habitus. 3.- A. (A.) albocostata lueri, habitus. 4.- A. (A.) albocostata albocostata, pigóforo en vista dorsal. 5.- A. (A.) albocostata carinata, pigóforo en vista dorsal. 6.- A. (A.) albocostata lueri, pigóforo en vista dorsal. 7.- A. (A.) albocostata albocostata, pigóforo en vista ventral. 8.- A. (A.) albocostata carinata, pigóforo en vista ventral. 9.- A. (A.) albocostata lueri, pigóforo en vista ventral." figureDoi="http://doi.org/10.5281/zenodo.12766006" httpUri="https://zenodo.org/record/12766006/files/figure.png" pageId="3" pageNumber="386">Figs. 6, 9</figureCitation>
), vista ventral con el ápice bilobulado, en vista dorsal se aprecia un puente apical con dos lóbulos gruesos y una fóvea media apreciable en vista caudal, parámeros con forma de bastón, lóbulo basal más delgado que el apical, lóbulo apical con un diente amplio redondeado en la base.
</emphasis>
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, 
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, 
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, 27/ 
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, leg. A. 
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♂ 
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(
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<emphasis id="57B29225D021A402FAD9FE7AB51DAFC5" bold="true" box="[1289,1293,387,413]" pageId="3" pageNumber="386">.</emphasis>
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<materialsCitation id="D5AE446AD021A402FF3EFE51B58FAF9A" box="[238,1439,424,450]" collectingDate="1968-02-09" collectionCode="DARC, IPUM, HRCC" country="Argentina" location="Epuyen" pageId="3" pageNumber="386" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Chubut" typeStatus="Paratipos">
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: 
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, 
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, 
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, 
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♂, 
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♀
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(
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; 
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), 
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♂, 
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<emphasis id="57B29225D021A402FAC0FE51B50CAF99" bold="true" box="[1296,1308,424,449]" pageId="3" pageNumber="386">1</emphasis>
♀
</specimenCount>
<emphasis id="57B29225D021A402FA93FE51B58FAF9A" bold="true" box="[1347,1439,424,450]" pageId="3" pageNumber="386">
(
<collectionCode id="03D7D6F2D021A402FA9AFE51B588AF9A" box="[1354,1432,424,450]" pageId="3" pageNumber="386">HRCC</collectionCode>
)
</emphasis>
</materialsCitation>
<emphasis id="57B29225D021A402FA4FFE51B5B3AF9A" bold="true" box="[1439,1443,424,450]" pageId="3" pageNumber="386">;</emphasis>
<materialsCitation id="D5AE446AD021A402FF66FE37B3C7AFBF" box="[182,983,461,487]" collectionCode="DARC" collectorName="El Bolson &amp; A. Kovacs" country="Argentina" location="Rio Negro" pageId="3" pageNumber="386" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="Paratipos">
<emphasis id="57B29225D021A402FF66FE37B336AFBF" bold="true" box="[182,806,461,487]" pageId="3" pageNumber="386">
<collectingCountry id="1DD10EA7D021A402FF66FE37B124AFBF" box="[182,308,462,487]" name="Argentina" pageId="3" pageNumber="386">Argentina</collectingCountry>
, 
<location id="601918ECD021A402FE93FE34B1D4AFBF" LSID="urn:lsid:plazi:treatment:ED6FFF21D020A402FF66FB5DB452AA61:601918ECD021A402FE93FE34B1D4AFBF" box="[323,452,461,487]" country="Argentina" name="Rio Negro" pageId="3" pageNumber="386">Río Negro</location>
, 
<collectorName id="C8332BE1D021A402FE04FE37B259AFBF" box="[468,585,461,487]" pageId="3" pageNumber="386">El Bolsón</collectorName>
, leg. 
<collectorName id="C8332BE1D021A402FD5FFE37B31BAFBF" box="[655,779,462,487]" pageId="3" pageNumber="386">A. Kovacs</collectorName>
, 
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</emphasis>
♂, 
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<emphasis id="57B29225D021A402FC9DFE37B349AFBF" bold="true" box="[845,857,462,487]" pageId="3" pageNumber="386">1</emphasis>
♀
</specimenCount>
<emphasis id="57B29225D021A402FCA9FE34B3C7AFBF" bold="true" box="[889,983,461,487]" pageId="3" pageNumber="386">
(
<collectionCode id="03D7D6F2D021A402FC50FE34B3DFAFBF" box="[896,975,461,487]" pageId="3" pageNumber="386">DARC</collectionCode>
)
</emphasis>
</materialsCitation>
<emphasis id="57B29225D021A402FC07FE34B3CBAFBF" bold="true" box="[983,987,461,487]" pageId="3" pageNumber="386">;</emphasis>
<materialsCitation id="D5AE446AD021A402FC37FE37B2E6AC54" collectingDate="1985-09-05" collectionCode="USNM" collectorName="L. Pena" country="Argentina" county="Rio Negro" location="El Maiten" municipality="Bariloche" pageId="3" pageNumber="386" specimenCount="1" specimenCount-male="1" typeStatus="Paratipos">
<emphasis id="57B29225D021A402FC37FE37B27BAC54" bold="true" pageId="3" pageNumber="386">
<collectingCountry id="1DD10EA7D021A402FC37FE37B475AFBF" box="[999,1125,462,487]" name="Argentina" pageId="3" pageNumber="386">Argentina</collectingCountry>
, 
<collectingCounty id="8C1836BBD021A402FBA4FE34B4E5AFBF" box="[1140,1269,461,487]" pageId="3" pageNumber="386">Río Negro</collectingCounty>
, 
<collectingMunicipality id="851DD44DD021A402FAD5FE37B56CAFBF" box="[1285,1404,462,487]" pageId="3" pageNumber="386">Bariloche</collectingMunicipality>
, 
<location id="601918ECD021A402FA5BFE37B11FAC54" LSID="urn:lsid:plazi:treatment:ED6FFF21D020A402FF66FB5DB452AA61:601918ECD021A402FA5BFE37B11FAC54" country="Argentina" county="Rio Negro" municipality="Bariloche" name="El Maiten" pageId="3" pageNumber="386">El Maiten</location>
, 
<date id="117868F7D021A402FEF0FE0AB1B8AC54" box="[288,424,499,524]" pageId="3" pageNumber="386" value="1985-09-05">
<collectingDate id="013C911FD021A402FEF0FE0AB1B8AC54" box="[288,424,499,524]" pageId="3" pageNumber="386" value="1985-09-05">5-IX-1985</collectingDate>
</date>
, leg. 
<collectorName id="C8332BE1D021A402FE21FE0AB25EAC54" box="[497,590,499,524]" pageId="3" pageNumber="386">L. Pena</collectorName>
, 
<specimenCount id="73C085BED021A402FD8FFE0AB27BAC54" box="[607,619,499,524]" count="1" pageId="3" pageNumber="386" type="male">1</specimenCount>
</emphasis>
♂ 
<emphasis id="57B29225D021A402FD5DFE0BB2E6AC54" bold="true" box="[653,758,498,524]" pageId="3" pageNumber="386">
(
<collectionCode id="03D7D6F2D021A402FD45FE0BB2FEAC54" box="[661,750,498,524]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="386" type="Museum">USNM</collectionCode>
)
</emphasis>
</materialsCitation>
<emphasis id="57B29225D021A402FD26FE0BB2EBAC54" bold="true" box="[758,763,498,524]" pageId="3" pageNumber="386">;</emphasis>
<materialsCitation id="D5AE446AD021A402FCD8FE0AB49EAC69" collectingDate="1986-01-22" collectionCode="AMNH" collectorName="Schuh &amp; Muddy" country="Argentina" location="12 km E of Esquivel" pageId="3" pageNumber="386" specimenCount="1" specimenCount-female="1" stateProvince="Chubut" typeStatus="Paratipos">
<emphasis id="57B29225D021A402FCD8FE0AB416AC69" bold="true" pageId="3" pageNumber="386">
<collectingCountry id="1DD10EA7D021A402FCD8FE0AB396AC54" box="[776,902,499,524]" name="Argentina" pageId="3" pageNumber="386">Argentina</collectingCountry>
, 
<collectingRegion id="A70280D5D021A402FC47FE0BB3E2AC54" box="[919,1010,498,524]" country="Argentina" name="Chubut" pageId="3" pageNumber="386">Chubut</collectingRegion>
, 
<location id="601918ECD021A402FBD3FE0AB519AC54" LSID="urn:lsid:plazi:treatment:ED6FFF21D020A402FF66FB5DB452AA61:601918ECD021A402FBD3FE0AB519AC54" box="[1027,1289,499,524]" country="Argentina" name="12 km E of Esquivel" pageId="3" pageNumber="386" stateProvince="Chubut">
<quantity id="A23EE3D2D021A402FBD3FE0AB45FAC54" box="[1027,1103,499,524]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="386" unit="km" value="12.0">12 km</quantity>
E of Esquivel
</location>
, 
<date id="117868F7D021A402FACAFE0AB5B0AC54" box="[1306,1440,499,524]" pageId="3" pageNumber="386" value="1986-01-22">
<collectingDate id="013C911FD021A402FACAFE0AB5B0AC54" box="[1306,1440,499,524]" pageId="3" pageNumber="386" value="1986-01-22">22-I-1986</collectingDate>
</date>
, 
<collectorName id="C8332BE1D021A402FF66FDE1B113AC6A" box="[182,259,536,562]" pageId="3" pageNumber="386">Schuh</collectorName>
&amp; Platnick leg. 
<collectorName id="C8332BE1D021A402FE6FFDE1B202AC69" box="[447,530,536,561]" pageId="3" pageNumber="386">Muddy</collectorName>
shore of pond w/scattered vegetation 
<specimenCount id="73C085BED021A402FC2AFDE1B416AC69" box="[1018,1030,536,561]" count="1" pageId="3" pageNumber="386" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="57B29225D021A402FBF3FDE1B49EAC69" bold="true" box="[1059,1166,536,561]" pageId="3" pageNumber="386">
(
<collectionCode id="03D7D6F2D021A402FBFBFDE1B495AC69" box="[1067,1157,536,561]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="386">AMNH</collectionCode>
)
</emphasis>
</materialsCitation>
<emphasis id="57B29225D021A402FB5EFDE1B483AC69" bold="true" box="[1166,1171,536,561]" pageId="3" pageNumber="386">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2DDC1DBCD021A402FF3EFDB6B452AA61" pageId="3" pageNumber="386" type="diagnosis">
<paragraph id="65794E37D021A402FF3EFDB6B563AD84" blockId="3.[182,1444,591,988]" pageId="3" pageNumber="386">
<emphasis id="57B29225D021A402FF3EFDB6B563AD84" bold="true" pageId="3" pageNumber="386">
Diagnosis diferencial: Esta subespecie puede reconocerse dentro del complejo de 
<taxonomicName id="A2C635B4D021A402FB20FDB6B5B3AC30" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[1264,1443,591,616]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="albocostata">
<emphasis id="57B29225D021A402FB20FDB6B5B3AC30" bold="true" box="[1264,1443,591,616]" italics="true" pageId="3" pageNumber="386">A. albocostata</emphasis>
</taxonomicName>
por las siguientes características: de 
<taxonomicName id="A2C635B4D021A402FD61FD8DB45AACD5" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[689,1098,628,653]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="subSpecies" species="albocostata" subGenus="Acledra" subSpecies="albocostata">
<emphasis id="57B29225D021A402FD61FD8DB45AACD5" bold="true" box="[689,1098,628,653]" italics="true" pageId="3" pageNumber="386">A. (A.) albocostata albocostata</emphasis>
</taxonomicName>
se diferencia por poseer callosidades laterales del escutelo más reducidas, también en la genitalia masculina, los lóbulos del puente superior del pigóforo son mucho más amplios en 
<taxonomicName id="A2C635B4D021A402FCBDFD46B487AC80" authorityName="Fȧndez &amp; Rider" authorityYear="2016" box="[877,1175,703,728]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="subSpecies" species="albocostata" subGenus="Acledra" subSpecies="lueri">
<emphasis id="57B29225D021A402FCBDFD46B487AC80" bold="true" box="[877,1175,703,728]" italics="true" pageId="3" pageNumber="386">A. (A.) albocostata lueri</emphasis>
</taxonomicName>
, carácter mucho más notorio en vista ventral. De 
<taxonomicName id="A2C635B4D021A402FDF8FD1DB39DACA5" authorityName="Breddin" authorityYear="1914" baseAuthorityName="Breddin" baseAuthorityYear="1914" box="[552,909,740,765]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="subSpecies" species="albocostata" subGenus="Acledra" subSpecies="carinata">
<emphasis id="57B29225D021A402FDF8FD1DB39DACA5" bold="true" box="[552,909,740,765]" italics="true" pageId="3" pageNumber="386">A. (A.) albocostata carinata</emphasis>
</taxonomicName>
se distingue por poseer un escutelo más ensanchado en el ápice, pronoto más compacto, en la genitalia femenina los paraterguitos 8 y 9 son más compactos, y en la genitalia masculina los lóbulos el puente superior del pigóforo son mucho más amplios. Fuera del grupo de 
<taxonomicName id="A2C635B4D021A402FDCCFCAAB2C1AD34" baseAuthorityName="Spinola" baseAuthorityYear="1852" box="[540,721,851,876]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="albocostata">
A. 
<emphasis id="57B29225D021A402FD91FCAAB2C1AD34" bold="true" box="[577,721,851,876]" italics="true" pageId="3" pageNumber="386">albocostata</emphasis>
</taxonomicName>
, esta subespecie tiene similitudes con 
<taxonomicName id="A2C635B4D021A402FB13FCAAB18CADC9" authority="Fa" authorityName="Fa" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="bichromocornis" subGenus="Acledra">
<emphasis id="57B29225D021A402FB13FCAAB162ADCA" bold="true" italics="true" pageId="3" pageNumber="386">Acledra (Acledra) bichromocornis</emphasis>
Fa
</taxonomicName>
̇ndez, 
<bibRefCitation id="015733C6D021A402FE21FC80B317ADCA" author="Rider, D. A. &amp; Carvajal, M. A." box="[497,775,888,914]" pageId="3" pageNumber="386" pagination="127 - 134" refId="ref3590" refString="Fa ˙ ndez, E. I.; Rider, D. A. &amp; Carvajal, M. A. 2014. A new species of Acledra s. str. (Hemiptera: Heteroptera: Pentatomidae) from the highlands of Argentina and Bolivia, with a checklist and key to the species of the nominate subgenus. Zootaxa, 3900 (1): 127 - 134." type="journal article" year="2014">Rider &amp; Carvajal, 2014</bibRefCitation>
y 
<taxonomicName id="A2C635B4D021A402FCF7FC80B4EEADCA" authority="Berg, 1878" authorityName="Berg" authorityYear="1878" box="[807,1278,889,914]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="gregalis" subGenus="Acledra">
<emphasis id="57B29225D021A402FCF7FC80B47EADCA" bold="true" box="[807,1134,889,914]" italics="true" pageId="3" pageNumber="386">Acledra (Acledra) gregalis</emphasis>
Berg, 1878
</taxonomicName>
de las cuales se distingue fácilmente por poseer una carena marfileña central a lo largo el escutelo; mientras que en 
<emphasis id="57B29225D021A402FF0AFC3AB1FFAD84" bold="true" box="[218,495,963,988]" italics="true" pageId="3" pageNumber="386">
A. (A). 
<taxonomicName id="A2C635B4D021A402FEE2FC3AB1FFAD84" box="[306,495,963,988]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="bichromocornis">bichromocornis</taxonomicName>
</emphasis>
ésta está ausente, y 
<emphasis id="57B29225D021A402FD28FC3AB3C5AD84" bold="true" box="[760,981,963,988]" italics="true" pageId="3" pageNumber="386">
en 
<taxonomicName id="A2C635B4D021A402FCCCFC3AB3C5AD84" authorityName="Berg" authorityYear="1878" box="[796,981,963,988]" class="Insecta" family="Pentatomidae" genus="Acledra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="gregalis" subGenus="Acledra">A. (A.) gregalis</taxonomicName>
</emphasis>
ésta se restringe a la zona basal.
</emphasis>
</paragraph>
<paragraph id="65794E37D021A402FF3EFC03B452AA61" blockId="3.[182,1444,1018,1081]" pageId="3" pageNumber="386">
<emphasis id="57B29225D021A402FF3EFC03B452AA61" bold="true" pageId="3" pageNumber="386">
Etimología: Dedicamos esta subespecie a Alfredo Lüer, en reconocimiento a sus valiosas recolecciones de heterópteros, incluyendo el 
<typeStatus id="BA7DF095D021A402FD35FBD9B358AA61" box="[741,840,1056,1081]" pageId="3" pageNumber="386" type="holotype">holotipo</typeStatus>
de esta subespecie.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>