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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1146.95004" ID-Pensoft-Pub="1313-2970-1146-135" ID-Pensoft-UUID="73ABDE9A1ED35289B5F3433C37ECCA59" ID-ZooBank="84DB12238E0F4837BFD25616E692DAF1" ModsDocID="1313-2970-1146-135" checkinTime="1675846402594" checkinUser="pensoft" docAuthor="Hookabe, Natsumi, Kohtsuka, Hisanori, Fujiwara, Yoshihiro, Tsuchida, Shinji &amp; Ueshima, Rei" docDate="2023" docId="E64BFBDD40A559129F6C3659A7D127E0" docLanguage="en" docName="ZooKeys 1146: 135-146" docOrigin="ZooKeys 1146" docPubDate="2023-02-07" docSource="http://dx.doi.org/10.3897/zookeys.1146.95004" docTitle="Tetrastemma shohoense Hookabe &amp; Kohtsuka &amp; Fujiwara &amp; Tsuchida &amp; Ueshima 2023, sp. nov." docType="treatment" docUuid="9D1CD900-900F-4114-8853-16C480FDD75D" docUuidSource="ZooBank" docVersion="1" id="73ABDE9A1ED35289B5F3433C37ECCA59" lastPageNumber="135" masterDocId="73ABDE9A1ED35289B5F3433C37ECCA59" masterDocTitle="Three new species in Tetrastemma Ehrenberg, 1828 (Nemertea, Monostilifera) from sublittoral to upper bathyal zones of the northwestern Pacific" masterLastPageNumber="146" masterPageNumber="135" pageNumber="135" updateTime="1675846402594" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Three new species in Tetrastemma Ehrenberg, 1828 (Nemertea, Monostilifera) from sublittoral to upper bathyal zones of the northwestern Pacific</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hookabe, Natsumi</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3498-8614</mods:nameIdentifier>
<mods:affiliation>School of Science, The University of Tokyo, 113 - 0033, Bunkyo, Japan</mods:affiliation>
<mods:nameIdentifier type="email">sofeechan312@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kohtsuka, Hisanori</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4440-2776</mods:nameIdentifier>
<mods:affiliation>Misaki Marine Biological Station (MMBS), School of Science, The University of Tokyo, 238 - 0225, Miura, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fujiwara, Yoshihiro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1833-1866</mods:nameIdentifier>
<mods:affiliation>Research Institute for Global Change (RIGC), Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, 237 - 0061, Kanagawa, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tsuchida, Shinji</mods:namePart>
<mods:affiliation>Research Institute for Global Change (RIGC), Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, 237 - 0061, Kanagawa, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ueshima, Rei</mods:namePart>
<mods:affiliation>School of Science, The University of Tokyo, 113 - 0033, Bunkyo, Japan</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-07</mods:number>
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<mods:detail type="volume">
<mods:number>1146</mods:number>
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<mods:start>135</mods:start>
<mods:end>146</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1146.95004</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1146.95004</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1146-135</mods:identifier>
<mods:identifier type="ZooBank">84DB12238E0F4837BFD25616E692DAF1</mods:identifier>
<mods:identifier type="Pensoft-UUID">73ABDE9A1ED35289B5F3433C37ECCA59</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:9D1CD900-900F-4114-8853-16C480FDD75D" httpUri="http://treatment.plazi.org/id/E64BFBDD40A559129F6C3659A7D127E0" lastPageNumber="135" pageId="0" pageNumber="135">
<subSubSection pageId="0" pageNumber="135" type="nomenclature">
<paragraph pageId="0" pageNumber="135">
<taxonomicName LSID="https://zoobank.org/9D1CD900-900F-4114-8853-16C480FDD75D" authority="Hookabe &amp; Kohtsuka &amp; Fujiwara &amp; Tsuchida &amp; Ueshima, 2023" authorityName="Hookabe &amp; Kohtsuka &amp; Fujiwara &amp; Tsuchida &amp; Ueshima" authorityYear="2023" class="Enopla" family="Tetrastemmatidae" genus="Tetrastemma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tetrastemma shohoense" order="Hoplonemertea" pageId="0" pageNumber="135" phylum="Nemertea" rank="species" species="shohoense" status="sp. nov.">Tetrastemma shohoense</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="135">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="description">
<paragraph pageId="0" pageNumber="135">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">Fig. 2E-I [New Japanese name: shoho-kakubari-himomushi]</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="etymology">
<paragraph pageId="0" pageNumber="135">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="135">The species is named after the type locality, Shoho Seamount of the Nishi-Shichito Ridge, Japan.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="materials_examined">
<paragraph pageId="0" pageNumber="135">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="135">
<materialsCitation collectingDate="2020-11-29" collectorName="V Kaimei, NW Pacific." country="Japan" elevation="455" latitude="32.323166" location="Japan" longLatPrecision="12" longitude="138.74133" specimenCount="1" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="135">
<typeStatus>Holotype</typeStatus>
</emphasis>
: NMST-Nem-H-05, unsectioned complete specimen except for the posterior tip, fixed in
<normalizedToken originalValue="Bouins">Bouin's</normalizedToken>
fluid, posterior tip preserved in 99% ethanol, collected on
<collectingDate value="2020-11-29">November 29 2020</collectingDate>
by NH, by use of ROV
<emphasis italics="true" pageId="0" pageNumber="135">KM-ROV</emphasis>
(dive #123) during KM20-10C cruise of R/
<collectorName>
V
<emphasis italics="true" pageId="0" pageNumber="135">Kaimei</emphasis>
</collectorName>
, at a depth of
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.55" unit="m" value="455.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.55" unit="m" value="455.0">455 m</elevation>
</quantity>
, near the summit of Shoho Seamount of the Nishi-Shichito Ridge (
<geoCoordinate degrees="32" direction="north" minutes="19.39" orientation="latitude" precision="9" value="32.323166">32°19.39'N</geoCoordinate>
,
<geoCoordinate degrees="138" direction="east" minutes="44.48" orientation="longitude" precision="9" value="138.74133">138°44.48'E</geoCoordinate>
),
<collectingCountry name="Japan">Japan</collectingCountry>
,
<collectorName>NW Pacific.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="description">
<paragraph pageId="0" pageNumber="135">Description.</paragraph>
<paragraph pageId="0" pageNumber="135">
Head spatulate in profile (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2E-H</figureCitation>
), not well demarcated from body by anterior cephalic furrows (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2E</figureCitation>
). Before anesthetization, body of a live specimen 5.5 mm long and 0.3 mm wide. Background body color generally white, tinged with bright yellow to orange, and almost transparent (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2E</figureCitation>
). Head with a red rectangle cephalic patch without extending behind a posterior pair of eyes (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2E, H</figureCitation>
). Anterior pair of cephalic furrows present (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2F</figureCitation>
) but posterior one not well distinguished. Cerebral ganglia and blood uncolored (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2G, H</figureCitation>
). Alimentary canals visible as bright yellow organs through body wall (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2E</figureCitation>
). Proboscis pale, extending about 3/4 of the body length (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2E</figureCitation>
). Four brown eyes present; anterior pair slightly larger than posterior ones (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2H</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="135">
Stylet basis cylindrical, 55.0
<normalizedToken originalValue="μm">μm</normalizedToken>
in length and 25.0
<normalizedToken originalValue="μm">μm</normalizedToken>
in maximum width; central stylet smooth, 47.0
<normalizedToken originalValue="μm">μm</normalizedToken>
in length; (stylet length) / (basis length) ratio 0.85 (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2I</figureCitation>
). Two accessory stylet pouches present, each containing two stylets (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype specimens of new Tetrastemma species; photographs were taken in life by NH A-C T. album sp. nov. A complete body, dorsal view B head, dorsal view C head, ventral view D T. persona sp. nov., complete body, dorsal view E-I T. shohoense sp. nov. E complete body F head, ventral view G squeezed specimen under a cover slip, complete body, dorsal view H head, dorsal view I stylet apparatus. Abbreviations: acf, anterior cephalic furrow; pcf, posterior cephalic furrow, cg, cerebral ganglia; co, cerebral organ; pb, proboscis; pp, proboscis pore. Scale bars: 2 mm (A); 500 μm (B, C, G); 1 mm (D, E); 100 μm (F, H); 50 μm (I)." figureDoi="10.3897/zookeys.1146.95004.figure2" httpUri="https://binary.pensoft.net/fig/806733" pageId="0" pageNumber="135">2I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="type locality">
<paragraph pageId="0" pageNumber="135">Type locality and distribution.</paragraph>
<paragraph pageId="0" pageNumber="135">
The species is only known from the type locality, Shoho Seamount of the Nishi-Shichito Ridge, Japan, at a depth of 455 m (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites of the specimens examined in the present study." figureDoi="10.3897/zookeys.1146.95004.figure1" httpUri="https://binary.pensoft.net/fig/806732" pageId="0" pageNumber="135">1</figureCitation>
), among the sandy sediments on rocky substrates.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="135" type="remarks">
<paragraph pageId="0" pageNumber="135">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="135">
Having a dark cephalic patch and cylindrical stylet basis and lacking a longitudinal line on the dorsal surface of the body,
<taxonomicName class="Hoplonemertea" kingdom="Animalia" lsidName="T. shohoense" order="Monostilifera" pageId="0" pageNumber="135" phylum="Nemertea" rank="species" species="shohoense">
<emphasis italics="true" pageId="0" pageNumber="135">T. shohoense</emphasis>
</taxonomicName>
sp. nov. resembles
<taxonomicName class="Hoplonemertea" kingdom="Animalia" lsidName="T. freyae" order="Monostilifera" pageId="0" pageNumber="135" phylum="Nemertea" rank="species" species="freyae">
<emphasis italics="true" pageId="0" pageNumber="135">T. freyae</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4808.2.10" author="Chernyshev, AV" journalOrPublisher="Zootaxa" pageId="0" pageNumber="135" pagination="379 - 383" refId="B5" refString="Chernyshev, AV, Polyakova, NE, Vignesh, MS, Jain, RP, Sanjeevi, P, Norenburg, JL, Rajesh, RP, 2020. A histology-free description of a new species of the genus Tetrastemma (Nemertea: Hoplonemertea: Monostilifera) from Hawaii and India. Zootaxa 4808 (2): 379 - 383, DOI: https://doi.org/10.11646/zootaxa.4808.2.10" title="A histology-free description of a new species of the genus Tetrastemma (Nemertea: Hoplonemertea: Monostilifera) from Hawaii and India." url="https://doi.org/10.11646/zootaxa.4808.2.10" volume="4808" year="2020">Chernyshev et al., 2020</bibRefCitation>
originally described based on Hawaiian and Indian specimens. The new species is differentiated from
<taxonomicName class="Hoplonemertea" kingdom="Animalia" lsidName="T. freyae" order="Monostilifera" pageId="0" pageNumber="135" phylum="Nemertea" rank="species" species="freyae">
<emphasis italics="true" pageId="0" pageNumber="135">T. freyae</emphasis>
</taxonomicName>
in the color of the cephalic patch as well as the non-flared posterior margin of the cylindrical stylet basis.
</paragraph>
<paragraph pageId="0" pageNumber="135">
A genetic distance based on COI between
<taxonomicName class="Hoplonemertea" kingdom="Animalia" lsidName="T. shohoense" order="Monostilifera" pageId="0" pageNumber="135" phylum="Nemertea" rank="species" species="shohoense">
<emphasis italics="true" pageId="0" pageNumber="135">T. shohoense</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Hoplonemertea" kingdom="Animalia" lsidName="T. freyae" order="Monostilifera" pageId="0" pageNumber="135" phylum="Nemertea" rank="species" species="freyae">
<emphasis italics="true" pageId="0" pageNumber="135">T. freyae</emphasis>
</taxonomicName>
(specimens from Hawaii (MT247877) and India (MT247878) was 12.6%; the value is comparable with interspecific values observed among
<taxonomicName class="Hoplonemertea" kingdom="Animalia" lsidName="" order="Monostilifera" pageId="0" pageNumber="135" phylum="Nemertea" rank="order">Monostilifera</taxonomicName>
(e.g.,
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0155541" author="Sundberg, P" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="135" refId="B30" refString="Sundberg, P, Kvist, S, Strand, M, 2016. Evaluating the utility of single-locus DNA barcoding for the identification of ribbon worms (phylum Nemertea). PLoS ONE 11(5): e0155541. https://doi.org/10.1371/journal.pone.0155541" title="Evaluating the utility of single-locus DNA barcoding for the identification of ribbon worms (phylum Nemertea). PLoS ONE 11 (5): e 0155541." url="https://doi.org/10.1371/journal.pone.0155541" year="2016">Sundberg et al. 2016</bibRefCitation>
;
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).
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