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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1072.72978" ID-GBIF-Dataset="5abb2e98-44e8-4554-9e15-fa892088595d" ID-PMC="PMC8629907" ID-Pensoft-Pub="1313-2970-1072-129" ID-Pensoft-UUID="9D8AA58A90535BAAB59216914F04E621" ID-PubMed="34899010" ModsDocID="1313-2970-1072-129" checkinTime="1637640646787" checkinUser="pensoft" docAuthor="Ng, Peter K. L. &amp; Yang, Chien-Hui" docDate="2021" docId="E558347305695F91A87119A9F9822336" docLanguage="en" docName="ZooKeys 1072: 129-165" docOrigin="ZooKeys 1072" docPubDate="2021-11-22" docSource="http://dx.doi.org/10.3897/zookeys.1072.72978" docTitle="Dicranodromia robusta Ng &amp; Yang 2021, sp. nov." docType="treatment" docVersion="4" id="9D8AA58A90535BAAB59216914F04E621" lastPageNumber="129" masterDocId="9D8AA58A90535BAAB59216914F04E621" masterDocTitle="On two new species of deep-sea carrier crabs (Crustacea, Brachyura, Homolodromiidae, Dicranodromia) from Taiwan and the Philippines, with notes on other Indo-West Pacific species" masterLastPageNumber="165" masterPageNumber="129" pageNumber="129" updateTime="1668151158991" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>On two new species of deep-sea carrier crabs (Crustacea, Brachyura, Homolodromiidae, Dicranodromia) from Taiwan and the Philippines, with notes on other Indo-West Pacific species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5946-0608</mods:nameIdentifier>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Chien-Hui</mods:namePart>
<mods:affiliation>National Taiwan Ocean University, 2 Pei-Ning Road, Keelung 20224, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">chyang@ntou.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1072</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>165</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1072.72978</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1072.72978</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1072-129</mods:identifier>
<mods:identifier type="Pensoft-UUID">9D8AA58A90535BAAB59216914F04E621</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="129" type="systematic account">
<treatment ID-GBIF-Taxon="190549118" LSID="urn:lsid:zoobank.org:act:ADCE5085-D836-4A3E-8CF9-5BEB79ED3FE1" httpUri="http://treatment.plazi.org/id/E558347305695F91A87119A9F9822336" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
<taxonomicName LSID="http://zoobank.org/ADCE5085-D836-4A3E-8CF9-5BEB79ED3FE1" authority="Ng &amp; Yang, 2021" authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">Dicranodromia robusta</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="129">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">Figures 17</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of left chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure18" httpUri="https://binary.pensoft.net/fig/612924" pageId="0" pageNumber="129">, 18</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A right P 3 B right P 5 C right P 5 propodus and dactylus D left P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca." figureDoi="10.3897/zookeys.1072.72978.figure19" httpUri="https://binary.pensoft.net/fig/612925" pageId="0" pageNumber="129">, 19</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">, 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">, 21C, G-J, N-P</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="materials_examined">
<paragraph pageId="0" pageNumber="129">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="129">
<materialsCitation collectingDate="2017-11-26" country="Philippines" county="Balut Island" latitude="5.4" location="Occidental Province" longLatPrecision="930" longitude="125.375" municipality="Islands" specimenCount="♀" stateProvince="Sarangani" typeStatus="Holotype">
<collectingCountry name="Philippines">Philippines</collectingCountry>
:
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">
<typeStatus>Holotype</typeStatus>
</emphasis>
<specimenCount></specimenCount>
(19.6
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" unit="mm" value="26.4">26.4 mm</quantity>
), ca.
<geoCoordinate degrees="5" direction="north" minutes="24" orientation="latitude" precision="925" value="5.4">5°24'N</geoCoordinate>
,
<geoCoordinate degrees="125" direction="east" minutes="22.5" orientation="longitude" precision="92" value="125.375">125°22.5'E</geoCoordinate>
,
<collectingCounty>Balut Island</collectingCounty>
,
<collectingRegion country="Philippines" name="Sarangani">Sarangani</collectingRegion>
<collectingMunicipality>Islands</collectingMunicipality>
,
<collectingRegion country="Philippines" name="Davao">Davao</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:E558347305695F91A87119A9F9822336:8BDD4602960FBD53EECD098254F13CB0" country="Philippines" county="Balut Island" latitude="5.4" longLatPrecision="930" longitude="125.375" municipality="Islands" name="Occidental Province" stateProvince="Sarangani">Occidental Province</location>
, south of
<location LSID="urn:lsid:plazi:treatment:E558347305695F91A87119A9F9822336:AA876393F5563AF08DA1AFB7ED205844" country="Philippines" county="Balut Island" latitude="5.4" longLatPrecision="930" longitude="125.375" municipality="Islands" name="Mindanao Island" stateProvince="Sarangani">Mindanao Island</location>
, coll. tangle nets, local fishermen,
<collectingDate value="2017-11-26">26 Nov. 2017</collectingDate>
(ZRC 2018.0161)
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Philippines" county="Balut Island" latitude="5.4" location="Occidental Province" longLatPrecision="930" longitude="125.375" municipality="Islands" specimenCount="♂" stateProvince="Sarangani" typeStatus="Paratype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount></specimenCount>
(15.2
<normalizedToken originalValue="×">x</normalizedToken>
21.0 mm), same location as holotype, coll. tangle nets, local fishermen, 2017 (ZRC 2018.0095)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">
Carapace longitudinally subquadrate, widest across intestinal-mesobranchial regions; dorsal surface gently convex, lateral surfaces covered with low spinules, median part smooth, margins with scattered short stiff setae, not obscuring margins; short stiff setae present on pereiopods, thoracic sternum and pleon but not obscuring surface or margins. Branchiocardiac groove distinct, curving medially anteriorly. Each pseudorostral lobe triangular, inner margin straight, outer margin gently convex, directed anteriorly, inner margin entire; exorbital tooth dentiform, directed obliquely laterally, anterior margin with two or three spinules; supraorbital margin separated from external orbital tooth by shallow concave cleft, posterior part with five or six spinules; infraorbital margin with large dorsoventrally flattened lobe which is dentiform to linguiform, larger than exorbital tooth, distal part with spine, anterior margin with two spinules, prominently visible in dorsal view. Rostrum present as one sharp granule. Epistome covered with scattered granules on anterior half; posterior half gently upturned, with median fissure, surface not covered with spinules, posterior margin gently convex, median part entire, lateral part gently serrate. Basal antennal article subquadrate; surfaces covered by spinules and granules; anteroexternal tooth short. Eyes with long peduncle. Third maxilliped relatively narrow; merus subovate with low anterointernal lobe, shorter than ischium; ischium subtrapezoidal, distal half slightly wider than proximal part; palp (carpus, propodus, dactylus) long, reaching to median part of ischium when folded; exopod with proximal third widest. Chelipeds covered with stiff setae on most parts; merus and carpus with margins uneven or lined with granules; palm relatively short, subdorsal and subventral margins with low sharp granules, median part smooth; fingers thick, wide, occluding surface hollowed; pollex with deep U-shaped depression distally. P2 and P3 relatively short, P3 longer than P2; merus with low tooth on distal extensor margin, length to width ratio of P2 and P3 merus 4.2 and 3.9, respectively; margins unarmed; propodus almost straight, unarmed, length to width ratio of P2 and P3 propodus 5.2 and 6.4, respectively; dactylus curved, flexor margin lined with 8 or 9 spines, terminating in strongly gently curved claw, propodus about 2.4
<normalizedToken originalValue="×">x</normalizedToken>
length of dactylus. P4 stouter, shorter than P5; length to width ratio of P4 and P5 merus 2.4 and 3.4, respectively; margins of merus unarmed; P4 and P5 propodus with submedian spinule on distal third of outer surface, length to width ratio of P4 and P5 propodus 2.3 and 3.6, respectively, distal margin fringed by sharp spines bracketing dactylus; dactylus claw-like, strongly incurved, extensor margin with median spine or absent, flexor margin with 2-4 spines. Thoracic sternite 7 with low transverse ridge from posterior inner part of female gonopore, lateral part high, forming triangular tubercle, curving posteriorly to join oblique ridge formed by posterior part of sternite 7, just before suture with sternite 8, groove between sternites 7 and 8 curve to join spermathecal aperture at base of triangular tubercle. Male and female pleons with 6 free somites and telson; male telson distinctly elongate, triangular with gently convex lateral margins; female telson triangular, with gently convex margins. G1 stout, endopod distally covered by dense long setae, subdistal part of outer margin with two lobes, the distal one being more prominent; G2 endopod gradually tapering to sharp tip.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="variation">
<paragraph pageId="0" pageNumber="129">Variation.</paragraph>
<paragraph pageId="0" pageNumber="129">
In the holotype female, the left P5 dactylus has a prominent spine on the extensor margin (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">21I</figureCitation>
), but there is none on the right side (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">21H</figureCitation>
). The P5 dactyli of the paratype male are armed a spinule on the extensor margin. Both specimens possess the spine on the outer surface of the P5 propodus (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">21H, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="etymology">
<paragraph pageId="0" pageNumber="129">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="129">
The species is named after the Latin
<emphasis italics="true" pageId="0" pageNumber="129">robusta</emphasis>
for stout, alluding to the stocky appearance of the species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="remarks">
<paragraph pageId="0" pageNumber="129">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="129">
The most diagnostic character of
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">D. robusta</emphasis>
</taxonomicName>
sp. nov. is the large dorsoventrally flattened infraorbital tooth, which is dentiform to linguiform, clearly visible in dorsal view, and distinctly larger than the exorbital tooth (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">17B-D</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">20B</figureCitation>
). No other Indo-West Pacific species of
<taxonomicName authorityName="A. Milne-Edwards" authorityYear="1880" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia</emphasis>
</taxonomicName>
has such a large and wide infraorbital tooth. The long anteroexternal tooth on the basal antennal article allies
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">D. robusta</emphasis>
</taxonomicName>
with
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. martini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="0" pageNumber="129">D. martini</emphasis>
</taxonomicName>
(cf.
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 20B),
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. baffini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="baffini">
<emphasis italics="true" pageId="0" pageNumber="129">D. baffini</emphasis>
</taxonomicName>
(cf.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10330" author="Alcock, A" journalOrPublisher="Keelung, Taiwan" pageId="0" pageNumber="129" refId="B3" refString="Alcock, A, 1899. An account of the Deep-Sea Brachyura collected by the Royal Indian marine survey ship INVESTIGATOR. Printed by order of the Trustees of the Indian Museum, Calcutta, 87 pp, pls 1-4. https://doi.org/10.5962/bhl.title.10330" title="An account of the Deep-Sea Brachyura collected by the Royal Indian marine survey ship INVESTIGATOR. Printed by order of the Trustees of the Indian Museum, Calcutta, 87 pp, pls 1 - 4." url="https://doi.org/10.5962/bhl.title.10330" year="1899">Alcock 1899</bibRefCitation>
: pl. 2 fig. 1a;
<bibRefCitation author="Alcock, A" journalOrPublisher="Keelung, Taiwan" pageId="0" pageNumber="129" refId="B4" refString="Alcock, A, 1901. Catalogue of the Indian Decapod Crustacea in the collection of the Indian Museum. Part 1. Brachyura. Fasciculus 1. lntroduction and Dromides or Dromiacea (Brachyura Primigenia). Trustees of the Indian Museum, Calcutta. 1-80." title="Catalogue of the Indian Decapod Crustacea in the collection of the Indian Museum. Part 1. Brachyura. Fasciculus 1. lntroduction and Dromides or Dromiacea (Brachyura Primigenia). Trustees of the Indian Museum, Calcutta. 1 - 80." year="1901">Alcock 1901</bibRefCitation>
: pl. 1 fig. 1a),
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. danielae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="danielae">
<emphasis italics="true" pageId="0" pageNumber="129">D. danielae</emphasis>
</taxonomicName>
(cf.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1029.1.3" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="39 - 46" refId="B26" refString="Ng, PKL, McLay, CL, 2005. Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura). Zootaxa 1029: 39 - 46, DOI: https://doi.org/10.11646/zootaxa.1029.1.3" title="Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1029.1.3" volume="1029" year="2005">Ng and McLay 2005</bibRefCitation>
: fig. 3A, B) and
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. chenae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="chenae">
<emphasis italics="true" pageId="0" pageNumber="129">D. chenae</emphasis>
</taxonomicName>
(cf.
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="129" pagination="47 - 53" refId="B28" refString="Ng, PKL, Naruse, T, 2007. On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species. Raffles Bulletin of Zoology, Supplement 16: 47 - 53" title="On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species." volume="16" year="2007">Ng and Naruse 2007</bibRefCitation>
: fig. 5b) but the structure of the infraorbital tooth easily distinguishes it from them.
</paragraph>
<caption doi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 17.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia robusta</emphasis>
</taxonomicName>
sp. nov., holotype ♀ (19.6
<normalizedToken originalValue="×">x</normalizedToken>
26.4 mm) (ZRC 2018.0161), Philippines
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
overall view
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
dorsal view of carapace
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
front and anterior part of carapace
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
right front, orbit and first anterolateral spine
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1072.72978.figure18" httpUri="https://binary.pensoft.net/fig/612924" pageId="0" pageNumber="129" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 18.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia robusta</emphasis>
</taxonomicName>
sp. nov., holotype ♀ (19.6
<normalizedToken originalValue="×">x</normalizedToken>
26.4 mm) (ZRC 2018.0161), Philippines
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
ventral view of cephalothorax
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
left third maxilliped
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
frontal view of cephalothorax
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
epistome, antennules, antennae and orbits
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
dorsal view of left chela
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
left chela.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1072.72978.figure19" httpUri="https://binary.pensoft.net/fig/612925" pageId="0" pageNumber="129" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 19.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia robusta</emphasis>
</taxonomicName>
sp. nov., holotype ♀ (19.6
<normalizedToken originalValue="×">x</normalizedToken>
26.4 mm) (ZRC 2018.0161), Philippines
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
right P3
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
right P5
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
right P5 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
left P4 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
anterior thoracic sternum and spermatheca
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
posterior thoracic sternum showing spermatheca.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="129">
The carapace shape of
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">D. robusta</emphasis>
</taxonomicName>
is distinctly more quadrate (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">17A, B</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">20A, B</figureCitation>
) than the more pyriform
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. martini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="0" pageNumber="129">D. martini</emphasis>
</taxonomicName>
described from the Philippines (cf. Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dicranodromia martini Guinot, 1995, ♀ (28.1 v 34.2 mm) (ZRC 2007.0106), Philippines A overall view B right third maxilliped C dorsal view of carapace D anterior part of carapace (right side denuded) E frontal view of cephalothorax." figureDoi="10.3897/zookeys.1072.72978.figure4" httpUri="https://binary.pensoft.net/fig/612909" pageId="0" pageNumber="129">4A, C</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dicranodromia martini Guinot, 1995, ♂ (12.3 x 16.6 mm) (ZRC 2007.0105), Philippines A overall view B dorsal view of carapace C anterior part of carapace (partially denuded) D male telson." figureDoi="10.3897/zookeys.1072.72978.figure6" httpUri="https://binary.pensoft.net/fig/612912" pageId="0" pageNumber="129">6A, B</figureCitation>
;
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 19b;
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="129" pagination="47 - 53" refId="B28" refString="Ng, PKL, Naruse, T, 2007. On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species. Raffles Bulletin of Zoology, Supplement 16: 47 - 53" title="On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species." volume="16" year="2007">Ng and Naruse 2007</bibRefCitation>
: fig. 1a-c); the posterior margin of the epistome is entire (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of left chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure18" httpUri="https://binary.pensoft.net/fig/612924" pageId="0" pageNumber="129">18C</figureCitation>
) (versus margin gently crenulate in subventral view in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. martini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="0" pageNumber="129">D. martini</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 20B); P2 and P3 are prominently shorter with the dactylus especially short (e.g., P3 merus 3.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, propodus 6.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A right P 3 B right P 5 C right P 5 propodus and dactylus D left P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca." figureDoi="10.3897/zookeys.1072.72978.figure19" httpUri="https://binary.pensoft.net/fig/612925" pageId="0" pageNumber="129">19A</figureCitation>
) (versus P3 merus 7.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, propodus 7.2
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. martini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="0" pageNumber="129">D. martini</emphasis>
</taxonomicName>
, cf. Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dicranodromia martini Guinot, 1995, ♀ (28.1 v 34.2 mm) (ZRC 2007.0106), Philippines A overall view B right third maxilliped C dorsal view of carapace D anterior part of carapace (right side denuded) E frontal view of cephalothorax." figureDoi="10.3897/zookeys.1072.72978.figure4" httpUri="https://binary.pensoft.net/fig/612909" pageId="0" pageNumber="129">4A</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dicranodromia martini Guinot, 1995, ♀ (28.1 x 34.2 mm) (ZRC 2007.0106), Philippines A epistome, antennules, antennae and orbits B left chela C right P 3 D right P 5 E right P 5 dactylus F right P 4 G telson H posterior thoracic sternum showing spermatheca and female gonopores." figureDoi="10.3897/zookeys.1072.72978.figure5" httpUri="https://binary.pensoft.net/fig/612911" pageId="0" pageNumber="129">5C</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dicranodromia martini Guinot, 1995, ♂ (12.3 x 16.6 mm) (ZRC 2007.0105), Philippines A overall view B dorsal view of carapace C anterior part of carapace (partially denuded) D male telson." figureDoi="10.3897/zookeys.1072.72978.figure6" httpUri="https://binary.pensoft.net/fig/612912" pageId="0" pageNumber="129">6A</figureCitation>
;
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: figs 19a, e, 20C); P4 and P5 are much shorter (e.g., P4 merus just reaching branchiocardiac groove when folded dorsally, Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">17A</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A right P 3 B right P 5 C right P 5 propodus and dactylus D left P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca." figureDoi="10.3897/zookeys.1072.72978.figure19" httpUri="https://binary.pensoft.net/fig/612925" pageId="0" pageNumber="129">19B</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">20A</figureCitation>
) (versus P4 merus long, reaching beyond branchiocardiac groove when folded dorsally in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. martini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="0" pageNumber="129">D. martini</emphasis>
</taxonomicName>
, cf. Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dicranodromia martini Guinot, 1995, ♀ (28.1 v 34.2 mm) (ZRC 2007.0106), Philippines A overall view B right third maxilliped C dorsal view of carapace D anterior part of carapace (right side denuded) E frontal view of cephalothorax." figureDoi="10.3897/zookeys.1072.72978.figure4" httpUri="https://binary.pensoft.net/fig/612909" pageId="0" pageNumber="129">4A</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dicranodromia martini Guinot, 1995, ♀ (28.1 x 34.2 mm) (ZRC 2007.0106), Philippines A epistome, antennules, antennae and orbits B left chela C right P 3 D right P 5 E right P 5 dactylus F right P 4 G telson H posterior thoracic sternum showing spermatheca and female gonopores." figureDoi="10.3897/zookeys.1072.72978.figure5" httpUri="https://binary.pensoft.net/fig/612911" pageId="0" pageNumber="129">5D</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dicranodromia martini Guinot, 1995, ♂ (12.3 x 16.6 mm) (ZRC 2007.0105), Philippines A overall view B dorsal view of carapace C anterior part of carapace (partially denuded) D male telson." figureDoi="10.3897/zookeys.1072.72978.figure6" httpUri="https://binary.pensoft.net/fig/612912" pageId="0" pageNumber="129">6A</figureCitation>
;
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 19a); and the male telson is lingulate (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">20C</figureCitation>
) (versus more elongate in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. martini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="0" pageNumber="129">D. martini</emphasis>
</taxonomicName>
, cf. Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dicranodromia martini Guinot, 1995, ♂ (12.3 x 16.6 mm) (ZRC 2007.0105), Philippines A overall view B dorsal view of carapace C anterior part of carapace (partially denuded) D male telson." figureDoi="10.3897/zookeys.1072.72978.figure6" httpUri="https://binary.pensoft.net/fig/612912" pageId="0" pageNumber="129">6D</figureCitation>
;
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 19c).
</paragraph>
<caption doi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 20.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia robusta</emphasis>
</taxonomicName>
sp. nov., paratype ♂ (15.2
<normalizedToken originalValue="×">x</normalizedToken>
21.0 mm) (ZRC 2018.0095), Philippines
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
overall view
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
dorsal view of carapace
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
pleon
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
anterior thoracic sternum and G1s in situ.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="129">
Compared to
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. baffini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="baffini">
<emphasis italics="true" pageId="0" pageNumber="129">D. baffini</emphasis>
</taxonomicName>
from the Indian Ocean,
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">D. robusta</emphasis>
</taxonomicName>
has a more quadrate carapace (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">17A, B</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">20A, B</figureCitation>
) (versus more pyriform in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. baffini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="baffini">
<emphasis italics="true" pageId="0" pageNumber="129">D. baffini</emphasis>
</taxonomicName>
, cf.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10330" author="Alcock, A" journalOrPublisher="Keelung, Taiwan" pageId="0" pageNumber="129" refId="B3" refString="Alcock, A, 1899. An account of the Deep-Sea Brachyura collected by the Royal Indian marine survey ship INVESTIGATOR. Printed by order of the Trustees of the Indian Museum, Calcutta, 87 pp, pls 1-4. https://doi.org/10.5962/bhl.title.10330" title="An account of the Deep-Sea Brachyura collected by the Royal Indian marine survey ship INVESTIGATOR. Printed by order of the Trustees of the Indian Museum, Calcutta, 87 pp, pls 1 - 4." url="https://doi.org/10.5962/bhl.title.10330" year="1899">Alcock 1899</bibRefCitation>
: pl. 2 fig. 1a;
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 13;
<bibRefCitation DOI="https://doi.org/10.1016/j.rsma.2020.101117" author="Padate, VP" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="129" refId="B32" refString="Padate, VP, Amritha, KM, Cubelio, SS, Saravanane, N, Sudhakar, M, Ng, PKL, 2020. Deep-water Brachyura from the surveys of the FORV Sagar Sampada off the Andaman and Nicobar archipelagos, India. Regional Studies in Marine Science 35: e101117. https://doi.org/10.1016/j.rsma.2020.101117" title="Deep-water Brachyura from the surveys of the FORV Sagar Sampada off the Andaman and Nicobar archipelagos, India. Regional Studies in Marine Science 35: e 101117." url="https://doi.org/10.1016/j.rsma.2020.101117" year="2020">Padate et al. 2020</bibRefCitation>
: fig. 2a); and the P2 and P3 dactylus is distinctly shorter (Figs
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A right P 3 B right P 5 C right P 5 propodus and dactylus D left P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca." figureDoi="10.3897/zookeys.1072.72978.figure19" httpUri="https://binary.pensoft.net/fig/612925" pageId="0" pageNumber="129">19A</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">21G</figureCitation>
) (versus longer in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. baffini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="baffini">
<emphasis italics="true" pageId="0" pageNumber="129">D. baffini</emphasis>
</taxonomicName>
, cf.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10330" author="Alcock, A" journalOrPublisher="Keelung, Taiwan" pageId="0" pageNumber="129" refId="B3" refString="Alcock, A, 1899. An account of the Deep-Sea Brachyura collected by the Royal Indian marine survey ship INVESTIGATOR. Printed by order of the Trustees of the Indian Museum, Calcutta, 87 pp, pls 1-4. https://doi.org/10.5962/bhl.title.10330" title="An account of the Deep-Sea Brachyura collected by the Royal Indian marine survey ship INVESTIGATOR. Printed by order of the Trustees of the Indian Museum, Calcutta, 87 pp, pls 1 - 4." url="https://doi.org/10.5962/bhl.title.10330" year="1899">Alcock 1899</bibRefCitation>
: pl. 2 fig. 1a;
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 13). With regards to the relatively shorter P2 and P3 dactyli,
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">D. robusta</emphasis>
</taxonomicName>
resembles
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. chenae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="chenae">
<emphasis italics="true" pageId="0" pageNumber="129">D. chenae</emphasis>
</taxonomicName>
, described from a single large ovigerous female from the central Philippines.
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia robusta</emphasis>
</taxonomicName>
, however, can easily be distinguished in having the outer margin of the pseudorostral lobe is almost straight and the structure is directed anteriorly (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">17B-D</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">20B</figureCitation>
) (versus outer margin of the pseudorostral lobe is distinctly convex with the structure gradually curved inwards towards the median in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. chenae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="chenae">
<emphasis italics="true" pageId="0" pageNumber="129">D. chenae</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="129" pagination="47 - 53" refId="B28" refString="Ng, PKL, Naruse, T, 2007. On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species. Raffles Bulletin of Zoology, Supplement 16: 47 - 53" title="On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species." volume="16" year="2007">Ng and Naruse 2007</bibRefCitation>
: fig. 5A); the ischium of the third maxilliped is short and rectangular (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of left chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure18" httpUri="https://binary.pensoft.net/fig/612924" pageId="0" pageNumber="129">18B</figureCitation>
) (versus distinctly longer and more slender in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. chenae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="chenae">
<emphasis italics="true" pageId="0" pageNumber="129">D. chenae</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="129" pagination="47 - 53" refId="B28" refString="Ng, PKL, Naruse, T, 2007. On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species. Raffles Bulletin of Zoology, Supplement 16: 47 - 53" title="On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species." volume="16" year="2007">Ng and Naruse 2007</bibRefCitation>
: fig. 5b); the female telson is relatively more elongate (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of left chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure18" httpUri="https://binary.pensoft.net/fig/612924" pageId="0" pageNumber="129">18A</figureCitation>
) (versus proportionately wider and shorter in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. chenae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="chenae">
<emphasis italics="true" pageId="0" pageNumber="129">D. chenae</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="129" pagination="47 - 53" refId="B28" refString="Ng, PKL, Naruse, T, 2007. On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species. Raffles Bulletin of Zoology, Supplement 16: 47 - 53" title="On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species." volume="16" year="2007">Ng and Naruse 2007</bibRefCitation>
: fig. 2b); and the spermatheca is on the prominently raised part around the suture between sternites 7 and 8 and ends at the centre of the triangular tubercle on sternite 6 (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A right P 3 B right P 5 C right P 5 propodus and dactylus D left P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca." figureDoi="10.3897/zookeys.1072.72978.figure19" httpUri="https://binary.pensoft.net/fig/612925" pageId="0" pageNumber="129">19F</figureCitation>
) (versus spermatheca is not prominently raised and ends at the base of the triangular tubercle in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. chenae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="chenae">
<emphasis italics="true" pageId="0" pageNumber="129">D. chenae</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="129" pagination="47 - 53" refId="B28" refString="Ng, PKL, Naruse, T, 2007. On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species. Raffles Bulletin of Zoology, Supplement 16: 47 - 53" title="On two species of deep sea homoloid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, with description of a new species." volume="16" year="2007">Ng and Naruse 2007</bibRefCitation>
: fig. 8).
</paragraph>
<caption doi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 21.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia erinaceus</emphasis>
</taxonomicName>
sp. nov., holotype ♀ (14.0
<normalizedToken originalValue="×">x</normalizedToken>
18.0 mm) (NTOU B00126), Taiwan
<emphasis bold="true" pageId="0" pageNumber="129">B, D-F, K-M</emphasis>
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">D. erinaceus</emphasis>
</taxonomicName>
sp. nov., paratype ♂ (8.2
<normalizedToken originalValue="×">x</normalizedToken>
12.5 mm) (ZRC 2021.0085), Taiwan
<emphasis bold="true" pageId="0" pageNumber="129">C, G-J, N-P</emphasis>
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">D. robusta</emphasis>
</taxonomicName>
sp. nov., paratype ♂ (15.2
<normalizedToken originalValue="×">x</normalizedToken>
21.0 mm) (ZRC 2018.0095), Philippines
<emphasis bold="true" pageId="0" pageNumber="129">A-C</emphasis>
right anterior part of carapace (setae removed or not drawn)
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
right P3 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
right P5 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
right P4 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">G</emphasis>
right P3 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">H</emphasis>
right P5 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">I</emphasis>
left P5 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">J</emphasis>
left P4 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">K, N</emphasis>
left G1 (ventral view)
<emphasis bold="true" pageId="0" pageNumber="129">L, O</emphasis>
left G1 (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="129">M, P</emphasis>
left G2. Setae for all structures not figured. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia robusta" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="robusta">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia robusta</emphasis>
</taxonomicName>
can be separated from
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. danielae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="danielae">
<emphasis italics="true" pageId="0" pageNumber="129">D. danielae</emphasis>
</taxonomicName>
in having the exorbital tooth distinctly triangular to linguiform (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">17B-D</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">21C</figureCitation>
) (versus subtrapezoidal in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. danielae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="danielae">
<emphasis italics="true" pageId="0" pageNumber="129">D. danielae</emphasis>
</taxonomicName>
, cf. Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Dicranodromia danielae Ng &amp; McLay, 2005, holotype ovigerous ♀ (broken, 10.8 x 14.2 mm) (ZRC 2005.0094), Philippines A dorsal view of carapace B frontal region showing rostrum C epistome, antennules, antennae and orbits D right chela E right P 3 F left P 5 G left P 5 propodus and dactylus H left P 4 propodus and dactylus I right third maxilliped." figureDoi="10.3897/zookeys.1072.72978.figure12" httpUri="https://binary.pensoft.net/fig/612918" pageId="0" pageNumber="129">12A, B</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1029.1.3" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="39 - 46" refId="B26" refString="Ng, PKL, McLay, CL, 2005. Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura). Zootaxa 1029: 39 - 46, DOI: https://doi.org/10.11646/zootaxa.1029.1.3" title="Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1029.1.3" volume="1029" year="2005">Ng and McLay 2005</bibRefCitation>
: figs 1B, 4A); the posterior margin of the epistome is entire (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of left chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure18" httpUri="https://binary.pensoft.net/fig/612924" pageId="0" pageNumber="129">18C</figureCitation>
) (versus clearly serrate in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. danielae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="danielae">
<emphasis italics="true" pageId="0" pageNumber="129">D. danielae</emphasis>
</taxonomicName>
, cf. Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Dicranodromia danielae Ng &amp; McLay, 2005, holotype ovigerous ♀ (broken, 10.8 x 14.2 mm) (ZRC 2005.0094), Philippines A dorsal view of carapace B frontal region showing rostrum C epistome, antennules, antennae and orbits D right chela E right P 3 F left P 5 G left P 5 propodus and dactylus H left P 4 propodus and dactylus I right third maxilliped." figureDoi="10.3897/zookeys.1072.72978.figure12" httpUri="https://binary.pensoft.net/fig/612918" pageId="0" pageNumber="129">12C</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1029.1.3" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="39 - 46" refId="B26" refString="Ng, PKL, McLay, CL, 2005. Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura). Zootaxa 1029: 39 - 46, DOI: https://doi.org/10.11646/zootaxa.1029.1.3" title="Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1029.1.3" volume="1029" year="2005">Ng and McLay 2005</bibRefCitation>
: fig. 4C); the median part of the outer surface of the chela is granular (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of left chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure18" httpUri="https://binary.pensoft.net/fig/612924" pageId="0" pageNumber="129">18E, F</figureCitation>
) (versus smooth in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. danielae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="danielae">
<emphasis italics="true" pageId="0" pageNumber="129">D. danielae</emphasis>
</taxonomicName>
, cf. Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Dicranodromia danielae Ng &amp; McLay, 2005, holotype ovigerous ♀ (broken, 10.8 x 14.2 mm) (ZRC 2005.0094), Philippines A dorsal view of carapace B frontal region showing rostrum C epistome, antennules, antennae and orbits D right chela E right P 3 F left P 5 G left P 5 propodus and dactylus H left P 4 propodus and dactylus I right third maxilliped." figureDoi="10.3897/zookeys.1072.72978.figure12" httpUri="https://binary.pensoft.net/fig/612918" pageId="0" pageNumber="129">12D</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1029.1.3" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="39 - 46" refId="B26" refString="Ng, PKL, McLay, CL, 2005. Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura). Zootaxa 1029: 39 - 46, DOI: https://doi.org/10.11646/zootaxa.1029.1.3" title="Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1029.1.3" volume="1029" year="2005">Ng and McLay 2005</bibRefCitation>
: fig. 3A, B); and P2-P5 are all proportionately longer with the flexor margins of the meri not spinate (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A overall view B dorsal view of carapace C front and anterior part of carapace D right front, orbit and first anterolateral spine" figureDoi="10.3897/zookeys.1072.72978.figure17" httpUri="https://binary.pensoft.net/fig/612923" pageId="0" pageNumber="129">17A</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Dicranodromia robusta sp. nov., holotype ♀ (19.6 x 26.4 mm) (ZRC 2018.0161), Philippines A right P 3 B right P 5 C right P 5 propodus and dactylus D left P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca." figureDoi="10.3897/zookeys.1072.72978.figure19" httpUri="https://binary.pensoft.net/fig/612925" pageId="0" pageNumber="129">19A</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Dicranodromia robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure20" httpUri="https://binary.pensoft.net/fig/612926" pageId="0" pageNumber="129">20A</figureCitation>
) (versus relatively shorter in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. danielae" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="danielae">
<emphasis italics="true" pageId="0" pageNumber="129">D. danielae</emphasis>
</taxonomicName>
with the meri of P2 and P3 distinctly spinate, cf. Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Dicranodromia danielae Ng &amp; McLay, 2005, holotype ovigerous ♀ (broken, 10.8 x 14.2 mm) (ZRC 2005.0094), Philippines A dorsal view of carapace B frontal region showing rostrum C epistome, antennules, antennae and orbits D right chela E right P 3 F left P 5 G left P 5 propodus and dactylus H left P 4 propodus and dactylus I right third maxilliped." figureDoi="10.3897/zookeys.1072.72978.figure12" httpUri="https://binary.pensoft.net/fig/612918" pageId="0" pageNumber="129">12E, F</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1029.1.3" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="39 - 46" refId="B26" refString="Ng, PKL, McLay, CL, 2005. Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura). Zootaxa 1029: 39 - 46, DOI: https://doi.org/10.11646/zootaxa.1029.1.3" title="Dicranodromia danielae, a new species of homolodromiid crab from the Philippines (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1029.1.3" volume="1029" year="2005">Ng and McLay 2005</bibRefCitation>
: fig. 1A).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>