193 lines
20 KiB
XML
193 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.241.3089" ID-GBIF-Dataset="176d5f10-f453-4868-b0db-71b9655bbb3f" ID-PMC="PMC3559106" ID-Pensoft-Pub="1313-2970-241-55" ID-PubMed="23372413" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-241-55" ModsDocOrigin="ZooKeys 241" ModsDocTitle="A revision of the Cautires obsoletus species group from Java (Coleoptera, Lycidae)" checkinTime="1451248561069" checkinUser="pensoft" docAuthor="Bocak, Ladislav" docDate="2012" docId="D6A989A31AD9CD6A14144746EE0B27EB" docLanguage="en" docName="ZooKeys 241: 55-66" docOrigin="ZooKeys 241" docSource="http://dx.doi.org/10.3897/zookeys.241.3089" docTitle="Cautires obsoletus Waterhouse 1878" docType="treatment" docVersion="3" lastPageNumber="59" masterDocId="8863FFCD2F20FF8EFF895F12FFBBFFEE" masterDocTitle="A revision of the Cautires obsoletus species group from Java (Coleoptera, Lycidae)" masterLastPageNumber="66" masterPageNumber="55" pageNumber="58" updateTime="1668154707921" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the Cautires obsoletus species group from Java (Coleoptera, Lycidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bocak, Ladislav</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>241</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>55</mods:start>
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<mods:end>66</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.241.3089</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.241.3089</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-241-55</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152037879" LSID="urn:lsid:plazi:treatment:D6A989A31AD9CD6A14144746EE0B27EB" httpUri="http://treatment.plazi.org/id/D6A989A31AD9CD6A14144746EE0B27EB" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
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<subSubSection pageId="3" pageNumber="58" type="nomenclature">
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName LSID="http://species-id.net/wiki/Cautires_obsoletus" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires obsoletus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="obsoletus">Cautires obsoletus (Waterhouse, 1878)</taxonomicName>
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Figs 6, 8, 12-13, 20
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="58" type="reference_group">
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides obsoletus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="obsoletus">Bulenides obsoletus</taxonomicName>
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<bibRefCitation author="Waterhouse, CO" journalOrPublisher="Transactions of the Entomological Society London" pageId="6" pageNumber="61" pagination="95 - 118" title="On the different forms occurring in the Coleopterous family Lycidae, with description of new genera and species." volume="1878" year="1878">Waterhouse 1878</bibRefCitation>
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: 109.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires obsoletus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="obsoletus">Cautires obsoletus</taxonomicName>
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(
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<bibRefCitation author="Waterhouse, CO" journalOrPublisher="Transactions of the Entomological Society London" pageId="6" pageNumber="61" pagination="95 - 118" title="On the different forms occurring in the Coleopterous family Lycidae, with description of new genera and species." volume="1878" year="1878">Waterhouse 1878</bibRefCitation>
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):
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<bibRefCitation author="Dudkova, P" journalOrPublisher="Zootaxa" pageId="6" pageNumber="61" pagination="28 - 48" title="A review of the Cautires obsoletus species group from Indo-Burma (Coleoptera: Lycidae)." volume="2527" year="2010">Dudkova and Bocak 2010</bibRefCitation>
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: 43.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides lineatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="lineatus">Bulenides lineatus</taxonomicName>
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<bibRefCitation pageId="3" pageNumber="58">Pic 1921</bibRefCitation>
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: 8.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires lineatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="lineatus">Cautires lineatus</taxonomicName>
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(
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<bibRefCitation pageId="3" pageNumber="58">Pic 1921</bibRefCitation>
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): Dudkova and Bocak: 42 (a junior secondary homonym of
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires lineatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="lineatus">Cautires lineatus</taxonomicName>
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(Hope in Gray 1831), syn. n.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires fruhstorferi" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="fruhstorferi">Cautires fruhstorferi</taxonomicName>
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Bocak and Dudkova 2010: 42 (a replacement name for
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides lineatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="lineatus">Bulenides lineatus</taxonomicName>
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), syn. n.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides javanicus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="javanicus">Bulenides javanicus</taxonomicName>
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<bibRefCitation author="Bourgeois, JM" journalOrPublisher="Annali del Museo civico di storia naturale Genova" pageId="6" pageNumber="61" pagination="640 - 666" title="Lycidae nouveaux ou peu connus." volume="43" year="1883">Bourgeois 1883</bibRefCitation>
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: 439.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires javanicus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="javanicus">Cautires javanicus</taxonomicName>
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(
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<bibRefCitation author="Bourgeois, JM" journalOrPublisher="Annali del Museo civico di storia naturale Genova" pageId="6" pageNumber="61" pagination="640 - 666" title="Lycidae nouveaux ou peu connus." volume="43" year="1883">Bourgeois 1883</bibRefCitation>
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):
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<bibRefCitation author="Dudkova, P" journalOrPublisher="Zootaxa" pageId="6" pageNumber="61" pagination="28 - 48" title="A review of the Cautires obsoletus species group from Indo-Burma (Coleoptera: Lycidae)." volume="2527" year="2010">Dudkova and Bocak 2010</bibRefCitation>
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: 42, syn. n.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides inhumeralis" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="inhumeralis">Bulenides inhumeralis</taxonomicName>
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<bibRefCitation pageId="3" pageNumber="58">Pic 1921</bibRefCitation>
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: 7.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires inhumeralis" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="inhumeralis">Cautires inhumeralis</taxonomicName>
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(
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<bibRefCitation pageId="3" pageNumber="58">Pic 1921</bibRefCitation>
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):
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<bibRefCitation author="Dudkova, P" journalOrPublisher="Zootaxa" pageId="6" pageNumber="61" pagination="28 - 48" title="A review of the Cautires obsoletus species group from Indo-Burma (Coleoptera: Lycidae)." volume="2527" year="2010">Dudkova and Bocak 2010</bibRefCitation>
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: 42, syn. n.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides nigromaculatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="nigromaculatus">Bulenides nigromaculatus</taxonomicName>
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<bibRefCitation author="Pic, M" journalOrPublisher="L'Echange" pageId="6" pageNumber="61" pagination="5 - 8" title="Malacodermes exotiques." volume="419" year="1925 a">Pic 1925a</bibRefCitation>
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: 7.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires nigromaculatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="nigromaculatus">Cautires nigromaculatus</taxonomicName>
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(
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<bibRefCitation author="Pic, M" journalOrPublisher="L'Echange" pageId="6" pageNumber="61" pagination="5 - 8" title="Malacodermes exotiques." volume="419" year="1925 a">Pic 1925a</bibRefCitation>
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):
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<bibRefCitation author="Dudkova, P" journalOrPublisher="Zootaxa" pageId="6" pageNumber="61" pagination="28 - 48" title="A review of the Cautires obsoletus species group from Indo-Burma (Coleoptera: Lycidae)." volume="2527" year="2010">Dudkova and Bocak 2010</bibRefCitation>
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: 43, syn. n.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides pudicus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="pudicus">Bulenides pudicus</taxonomicName>
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<bibRefCitation author="Kleine, R" journalOrPublisher="Treubia" pageId="6" pageNumber="61" pagination="245 - 290" title="Zur Kenntnis der Lycidenfauna Javas." volume="9" year="1931">Kleine 1931</bibRefCitation>
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: 257.
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</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires pudicus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="pudicus">Cautires pudicus</taxonomicName>
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(
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<bibRefCitation author="Kleine, R" journalOrPublisher="Treubia" pageId="6" pageNumber="61" pagination="245 - 290" title="Zur Kenntnis der Lycidenfauna Javas." volume="9" year="1931">Kleine 1931</bibRefCitation>
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):
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<bibRefCitation author="Dudkova, P" journalOrPublisher="Zootaxa" pageId="6" pageNumber="61" pagination="28 - 48" title="A review of the Cautires obsoletus species group from Indo-Burma (Coleoptera: Lycidae)." volume="2527" year="2010">Dudkova and Bocak 2010</bibRefCitation>
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: 43, syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="58" type="type material">
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<paragraph pageId="3" pageNumber="58">Type material.</paragraph>
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<paragraph pageId="3" pageNumber="58">
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Female, holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides obsoletus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="obsoletus">Bulenides obsoletus</taxonomicName>
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. [Indonesia] Java (without further data, BMNH). Female, holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides lineatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="lineatus">Bulenides lineatus</taxonomicName>
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. [Indonesia] Java occident., Sukabumi, 2000', 1893, H. Fruhstorfer (MHNP). Male, holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides javanicus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="javanicus">Bulenides javanicus</taxonomicName>
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. [Indonesia, Java] Giava, Tcibodas, Ott. 1874, O. Beccari (MHNP). Male, holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides inhumeralis" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="inhumeralis">Bulenides inhumeralis</taxonomicName>
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. [Indonesia] Bogor, 1000',
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<normalizedToken originalValue="v–vi">v-vi</normalizedToken>
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96, I. Z. Kannegieter (MHNP). Male, holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides nigromaculatus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="nigromaculatus">Bulenides nigromaculatus</taxonomicName>
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. [Indonesia] Java occident., Sukabumi 2000', H. Fruhstorfer (MHNP). Male, holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bulenides pudicus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="pudicus">Bulenides pudicus</taxonomicName>
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. [Indonesia] Toegoe, West-Jawa-Pasteur (without further data, MIZW).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="58" type="differential diagnosis">
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<paragraph pageId="3" pageNumber="58">Differential diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="58">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires obsoletus" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="obsoletus">Cautires obsoletus</taxonomicName>
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differs from the other Javanese species in the black pronotum (Fig. 8) and a very oblique border between the bright and dark parts of elytra (Fig. 6). The basal part of elytra is brown to reddish brown and differs from the brightly orange coloration of the other species in Java.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58" type="redescription">
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<paragraph pageId="3" pageNumber="58">Redescription</paragraph>
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<paragraph lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
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.Male. Body small to medium-sized, dorso-ventrally flattened, slender; head, body, posterior half of elytra and appendages dark-brown to black; basal half of elytra brown to reddish brown (Fig. 6), body covered with dense pubescence. Head small, partly hidden in pronotum, clypeus slightly concave, labrum simply rounded, mandibles slender, strongly curved, maxillary palpi slender, apical palpomere pointed; labial palpi similar in shape. Eyes small, hemispherically prominent, their frontal interocular distance 1.15 times eye diameter. Antennae shortly flabellate, 11-segmented, covered with short, dark colored setae (Fig. 20). Pronotum flat, only slightly transverse, 1.15 times wider at base than length at midline; frontal margin projected forward; lateral margins slightly elevated, concave, posterior angles acutely projected, pronotum with moderately wide median areola, attached to middle of basal margin of pronotum, connected to anterior
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<pageBreakToken pageId="4" pageNumber="59" start="start">margin</pageBreakToken>
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by keel occupying one third of midline, lateral keels absent (Fig. 8). Elytra flat, with well developed four primary longitudinal costae; secondary costae considerably weaker, cells regular, mostly slightly longitudinal. Legs laterally flattened, covered with dark colored setae. Male genitalia with phallus widest at midlength and gradually narrowed to apex (Figs 12-13). Female. Body medium-sized, antennae serrate, ovipositor with short valvifers.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="59" type="measurements">
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<paragraph pageId="4" pageNumber="59">Measurements.</paragraph>
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<paragraph pageId="4" pageNumber="59">BL 6.45 mm, PL 1.01 mm, PW 1.36 mm, HW 1.47 mm, Edist 0.51 mm, Ediam 0.44 mm.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="59" type="distribution">
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<paragraph pageId="4" pageNumber="59">Distribution.</paragraph>
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<paragraph pageId="4" pageNumber="59">
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires obsoletus" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="obsoletus">Cautires obsoletus</taxonomicName>
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is known only from several localities in Western Java.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="59" type="material examined">
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<paragraph pageId="4" pageNumber="59">Material examined.</paragraph>
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<paragraph pageId="4" pageNumber="59">2 males, 1 female, Indonesia, W. Java, Puncak Pass nr Bogor, 23. Mar. 1992, H. Arimoto, lgt.; male, [Indonesia] West Java, Puncak Pass, 22. Mar. 1993, Y. Miyake leg. (KMTC, LMBC).</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="59" type="remark">
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<paragraph pageId="4" pageNumber="59">Remark.</paragraph>
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<paragraph pageId="4" pageNumber="59">
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The holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires obsoletus" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="obsoletus">Cautires obsoletus</taxonomicName>
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is a strongly damaged female with only a part of one elytron preserved. The basal part of the elytron is testaceous and the loss of reddish coloration may be caused by long-term exposure to light. The holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires lineatus" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="lineatus">Cautires lineatus</taxonomicName>
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is also a female and it differs only in the darker hue of the bright part of the elytra, but it resembles the holotype of
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires obsoletus" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="obsoletus">Cautires obsoletus</taxonomicName>
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in its body shape. The female genitalia of all available specimens are very similar.Holotypes of four other species are males and they do not differ in comparable characters such as body shape and color pattern. Therefore, all these species are considered junior subjective synonyms of
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<taxonomicName class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cautires obsoletus" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="obsoletus">Cautires obsoletus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |