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<document id="65686F4359B8EAB6DC848DE7AAC20B5A" ID-CLB-Dataset="131926" ID-DOI="10.11646/zootaxa.5178.4.4" ID-GBIF-Dataset="69f03f3e-68ec-43c9-a3d3-75927165afae" ID-ISSN="1175-5326" ID-Zenodo-Dep="7031814" ID-ZooBank="60009AB8-4DAC-418E-8378-9155DDDD024B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661769161681" checkinUser="plazi" docAuthor="Nakamine, Hiroshi, Yamamoto, Shûhei &amp; Takahashi, Yui" docDate="2022" docId="D679951B82364C60FF765F29E5E0CAEF" docLanguage="en" docName="zootaxa.5178.4.4.pdf" docOrigin="Zootaxa 5178 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaeomegalomus burmiticus Nakamine &amp; Yamamoto &amp; Takahashi 2022, sp. nov." docType="treatment" docVersion="10" lastPageNumber="386" masterDocId="2A40ED6382354C66FFE15A4CE77CC81E" masterDocTitle="Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae)" masterLastPageNumber="390" masterPageNumber="380" pageNumber="383" updateTime="1699374251715" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4EEE5FD2CBA60F9B0C8E279ED0F8E71F">Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae)</mods:title>
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<mods:namePart id="973DA1979B4F6954A4E4414CAF631EAA">Nakamine, Hiroshi</mods:namePart>
<mods:affiliation id="598C59EE2DC3ECF85045571FE4D8DF14">Minoh Park Insect Museum, Minoh Park 1 - 18, Minoh City, Osaka 562 - 0002, Japan.</mods:affiliation>
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<mods:namePart id="9AC26FC05C78462872C5295EFDCFB458">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="9A34267FD9ECD868429E1AFE68B6D5B6">Hokkaio University Museum, Hokkaido University, Kita 10 - jo, Nishi 8 - chome, Kita-ku, Sapporo 060 - 0810, Japan.</mods:affiliation>
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<mods:namePart id="635AB03846105D1F7DF73630964D5824">Takahashi, Yui</mods:namePart>
<mods:affiliation id="5115D22004B76A8B566D66EA0B9F3EA2">Keio Yochisha Elementary School, Ebisu 2 - 35 - 1, Shibuya-ku, Tokyo 150 - 0013, Japan.</mods:affiliation>
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<treatment id="D679951B82364C60FF765F29E5E0CAEF" ID-DOI="http://doi.org/10.5281/zenodo.7037080" ID-GBIF-Taxon="199861244" ID-Zenodo-Dep="7037080" LSID="urn:lsid:plazi:treatment:D679951B82364C60FF765F29E5E0CAEF" httpUri="http://treatment.plazi.org/id/D679951B82364C60FF765F29E5E0CAEF" lastPageId="6" lastPageNumber="386" pageId="3" pageNumber="383">
<subSubSection id="16CA778682364C65FF765F29E50BCD61" box="[151,631,1381,1408]" pageId="3" pageNumber="383" type="nomenclature">
<paragraph id="5E6F240D82364C65FF765F29E50BCD61" blockId="3.[151,631,1381,1444]" box="[151,631,1381,1408]" pageId="3" pageNumber="383">
<heading id="0527936182364C65FF765F29E50BCD61" bold="true" box="[151,631,1381,1408]" fontSize="11" level="1" pageId="3" pageNumber="383" reason="1">
<emphasis id="6CA4F81F82364C65FF765F29E50BCD61" bold="true" box="[151,631,1381,1408]" pageId="3" pageNumber="383">
<taxonomicName id="99D05F8E82364C65FF765F29E56ECD9E" authority="Nakamine &amp; Yamamoto &amp; Takahashi, 2022" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi" authorityYear="2022" box="[151,530,1381,1408]" class="Insecta" family="Hemerobiidae" genus="Archaeomegalomus" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="383" phylum="Arthropoda" rank="species" species="burmiticus" status="sp. nov.">
<emphasis id="6CA4F81F82364C65FF765F29E56ECD9E" bold="true" box="[151,530,1381,1408]" italics="true" pageId="3" pageNumber="383">Archaeomegalomus burmiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7797456482364C65FDF85F29E50BCD61" box="[537,631,1381,1407]" pageId="3" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="16CA778682364C65FF765FC5E652CDBA" box="[151,302,1417,1444]" pageId="3" pageNumber="383" type="description">
<paragraph id="5E6F240D82364C65FF765FC5E652CDBA" blockId="3.[151,631,1381,1444]" box="[151,302,1417,1444]" pageId="3" pageNumber="383">
(
<figureCitation id="C6EB388882364C65FF7E5FC5E677CDBD" box="[159,267,1417,1443]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,951,975]" captionStartId-2="4.[151,250,1650,1674]" captionTargetBox-0="[216,1371,181,1919]" captionTargetBox-1="[151,1436,181,927]" captionTargetBox-2="[151,1436,181,1626]" captionTargetId-0="figure-18@2.[216,1371,181,1919]" captionTargetId-1="figure-21@3.[151,1436,181,927]" captionTargetId-2="figure-18@4.[151,1436,181,1626]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Habitus and wing detail of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid- Cretaceous Kachin amber. A, left lateral view; B, right lateral view. Scale bar: 1.0 mm." captionText-1="FIGURE 2. Wing venation of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, right forewing; B, left forewing; C, right hind wing; D, left hind wing. Abbreviations: A1A3: first to third anal vein; CuA: cubitus anterior; CuP: cubitus posterior; hv: humeral vein; MA: media anterior; MP: media posterior; RA: radius anterior; RP: radius posterior; Sc: subcosta. Scale bar: 1.0 mm." captionText-2="FIGURE 3. Details of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, Sc and RA on right forewing; B, Sc and RA on left hind wing; C, right lateral view of head and thorax; D, dorsal view of head and thorax; E, left metatibia and tarsus; F, right lateral view of terminalia.Abbreviations: ect: ectoproct; gx9: gonocoxite 9; hp: humeral plate; hv: humeral veinlet; lp: labial palpus; mp: maxillary palpus; mtti: metatibia; RA: radius anterior; Sc: subcosta; st: gonostylus of gonocoxite 9; ts: tibial spurs. Scale bars: 0.1 mm for A and B; 0.5 mm for CE; 0.2 mm for F." figureDoi-0="http://doi.org/10.5281/zenodo.7031816" figureDoi-1="http://doi.org/10.5281/zenodo.7031819" figureDoi-2="http://doi.org/10.5281/zenodo.7031821" httpUri-0="https://zenodo.org/record/7031816/files/figure.png" httpUri-1="https://zenodo.org/record/7031819/files/figure.png" httpUri-2="https://zenodo.org/record/7031821/files/figure.png" pageId="3" pageNumber="383">Figs. 13</figureCitation>
,
<figureCitation id="C6EB388882364C65FEF65FC5E659CDBA" box="[279,293,1417,1444]" captionStart="FIGURE 5" captionStartId="7.[151,250,1873,1897]" captionTargetBox="[151,1436,381,1850]" captionTargetId="figure-120@7.[151,1436,381,1850]" captionTargetPageId="7" captionText="FIGURE 5. Distribution map and forewing venation of Mesozoic Hemerobiidae. A, distribution map of Mesozoic Hemerobiidae; B, Promegalomus anomalus (modified after Panfilov 1980, fig. 91); C, Cretomerobius disjunctus (modified after Ponomarenko 1992, fig. 4); D, Cretoneuronema jarzembowskii (modified after Liu et al. 2022, fig. 3); E, Archaeomegalomus burmiticus gen. et sp. nov.; F, Hemeroberotha sinefurca (modified after Makarkin &amp; Gröhn 2020, fig. 4); G, Plesiorobius canadensis (modified after Klimaszewski &amp; Kevan 1986, fig. 2)." figureDoi="http://doi.org/10.5281/zenodo.7031825" httpUri="https://zenodo.org/record/7031825/files/figure.png" pageId="3" pageNumber="383">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="16CA778682364C65FF765F9DE519CDF5" box="[151,613,1489,1516]" pageId="3" pageNumber="383" type="diagnosis">
<paragraph id="5E6F240D82364C65FF765F9DE519CDF5" blockId="3.[151,1437,1489,2020]" box="[151,613,1489,1516]" pageId="3" pageNumber="383">
<emphasis id="6CA4F81F82364C65FF765F9DE66ECDF5" bold="true" box="[151,274,1489,1515]" pageId="3" pageNumber="383">Diagnosis.</emphasis>
As for the genus (
<emphasis id="6CA4F81F82364C65FE3E5F9DE529CDF5" box="[479,597,1489,1515]" italics="true" pageId="3" pageNumber="383">vide supra</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="16CA778682364C60FF265FB9E363C95F" lastPageId="6" lastPageNumber="386" pageId="3" pageNumber="383" type="description">
<paragraph id="5E6F240D82364C65FF265FB9E24ECE11" blockId="3.[151,1437,1489,2020]" box="[199,1330,1525,1552]" pageId="3" pageNumber="383">
<emphasis id="6CA4F81F82364C65FF265FB9E625CE11" bold="true" box="[199,345,1525,1551]" pageId="3" pageNumber="383">Description.</emphasis>
Body. Length ca.
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as preserved (measured from vertex to apex of the abdomen).
</paragraph>
<paragraph id="5E6F240D82364C65FF265C55E2F7CE65" blockId="3.[151,1437,1489,2020]" pageId="3" pageNumber="383">Head. Eyes large, protruding laterally. Antennae: scapus relatively short, ca. 2 times as long as wide; pedicellus one-half length of scape; both scattered with thin setae; flagellum moniliform, relatively long, composed of 38 flagellomeres, with scattered fine setae on each segment. Maxillary palpi 4 or 5 segmented; labial palpi 3 segmented.</paragraph>
<paragraph id="5E6F240D82364C65FF265CC9E63ECEDA" blockId="3.[151,1437,1489,2020]" pageId="3" pageNumber="383">Thorax. Pronotum rather short, poorly preserved, covered with thin setae. Mesothorax and metathorax covered with thin setae.</paragraph>
<paragraph id="5E6F240D82364C65FF265C82E218CFFD" blockId="3.[151,1437,1489,2020]" pageId="3" pageNumber="383">
Legs. Foreleg slender; procoxa poorly preserved, relatively long; profemur ca.
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long, scarce setae; protibia ca.
<quantity id="992889E882364C65FF1B5CBDE62DCF12" box="[250,337,1777,1804]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="383" unit="mm" value="0.5">0.5 mm</quantity>
long, slightly swollen at the middle, covered with dense thin setae, paired tibial spurs; protarsus five segmented, all tarsomeres covered with thin setae, pretarsal claws thin, small. Midleg slender; mesocoxa short; mesofemur ca.
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long, scarce setae; mesotibia ca.
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long, only slightly swollen at the middle to apex, covered with dense thin setae, paired tibial spurs; mesotarsus five segmented, all tarsomeres covered with thin setae, pretarsal claws thin, small. Hindleg slender; metacoxa short; metafemur ca.
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long, scarce setae; metatibia ca.
<quantity id="992889E882364C65FF765DE9E790CFDE" box="[151,236,1957,1984]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="383" unit="mm" value="1.5">1.5 mm</quantity>
long, slightly curved, only slightly swollen at the middle to apex, covered with dense thin setae, paired tibial spurs (
<figureCitation id="C6EB388882364C65FF035D85E645CFFD" box="[226,313,1993,2019]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Details of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, Sc and RA on right forewing; B, Sc and RA on left hind wing; C, right lateral view of head and thorax; D, dorsal view of head and thorax; E, left metatibia and tarsus; F, right lateral view of terminalia.Abbreviations: ect: ectoproct; gx9: gonocoxite 9; hp: humeral plate; hv: humeral veinlet; lp: labial palpus; mp: maxillary palpus; mtti: metatibia; RA: radius anterior; Sc: subcosta; st: gonostylus of gonocoxite 9; ts: tibial spurs. Scale bars: 0.1 mm for A and B; 0.5 mm for CE; 0.2 mm for F." figureDoi="http://doi.org/10.5281/zenodo.7031821" httpUri="https://zenodo.org/record/7031821/files/figure.png" pageId="3" pageNumber="383">Fig. 3E</figureCitation>
); metatarsus five segmented, all tarsomeres covered with thin setae, pretarsal claws thin, small.
</paragraph>
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<paragraph id="5E6F240D82314C62FF765C3EE246CF04" blockId="4.[151,1437,1650,1819]" pageId="4" pageNumber="384">
<emphasis id="6CA4F81F82314C62FF765C3EE66ECE94" bold="true" box="[151,274,1650,1674]" pageId="4" pageNumber="384">FIGURE 3.</emphasis>
Details of
<taxonomicName id="99D05F8E82314C62FE605C3EE5C0CE94" authority="Nakamine &amp; Yamamoto &amp; Takahashi, 2022" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi" authorityYear="2022" box="[385,700,1650,1674]" class="Insecta" family="Hemerobiidae" genus="Archaeomegalomus" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="384" phylum="Arthropoda" rank="species" species="burmiticus" status="gen. et sp. nov.">
<emphasis id="6CA4F81F82314C62FE605C3EE5C0CE94" box="[385,700,1650,1674]" italics="true" pageId="4" pageNumber="384">Archaeomegalomus burmiticus</emphasis>
</taxonomicName>
<emphasis id="6CA4F81F82314C62FD235C3FE422CE95" bold="true" box="[706,862,1651,1675]" pageId="4" pageNumber="384">
<taxonomicNameLabel id="7797456482314C62FD235C3FE422CE95" box="[706,862,1651,1675]" pageId="4" pageNumber="384" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, Sc and RA on right forewing; B, Sc and RA on left hind wing; C, right lateral view of head and thorax; D, dorsal view of head and thorax; E, left metatibia and tarsus; F, right lateral view of terminalia. Abbreviations: ect: ectoproct; gx9: gonocoxite 9; hp: humeral plate; hv: humeral veinlet; lp: labial palpus; mp: maxillary palpus; mtti: metatibia; RA: radius anterior; Sc: subcosta; st: gonostylus of gonocoxite 9; ts: tibial spurs. Scale bars: 0.1 mm for A and B; 0.5 mm for CE; 0.2 mm for F.
</paragraph>
</caption>
<paragraph id="5E6F240D82314C63FF265D05E458CA23" blockId="4.[151,1437,1864,2035]" lastBlockId="5.[151,1436,151,573]" lastPageId="5" lastPageNumber="385" pageId="4" pageNumber="384">
Wings. Forewing hyaline (
<figureCitation id="C6EB388882314C62FE1D5D05E500CF7D" box="[508,636,1865,1891]" captionStart="FIGURE 2" captionStartId="3.[151,250,951,975]" captionTargetBox="[151,1436,181,927]" captionTargetId="figure-21@3.[151,1436,181,927]" captionTargetPageId="3" captionText="FIGURE 2. Wing venation of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, right forewing; B, left forewing; C, right hind wing; D, left hind wing. Abbreviations: A1A3: first to third anal vein; CuA: cubitus anterior; CuP: cubitus posterior; hv: humeral vein; MA: media anterior; MP: media posterior; RA: radius anterior; RP: radius posterior; Sc: subcosta. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7031819" httpUri="https://zenodo.org/record/7031819/files/figure.png" pageId="4" pageNumber="384">Fig. 2A, B</figureCitation>
), oval with subacute apex, ca.
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long, ca.
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wide (right forewing); ca.
<quantity id="992889E882314C62FEA05D20E6E6CF99" box="[321,410,1900,1927]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="384" unit="mm" value="5.3">5.3 mm</quantity>
long, ca. 2.0 mm wide (left forewing). Trichosors present along the entire wing margin, one trichosor between adjacent veins; marginal portions of veins and trichosors with tufts of thin setae; thin setae sparsely present on dorsal and ventral surfaces of longitudinal veins, but absent on crossveins. Costal space broad, narrowed distally; humeral veinlets recurrent, pectinately branched, most subcostal veinlets once to twice forked. Sc fused with RA at distal fourth (
<figureCitation id="C6EB388882304C63FE145ADBE52CC8AF" box="[501,592,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Details of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, Sc and RA on right forewing; B, Sc and RA on left hind wing; C, right lateral view of head and thorax; D, dorsal view of head and thorax; E, left metatibia and tarsus; F, right lateral view of terminalia.Abbreviations: ect: ectoproct; gx9: gonocoxite 9; hp: humeral plate; hv: humeral veinlet; lp: labial palpus; mp: maxillary palpus; mtti: metatibia; RA: radius anterior; Sc: subcosta; st: gonostylus of gonocoxite 9; ts: tibial spurs. Scale bars: 0.1 mm for A and B; 0.5 mm for CE; 0.2 mm for F." figureDoi="http://doi.org/10.5281/zenodo.7031821" httpUri="https://zenodo.org/record/7031821/files/figure.png" pageId="5" pageNumber="385">Fig. 3A</figureCitation>
). Subcostal space, proximal crossvein (1sc-r) located at origin of RP, intermediate crossvein (2sc-r) present. RP stemed single origin, with nine branches, most proximal branch deeply forked at the near stem, all branches terminally forked rather shallowly once or twice; five (right forewing) or six (left forewing) third series intra-crossveins and ten (left forewing) or eleven (right forewing) forth series intra-crossveins present. Three ra-rp crossveins (2ra-rp, 3ra-rp, and 4ra-rp) present. M divided into MA and MP, MP deeply forked; two third series intra-crossveins and three fourth series intra-crossveins present. Three r-m crossveins (2r-m, 3r-m, and 4r-m) present. Long radiomedial flexion line clearly discernible. Cu divided into CuA and CuP, CuP close to wing base; CuA pectinately branched, with four branches, most proximal branch deeply forked; CuP deeply forked; all branches terminally forked shallowly once or twice; one second series intra-crossvein and six fourth series intra-crossveins present. Three m-cu crossveins (1m-cu, 2m-cu, and 4m-cu) present. Short mediocubital flexion line clearly discernible. Three anal veins present, poorly preserved; A1 deeply forked. Two cu-a crossveins (1cu-a and 4cu-a) present. Cubitoanal flexion line clearly discernible.
</paragraph>
<caption id="0AAF748582304C63FF765820E5E2CA9A" ID-Table-UUID="0AAF748582304C63FF765820E5E2CA9A" box="[151,670,620,644]" httpUri="http://table.plazi.org/id/0AAF748582304C63FF765820E5E2CA9A" pageId="5" pageNumber="385" startId="5.[151,239,620,644]" targetBox="[166,1419,661,1587]" targetIsTable="true" targetPageId="5">
<paragraph id="5E6F240D82304C63FF765820E5E2CA9A" blockId="5.[151,670,620,644]" box="[151,670,620,644]" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF765820E676CA9A" bold="true" box="[151,266,620,644]" pageId="5" pageNumber="385">TABLE 1.</emphasis>
Venation in Mesozoic
<taxonomicName id="99D05F8E82304C63FDE15820E5E7CA9A" authorityName="Latreille" authorityYear="1802" box="[512,667,620,644]" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="385" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="5E6F240D82304C63FF4758D9E3A7CE2D" pageId="5" pageNumber="385">
<table id="2CD0D6AD8230B399FF4758D9E2F7CE2D" box="[166,1419,661,1587]" gridcols="5" gridrows="25" pageId="5" pageNumber="385">
<tr id="E0E0264F8230B399FF4758D9E2F7CAB3" box="[166,1419,661,685]" gridrow="0" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF4758D9E6C1CAB3" box="[166,445,661,685]" gridcol="0" gridrow="0" pageId="5" pageNumber="385">Taxon</th>
<th id="A3314F338230B399FE0058D9E43DCAB3" box="[481,833,661,685]" gridcol="1" gridrow="0" pageId="5" pageNumber="385">Deposit</th>
<th id="A3314F338230B399FC8558D9E490CAB3" box="[868,1004,661,685]" gridcol="2" gridrow="0" pageId="5" pageNumber="385">Distal part of</th>
<th id="A3314F338230B399FBED58D9E3EBCAB3" box="[1036,1175,661,685]" gridcol="3" gridrow="0" pageId="5" pageNumber="385">Number of</th>
<th id="A3314F338230B399FB5458D9E2F7CAB3" box="[1205,1419,661,685]" gridcol="4" gridrow="0" pageId="5" pageNumber="385">Reference</th>
</tr>
<tr id="E0E0264F8230B399FF4758F5E2F7CACF" box="[166,1419,697,721]" gridrow="1" pageId="5" pageNumber="385" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="A3314F338230B399FC8558F5E490CACF" box="[868,1004,697,721]" gridcol="2" gridrow="1" pageId="5" pageNumber="385">Sc and RA in</td>
<td id="A3314F338230B399FBED58F5E3EBCACF" box="[1036,1175,697,721]" gridcol="3" gridrow="1" pageId="5" pageNumber="385">stem of RP in</td>
</tr>
<tr id="E0E0264F8230B399FF475891E2F7CAEB" box="[166,1419,733,757]" gridrow="2" pageId="5" pageNumber="385" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="A3314F338230B399FC855891E490CAEB" box="[868,1004,733,757]" gridcol="2" gridrow="2" pageId="5" pageNumber="385">the forewing</td>
<td id="A3314F338230B399FBED5891E3EBCAEB" box="[1036,1175,733,757]" gridcol="3" gridrow="2" pageId="5" pageNumber="385">the forewing</td>
</tr>
<tr id="E0E0264F8230B399FF475949E2F7CB03" box="[166,1419,773,797]" gridrow="3" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF475949E6C1CB03" box="[166,445,773,797]" gridcol="0" gridrow="3" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475949E6DBCB03" box="[166,423,773,797]" italics="true" pageId="5" pageNumber="385">Promegalomus anomalus</emphasis>
</th>
<td id="A3314F338230B399FE005949E43DCB03" box="[481,833,773,797]" gridcol="1" gridrow="3" pageId="5" pageNumber="385">Late Jurassic, Kazakhstan</td>
<td id="A3314F338230B399FC855949E490CB03" box="[868,1004,773,797]" gridcol="2" gridrow="3" pageId="5" pageNumber="385">? (probably</td>
<td id="A3314F338230B399FBED5949E3EBCB03" box="[1036,1175,773,797]" gridcol="3" gridrow="3" pageId="5" pageNumber="385">three</td>
<td id="A3314F338230B399FB545949E2F7CB03" box="[1205,1419,773,797]" gridcol="4" gridrow="3" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FB545949E23BCB03" author="Panfilov, D. V." box="[1205,1351,773,797]" pageId="5" pageNumber="385" pagination="82 - 111" refId="ref5584" refString="Panfilov, D. V. (1980) New representatives of lacewings (Neuroptera) from the Jurassic of Karatau. In: Dolin, V. G., Panfilov, D. V., Ponomarenko, A. G. &amp; Pritykina, L. N. (Eds.), Fossil insects of the Mesozoic. Naukova Dumka, Kiev, pp. 82 - 111. [in Russian]" type="book chapter" year="1980">Panfilov 1980</bibRefCitation>
: fig.
</td>
</tr>
<tr id="E0E0264F8230B399FF475965E2F7CB5F" box="[166,1419,809,833]" gridrow="4" pageId="5" pageNumber="385" rowspan-1="1" rowspan-3="1">
<th id="A3314F338230B399FF475965E6C1CB5F" box="[166,445,809,833]" gridcol="0" gridrow="4" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FF475965E645CB5F" author="Panfilov, D. V." box="[166,313,809,833]" pageId="5" pageNumber="385" pagination="82 - 111" refId="ref5584" refString="Panfilov, D. V. (1980) New representatives of lacewings (Neuroptera) from the Jurassic of Karatau. In: Dolin, V. G., Panfilov, D. V., Ponomarenko, A. G. &amp; Pritykina, L. N. (Eds.), Fossil insects of the Mesozoic. Naukova Dumka, Kiev, pp. 82 - 111. [in Russian]" type="book chapter" year="1980">Panfilov, 1980</bibRefCitation>
</th>
<td id="A3314F338230B399FC855965E490CB5F" box="[868,1004,809,833]" gridcol="2" gridrow="4" pageId="5" pageNumber="385">fused)</td>
<td id="A3314F338230B399FB545965E2F7CB5F" box="[1205,1419,809,833]" gridcol="4" gridrow="4" pageId="5" pageNumber="385">91</td>
</tr>
<tr id="E0E0264F8230B399FF47591DE2F7CB77" box="[166,1419,849,873]" gridrow="5" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF47591DE6C1CB77" box="[166,445,849,873]" gridcol="0" gridrow="5" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF47591DE6E1CB77" box="[166,413,849,873]" italics="true" pageId="5" pageNumber="385">Purbemerobius medialis</emphasis>
</th>
<td id="A3314F338230B399FE00591DE43DCB77" box="[481,833,849,873]" gridcol="1" gridrow="5" pageId="5" pageNumber="385">Early Cretaceous, late Berreiasian,</td>
<td id="A3314F338230B399FC85591DE490CB77" box="[868,1004,849,873]" gridcol="2" gridrow="5" pageId="5" pageNumber="385">?</td>
<td id="A3314F338230B399FBED591DE3EBCB77" box="[1036,1175,849,873]" gridcol="3" gridrow="5" pageId="5" pageNumber="385">?</td>
<td id="A3314F338230B399FB54591DE2F7CB77" box="[1205,1419,849,873]" gridcol="4" gridrow="5" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FB54591EE20CCB77" author="Jepson, J. E. &amp; Makarkin, V. N. &amp; Coram, R. A." box="[1205,1392,849,873]" pageId="5" pageNumber="385" pagination="31 - 47" refId="ref4835" refString="Jepson, J. E., Makarkin, V. N. &amp; Coram, R. A. (2012) Lacewings (Insecta: Neuroptera) from the Lower Cretaceous Purbeck Limestone Group of southern England. Cretaceous Research, 34, 31 - 47. http: // doi. org / 10.1016 / j. cretres. 2011.10.001" type="journal article" year="2012">
Jepson
<emphasis id="6CA4F81F82304C63FAE1591EE248CB77" box="[1280,1332,849,873]" italics="true" pageId="5" pageNumber="385">et al.</emphasis>
2012
</bibRefCitation>
:
</td>
</tr>
<tr id="E0E0264F8230B399FF475939E2F7CB93" box="[166,1419,885,909]" gridrow="6" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1">
<th id="A3314F338230B399FF475939E6C1CB93" box="[166,445,885,909]" gridcol="0" gridrow="6" pageId="5" pageNumber="385">Jepson, Makarkin and Co-</th>
<td id="A3314F338230B399FE005939E43DCB93" box="[481,833,885,909]" gridcol="1" gridrow="6" pageId="5" pageNumber="385">Durlston Formation, England,</td>
<td id="A3314F338230B399FB545939E2F7CB93" box="[1205,1419,885,909]" gridcol="4" gridrow="6" pageId="5" pageNumber="385">fig. 16</td>
</tr>
<tr id="E0E0264F8230B399FF4759D5E2F7CBAF" box="[166,1419,921,945]" gridrow="7" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="A3314F338230B399FF4759D5E6C1CBAF" box="[166,445,921,945]" gridcol="0" gridrow="7" pageId="5" pageNumber="385">ram, 2012</th>
<td id="A3314F338230B399FE0059D5E43DCBAF" box="[481,833,921,945]" gridcol="1" gridrow="7" pageId="5" pageNumber="385">United Kingdom</td>
</tr>
<tr id="E0E0264F8230B399FF47598DE2F7CBC4" box="[166,1419,961,986]" gridrow="8" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF47598DE6C1CBC4" box="[166,445,961,986]" gridcol="0" gridrow="8" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF47598DE6D7CBC7" box="[166,427,961,985]" italics="true" pageId="5" pageNumber="385">Cretomerobius disjunctus</emphasis>
</th>
<td id="A3314F338230B399FE00598DE43DCBC4" box="[481,833,961,986]" gridcol="1" gridrow="8" pageId="5" pageNumber="385">Early Cretaceous, Mongolia</td>
<td id="A3314F338230B399FC85598DE490CBC4" box="[868,1004,961,986]" gridcol="2" gridrow="8" pageId="5" pageNumber="385">fused</td>
<td id="A3314F338230B399FBED598DE3EBCBC4" box="[1036,1175,961,986]" gridcol="3" gridrow="8" pageId="5" pageNumber="385">three</td>
<td id="A3314F338230B399FB54598DE2F7CBC4" box="[1205,1419,961,986]" gridcol="4" gridrow="8" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FB54598DE202CBC4" author="Ponomarenko, A. G." box="[1205,1406,961,986]" pageId="5" pageNumber="385" refId="ref5760" refString="Ponomarenko, A. G. (1992) New lacewings (Insecta, Neuroptera) from the Mesozoic of Mongolia. In: Grunt, T. A. (Ed.), New taxa of the fossil invertebrates of Mongolia. Transactions of the Joint Soviet - Mongolian Paleontological Expedition. Vol. 41. Nauka Press, Moscow, pp. 101 - 111. [in Russian]" type="book" year="1992">Ponomarenko 1992</bibRefCitation>
:
</td>
</tr>
<tr id="E0E0264F8230B399FF4759A9E2F7CBE0" box="[166,1419,997,1022]" gridrow="9" pageId="5" pageNumber="385" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="A3314F338230B399FF4759A9E6C1CBE0" box="[166,445,997,1022]" gridcol="0" gridrow="9" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FF4759A9E60ECBE0" author="Ponomarenko, A. G." box="[166,370,997,1022]" pageId="5" pageNumber="385" refId="ref5760" refString="Ponomarenko, A. G. (1992) New lacewings (Insecta, Neuroptera) from the Mesozoic of Mongolia. In: Grunt, T. A. (Ed.), New taxa of the fossil invertebrates of Mongolia. Transactions of the Joint Soviet - Mongolian Paleontological Expedition. Vol. 41. Nauka Press, Moscow, pp. 101 - 111. [in Russian]" type="book" year="1992">Ponomarenko, 1992</bibRefCitation>
</th>
<td id="A3314F338230B399FB5459A9E2F7CBE0" box="[1205,1419,997,1022]" gridcol="4" gridrow="9" pageId="5" pageNumber="385">fig. 4</td>
</tr>
<tr id="E0E0264F8230B399FF475E42E2F7CC38" box="[166,1419,1038,1062]" gridrow="10" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF475E42E6C1CC38" box="[166,445,1038,1062]" gridcol="0" gridrow="10" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475E42E6D6CC38" box="[166,426,1038,1062]" italics="true" pageId="5" pageNumber="385">Hemeroberotha sinefurca</emphasis>
</th>
<td id="A3314F338230B399FE005E42E43DCC38" box="[481,833,1038,1062]" gridcol="1" gridrow="10" pageId="5" pageNumber="385">Mid-Cretaceous, earliest</td>
<td id="A3314F338230B399FC855E42E490CC38" box="[868,1004,1038,1062]" gridcol="2" gridrow="10" pageId="5" pageNumber="385">separated</td>
<td id="A3314F338230B399FBED5E42E3EBCC38" box="[1036,1175,1038,1062]" gridcol="3" gridrow="10" pageId="5" pageNumber="385">single</td>
<td id="A3314F338230B399FB545E42E2F7CC38" box="[1205,1419,1038,1062]" gridcol="4" gridrow="10" pageId="5" pageNumber="385">Makarkin &amp; Gröhn</td>
</tr>
<tr id="E0E0264F8230B399FF475E7EE2F7CC54" box="[166,1419,1074,1098]" gridrow="11" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1">
<th id="A3314F338230B399FF475E7EE6C1CC54" box="[166,445,1074,1098]" gridcol="0" gridrow="11" pageId="5" pageNumber="385">Makarkin and Gröhn, 2020</th>
<td id="A3314F338230B399FE005E7EE43DCC54" box="[481,833,1074,1098]" gridcol="1" gridrow="11" pageId="5" pageNumber="385">Cenomanian, Myanmar</td>
<td id="A3314F338230B399FB545E7EE2F7CC54" box="[1205,1419,1074,1098]" gridcol="4" gridrow="11" pageId="5" pageNumber="385">2022: fig. 4</td>
</tr>
<tr id="E0E0264F8230B399FF475E16E2F7CC6C" box="[166,1419,1114,1138]" gridrow="12" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF475E16E6C1CC6C" box="[166,445,1114,1138]" gridcol="0" gridrow="12" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475E16E6DFCC6C" box="[166,419,1114,1138]" italics="true" pageId="5" pageNumber="385">
<taxonomicName id="99D05F8E82304C63FF475E16E62CCC6F" authority="Nakamine &amp; Yamamoto &amp; Takahashi, 2022" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi" authorityYear="2022" box="[166,336,1114,1137]" class="Insecta" family="Hemerobiidae" genus="Cretoneuronema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="genus" status="gen. nov.">Cretoneuronema</taxonomicName>
jarzem-
</emphasis>
</th>
<td id="A3314F338230B399FE005E16E43DCC6C" box="[481,833,1114,1138]" gridcol="1" gridrow="12" pageId="5" pageNumber="385">Mid-Cretaceous, earliest</td>
<td id="A3314F338230B399FC855E16E490CC6C" box="[868,1004,1114,1138]" gridcol="2" gridrow="12" pageId="5" pageNumber="385">fused</td>
<td id="A3314F338230B399FBED5E16E3EBCC6C" box="[1036,1175,1114,1138]" gridcol="3" gridrow="12" pageId="5" pageNumber="385">two</td>
<td id="A3314F338230B399FB545E16E2F7CC6C" box="[1205,1419,1114,1138]" gridcol="4" gridrow="12" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FB545E16E233CC6C" author="Liu, X. Y. &amp; Chen, Z. L. &amp; Zhuo, D." box="[1205,1359,1114,1138]" pageId="5" pageNumber="385" pagination="226 - 232" refId="ref5207" refString="Liu, X. Y., Chen, Z. L. &amp; Zhuo, D. (2022) Cretoneuronema gen. nov. (Neuroptera: Hemerobiidae), a new brown lacewing genus from the mid-Cretaceous Kachin amber. Palaeoentomology, 5 (3), 226 - 232. https: // doi. org / 10.11646 / palaeoentomology. 5.3.4" type="journal article" year="2022">
Liu
<emphasis id="6CA4F81F82304C63FB3E5E17E26ECC6C" box="[1247,1298,1114,1138]" italics="true" pageId="5" pageNumber="385">et al.</emphasis>
2022
</bibRefCitation>
: fig.3
</td>
</tr>
<tr id="E0E0264F8230B399FF475E32E2F7CC88" box="[166,1419,1150,1174]" gridrow="13" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="A3314F338230B399FF475E32E6C1CC88" box="[166,445,1150,1174]" gridcol="0" gridrow="13" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475E32E78FCC88" box="[166,243,1150,1174]" italics="true" pageId="5" pageNumber="385">bowskii</emphasis>
Liu, Chen and
</th>
<td id="A3314F338230B399FE005E32E43DCC88" box="[481,833,1150,1174]" gridcol="1" gridrow="13" pageId="5" pageNumber="385">Cenomanian, Myanmar</td>
</tr>
<tr id="E0E0264F8230B399FF475EEEE2F7CCA4" box="[166,1419,1186,1210]" gridrow="14" pageId="5" pageNumber="385" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="A3314F338230B399FF475EEEE6C1CCA4" box="[166,445,1186,1210]" gridcol="0" gridrow="14" pageId="5" pageNumber="385">Zhuo, 2022</th>
</tr>
<tr id="E0E0264F8230B399FF475E86E2F7CCFC" box="[166,1419,1226,1250]" gridrow="15" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF475E86E6C1CCFC" box="[166,445,1226,1250]" gridcol="0" gridrow="15" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475E86E6E3CCFC" box="[166,415,1226,1250]" italics="true" pageId="5" pageNumber="385">
<taxonomicName id="99D05F8E82304C63FF475E86E612CCFC" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi" authorityYear="2022" box="[166,366,1226,1250]" class="Insecta" family="Hemerobiidae" genus="Archaeomegalomus" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="385" phylum="Arthropoda" rank="genus">Archaeomegalomus</taxonomicName>
bur-
</emphasis>
</th>
<td id="A3314F338230B399FE005E86E43DCCFC" box="[481,833,1226,1250]" gridcol="1" gridrow="15" pageId="5" pageNumber="385">Mid-Cretaceous, earliest Cenoma-</td>
<td id="A3314F338230B399FC855E86E490CCFC" box="[868,1004,1226,1250]" gridcol="2" gridrow="15" pageId="5" pageNumber="385">fused</td>
<td id="A3314F338230B399FBED5E86E3EBCCFC" box="[1036,1175,1226,1250]" gridcol="3" gridrow="15" pageId="5" pageNumber="385">single</td>
<td id="A3314F338230B399FB545E86E2F7CCFC" box="[1205,1419,1226,1250]" gridcol="4" gridrow="15" pageId="5" pageNumber="385">this study</td>
</tr>
<tr id="E0E0264F8230B399FF475EA2E2F7CD19" box="[166,1419,1262,1287]" gridrow="16" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="A3314F338230B399FF475EA2E6C1CD19" box="[166,445,1262,1287]" gridcol="0" gridrow="16" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475EA3E793CD18" box="[166,239,1263,1286]" italics="true" pageId="5" pageNumber="385">miticus</emphasis>
gen. et sp. nov.
</th>
<td id="A3314F338230B399FE005EA2E43DCD19" box="[481,833,1262,1287]" gridcol="1" gridrow="16" pageId="5" pageNumber="385">nian, Myanmar</td>
</tr>
<tr id="E0E0264F8230B399FF475F5AE2F7CD31" box="[166,1419,1302,1327]" gridrow="17" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF475F5AE6C1CD31" box="[166,445,1302,1327]" gridcol="0" gridrow="17" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475F5AE6FBCD30" box="[166,391,1302,1326]" italics="true" pageId="5" pageNumber="385">Plesiorobius sibericus</emphasis>
</th>
<td id="A3314F338230B399FE005F5AE43DCD31" box="[481,833,1302,1327]" gridcol="1" gridrow="17" pageId="5" pageNumber="385">Late Cretaceous, Cenomanian,</td>
<td id="A3314F338230B399FC855F5AE490CD31" box="[868,1004,1302,1327]" gridcol="2" gridrow="17" pageId="5" pageNumber="385">fused</td>
<td id="A3314F338230B399FBED5F5AE3EBCD31" box="[1036,1175,1302,1327]" gridcol="3" gridrow="17" pageId="5" pageNumber="385">single</td>
<td id="A3314F338230B399FB545F5AE2F7CD31" box="[1205,1419,1302,1327]" gridcol="4" gridrow="17" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FB545F5AE228CD31" author="Makarkin, V. N." box="[1205,1364,1302,1327]" pageId="5" pageNumber="385" pagination="283 - 292" refId="ref5273" refString="Makarkin, V. N. (1994) Upper Cretaceous Neuroptera from Russia and Kazakhstan. Annales de la Societe Entomologique de France, New Series, 30 (3), 283 - 292." type="journal article" year="1994">Makarkin 1994</bibRefCitation>
: figs.
</td>
</tr>
<tr id="E0E0264F8230B399FF475F76E2F7CD4D" box="[166,1419,1338,1363]" gridrow="18" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1">
<th id="A3314F338230B399FF475F76E6C1CD4D" box="[166,445,1338,1363]" gridcol="0" gridrow="18" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FF475F76E634CD4D" author="Makarkin, V. N." box="[166,328,1338,1363]" pageId="5" pageNumber="385" pagination="283 - 292" refId="ref5273" refString="Makarkin, V. N. (1994) Upper Cretaceous Neuroptera from Russia and Kazakhstan. Annales de la Societe Entomologique de France, New Series, 30 (3), 283 - 292." type="journal article" year="1994">Makarkin, 1994</bibRefCitation>
</th>
<td id="A3314F338230B399FE005F76E43DCD4D" box="[481,833,1338,1363]" gridcol="1" gridrow="18" pageId="5" pageNumber="385">northeastern Russia</td>
<td id="A3314F338230B399FB545F76E2F7CD4D" box="[1205,1419,1338,1363]" gridcol="4" gridrow="18" pageId="5" pageNumber="385">912</td>
</tr>
<tr id="E0E0264F8230B399FF475F2FE2F7CD65" box="[166,1419,1379,1403]" gridrow="19" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF475F2FE6C1CD65" box="[166,445,1379,1403]" gridcol="0" gridrow="19" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475F2FE65ACD65" box="[166,294,1379,1403]" italics="true" pageId="5" pageNumber="385">Plesiorobius</emphasis>
cf.
<emphasis id="6CA4F81F82304C63FEAD5F2FE6C1CD65" box="[332,445,1379,1403]" italics="true" pageId="5" pageNumber="385">canadensis</emphasis>
</th>
<td id="A3314F338230B399FE005F2FE43DCD65" box="[481,833,1379,1403]" gridcol="1" gridrow="19" pageId="5" pageNumber="385">Late Cretaceous, Santonian,</td>
<td id="A3314F338230B399FC855F2FE490CD65" box="[868,1004,1379,1403]" gridcol="2" gridrow="19" pageId="5" pageNumber="385">?</td>
<td id="A3314F338230B399FBED5F2FE3EBCD65" box="[1036,1175,1379,1403]" gridcol="3" gridrow="19" pageId="5" pageNumber="385">single</td>
<td id="A3314F338230B399FB545F2FE2F7CD65" box="[1205,1419,1379,1403]" gridcol="4" gridrow="19" pageId="5" pageNumber="385">
<bibRefCitation id="3A4159FC82304C63FB545F2FE228CD65" author="Makarkin, V. N." box="[1205,1364,1379,1403]" pageId="5" pageNumber="385" pagination="283 - 292" refId="ref5273" refString="Makarkin, V. N. (1994) Upper Cretaceous Neuroptera from Russia and Kazakhstan. Annales de la Societe Entomologique de France, New Series, 30 (3), 283 - 292." type="journal article" year="1994">Makarkin 1994</bibRefCitation>
: fig.
</td>
</tr>
<tr id="E0E0264F8230B399FF475FCBE2F7CD81" box="[166,1419,1415,1439]" gridrow="20" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1">
<th id="A3314F338230B399FF475FCBE6C1CD81" box="[166,445,1415,1439]" gridcol="0" gridrow="20" pageId="5" pageNumber="385">Klimaszewski and Kevan,</th>
<td id="A3314F338230B399FE005FCBE43DCD81" box="[481,833,1415,1439]" gridcol="1" gridrow="20" pageId="5" pageNumber="385">Taymyr amber, northern Siberia</td>
<td id="A3314F338230B399FB545FCBE2F7CD81" box="[1205,1419,1415,1439]" gridcol="4" gridrow="20" pageId="5" pageNumber="385">13</td>
</tr>
<tr id="E0E0264F8230B399FF475FE7E2F7CDDD" box="[166,1419,1451,1475]" gridrow="21" pageId="5" pageNumber="385" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="A3314F338230B399FF475FE7E6C1CDDD" box="[166,445,1451,1475]" gridcol="0" gridrow="21" pageId="5" pageNumber="385">1986</th>
</tr>
<tr id="E0E0264F8230B399FF475F9FE2F7CDF5" box="[166,1419,1491,1515]" gridrow="22" pageId="5" pageNumber="385">
<th id="A3314F338230B399FF475F9FE6C1CDF5" box="[166,445,1491,1515]" gridcol="0" gridrow="22" pageId="5" pageNumber="385">
<emphasis id="6CA4F81F82304C63FF475F9FE6E1CDF5" box="[166,413,1491,1515]" italics="true" pageId="5" pageNumber="385">Plesiorobius canadensis</emphasis>
</th>
<td id="A3314F338230B399FE005F9FE43DCDF5" box="[481,833,1491,1515]" gridcol="1" gridrow="22" pageId="5" pageNumber="385">Late Cretaceous, Campanian,</td>
<td id="A3314F338230B399FC855F9FE490CDF5" box="[868,1004,1491,1515]" gridcol="2" gridrow="22" pageId="5" pageNumber="385">fused</td>
<td id="A3314F338230B399FBED5F9FE3EBCDF5" box="[1036,1175,1491,1515]" gridcol="3" gridrow="22" pageId="5" pageNumber="385">single</td>
<td id="A3314F338230B399FB545F9FE2F7CDF5" box="[1205,1419,1491,1515]" gridcol="4" gridrow="22" pageId="5" pageNumber="385">Klimaszewski &amp;</td>
</tr>
<tr id="E0E0264F8230B399FF475FBBE2F7CE11" box="[166,1419,1527,1551]" gridrow="23" pageId="5" pageNumber="385" rowspan-2="1" rowspan-3="1">
<th id="A3314F338230B399FF475FBBE6C1CE11" box="[166,445,1527,1551]" gridcol="0" gridrow="23" pageId="5" pageNumber="385">Klimaszewski and Kevan,</th>
<td id="A3314F338230B399FE005FBBE43DCE11" box="[481,833,1527,1551]" gridcol="1" gridrow="23" pageId="5" pageNumber="385">Canadian amber</td>
<td id="A3314F338230B399FB545FBBE2F7CE11" box="[1205,1419,1527,1551]" gridcol="4" gridrow="23" pageId="5" pageNumber="385">Kevan 1986: figs.</td>
</tr>
<tr id="E0E0264F8230B399FF475C57E2F7CE2D" box="[166,1419,1563,1587]" gridrow="24" pageId="5" pageNumber="385" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="A3314F338230B399FF475C57E6C1CE2D" box="[166,445,1563,1587]" gridcol="0" gridrow="24" pageId="5" pageNumber="385">1986</th>
<td id="A3314F338230B399FB545C57E2F7CE2D" box="[1205,1419,1563,1587]" gridcol="4" gridrow="24" pageId="5" pageNumber="385">1, 2</td>
</tr>
</table>
</paragraph>
<paragraph id="5E6F240D82304C60FF265C2FE582C8CB" blockId="5.[151,1437,1634,2021]" lastBlockId="6.[151,1437,151,753]" lastPageId="6" lastPageNumber="386" pageId="5" pageNumber="385">
Hind wing hyaline (
<figureCitation id="C6EB388882304C63FE4F5C2FE551CE63" box="[430,557,1635,1661]" captionStart="FIGURE 2" captionStartId="3.[151,250,951,975]" captionTargetBox="[151,1436,181,927]" captionTargetId="figure-21@3.[151,1436,181,927]" captionTargetPageId="3" captionText="FIGURE 2. Wing venation of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, right forewing; B, left forewing; C, right hind wing; D, left hind wing. Abbreviations: A1A3: first to third anal vein; CuA: cubitus anterior; CuP: cubitus posterior; hv: humeral vein; MA: media anterior; MP: media posterior; RA: radius anterior; RP: radius posterior; Sc: subcosta. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7031819" httpUri="https://zenodo.org/record/7031819/files/figure.png" pageId="5" pageNumber="385">Fig. 2C, D</figureCitation>
), elongate-ovoid, ca.
<quantity id="992889E882304C63FCC75C2EE403CE63" box="[806,895,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="385" unit="mm" value="4.6">4.6 mm</quantity>
long, ca.
<quantity id="992889E882304C63FC125C2FE330CE63" box="[1011,1100,1635,1661]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="385" unit="mm" value="1.8">1.8 mm</quantity>
wide (right hind wing); dark brown on pterostigmal area. Trichosors present from the pterostigmal area of costal margin to along the entire wing margin, one trichosor between adjacent veins; marginal portions of veins and trichosors with tufts of thin setae; thin setae sparsely present on dorsal and ventral surfaces of longitudinal veins, but absent on crossveins. Costal space narrowed; humeral lobe triangular, humeral plate bears dense bristles; almost subcostal veinlets simple, five veinlets forked at distal (left hind wing). Subcostal space one crossvein (3sc-r) present (
<figureCitation id="C6EB388882304C63FBDA5D5BE3EFCF2F" box="[1083,1171,1815,1841]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Details of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, Sc and RA on right forewing; B, Sc and RA on left hind wing; C, right lateral view of head and thorax; D, dorsal view of head and thorax; E, left metatibia and tarsus; F, right lateral view of terminalia.Abbreviations: ect: ectoproct; gx9: gonocoxite 9; hp: humeral plate; hv: humeral veinlet; lp: labial palpus; mp: maxillary palpus; mtti: metatibia; RA: radius anterior; Sc: subcosta; st: gonostylus of gonocoxite 9; ts: tibial spurs. Scale bars: 0.1 mm for A and B; 0.5 mm for CE; 0.2 mm for F." figureDoi="http://doi.org/10.5281/zenodo.7031821" httpUri="https://zenodo.org/record/7031821/files/figure.png" pageId="5" pageNumber="385">Fig. 3B</figureCitation>
). RP divided into seven branches; most proximal branche (RP1) loop like vein at the near stem (right hind wing), deeply forked; one second series intra-crossvein and eight third series intra-crossveins present. Two ra-rp crossveins (1ra-rp and 3ra-rp) present, 1ra-rp long and slightly sinuate. M divided into MA and MP, one intra-crossvein (3ma-mp) present. Three r-m crossveins (1r-m, 2r-m, and 3r-m) present, basal 1r-m long and sinuate. Cu divided into CuA and CuP; CuP close to wing base: CuA pectinately branched, with six (right hind wing) or five (left hind wing) branches; one third series intra-crossvein present. Two m-cu crossveins (1m-cu and 3m-cu) present. Anal vein poorly preserved; A1 deeply forked. Two cu-a crossveins (1cu-a and 3cu-a) present.
</paragraph>
<paragraph id="5E6F240D82334C60FF265A93E363C95F" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="386">
Abdomen. Visible in right lateral view (
<figureCitation id="C6EB388882334C60FD745A93E590C8E7" box="[661,748,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1626]" captionTargetId="figure-18@4.[151,1436,181,1626]" captionTargetPageId="4" captionText="FIGURE 3. Details of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid-Cretaceous Kachin amber. A, Sc and RA on right forewing; B, Sc and RA on left hind wing; C, right lateral view of head and thorax; D, dorsal view of head and thorax; E, left metatibia and tarsus; F, right lateral view of terminalia.Abbreviations: ect: ectoproct; gx9: gonocoxite 9; hp: humeral plate; hv: humeral veinlet; lp: labial palpus; mp: maxillary palpus; mtti: metatibia; RA: radius anterior; Sc: subcosta; st: gonostylus of gonocoxite 9; ts: tibial spurs. Scale bars: 0.1 mm for A and B; 0.5 mm for CE; 0.2 mm for F." figureDoi="http://doi.org/10.5281/zenodo.7031821" httpUri="https://zenodo.org/record/7031821/files/figure.png" pageId="6" pageNumber="386">Fig. 3F</figureCitation>
), poorly preserved, but posterior part rather well preserved. Tergite 6 and 7 developed each posterior margin. Terminal segment unclear. Gonocoxite 9 visible, scattered setae on surface; gonostylus of gonocoxite 9 well developed. Ectoproct poorly preserved.
</paragraph>
</subSubSection>
<subSubSection id="16CA778682334C60FF265B06E5CFC9EB" pageId="6" pageNumber="386" type="materials_examined">
<paragraph id="5E6F240D82334C60FF265B06E6BCC9B3" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="386">
<emphasis id="6CA4F81F82334C60FF265B06E64AC97A" bold="true" box="[199,310,330,356]" pageId="6" pageNumber="386">Material.</emphasis>
<materialsCitation id="EEB82E5082334C60FEDF5B07E6BCC9B3" ID-GBIF-Occurrence="3894254301" collectionCode="AMNH" pageId="6" pageNumber="386" specimenCount="1" typeStatus="holotype">
<typeStatus id="816B9AAF82334C60FEDF5B07E6AAC97B" box="[318,470,331,357]" pageId="6" pageNumber="386" type="holotype">HOLOTYPE</typeStatus>
: a nearly complete female adult (
<figureCitation id="C6EB388882334C60FCAD5B07E4F2C97B" box="[844,910,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[216,1371,181,1919]" captionTargetId="figure-18@2.[216,1371,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and wing detail of Archaeomegalomus burmiticus gen. et sp. nov., holotype (AMNH Bu-SY31) in mid- Cretaceous Kachin amber. A, left lateral view; B, right lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7031816" httpUri="https://zenodo.org/record/7031816/files/figure.png" pageId="6" pageNumber="386">Fig. 1</figureCitation>
), partly unobservable due to cracks, preserved in approximately
<quantity id="992889E882334C60FEBD5B23E6CBC997" box="[348,439,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="386" unit="mm" value="14.0">14 mm</quantity>
×
<quantity id="992889E882334C60FE2E5B22E559C997" box="[463,549,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="386" unit="mm" value="7.5">7.5 mm</quantity>
×
<quantity id="992889E882334C60FDA35B23E5FFC997" box="[578,643,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="386" unit="mm" value="4.0">4 mm</quantity>
flat drop-shaped, yellow, transparent amber piece; specimen accession number
<collectionCode id="38C1BCC882334C60FF135BDFE634C9B3" box="[242,328,403,429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="386">AMNH</collectionCode>
Bu-SY31.
</materialsCitation>
</paragraph>
<paragraph id="5E6F240D82334C60FF265BFBE5CFC9EB" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="386">
<materialsCitation id="EEB82E5082334C60FF265BFBE5D3C9EB" ID-GBIF-Occurrence="3894254302" country="Myanmar" county="lower Cenomanian" latitude="26.333334" location="Hukawng Valley" longLatPrecision="1242" longitude="96.6" municipality="upper Albian" pageId="6" pageNumber="386" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="6CA4F81F82334C60FF265BFBE578C9CE" bold="true" box="[199,516,438,465]" pageId="6" pageNumber="386">
<typeStatus id="816B9AAF82334C60FF265BFBE67EC9CF" box="[199,258,439,465]" pageId="6" pageNumber="386">Type</typeStatus>
locality and horizon.
</emphasis>
<location id="5B0F72D682334C60FDED5BFBE5B6C9CF" LSID="urn:lsid:plazi:treatment:D679951B82364C60FF765F29E5E0CAEF:5B0F72D682334C60FDED5BFBE5B6C9CF" box="[524,714,439,465]" country="Myanmar" county="lower Cenomanian" latitude="26.333334" longLatPrecision="1242" longitude="96.6" municipality="upper Albian" name="Hukawng Valley" pageId="6" pageNumber="386" stateProvince="Kachin">Hukawng Valley</location>
(
<geoCoordinate id="3BE442CA82334C60FD3A5BFBE432C9CF" box="[731,846,439,467]" degrees="26" direction="north" minutes="20" orientation="latitude" pageId="6" pageNumber="386" precision="925" value="26.333334">26° 20 N</geoCoordinate>
,
<geoCoordinate id="3BE442CA82334C60FCBA5BFBE4B6C9CF" box="[859,970,439,467]" degrees="96" direction="east" minutes="36" orientation="longitude" pageId="6" pageNumber="386" precision="925" value="96.6">96° 36 E</geoCoordinate>
),
<collectingRegion id="9C14EAEF82334C60FC015BFBE309C9CF" box="[992,1141,439,465]" country="Myanmar" name="Kachin" pageId="6" pageNumber="386">Kachin State</collectingRegion>
, northern
<collectingCountry id="26C7649D82334C60FB0B5BFBE227C9CF" box="[1258,1371,439,465]" name="Myanmar" pageId="6" pageNumber="386">Myanmar</collectingCountry>
; midCretaceous,
<collectingMunicipality id="BE0BBE7782334C60FEC35B97E6CFC9EB" box="[290,435,475,501]" pageId="6" pageNumber="386">upper Albian</collectingMunicipality>
to
<collectingCounty id="B70E5C8182334C60FE365B97E5D3C9EB" box="[471,687,475,501]" pageId="6" pageNumber="386">lower Cenomanian</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="16CA778682334C60FF265BB3E5D0CA23" pageId="6" pageNumber="386" type="etymology">
<paragraph id="5E6F240D82334C60FF265BB3E5D0CA23" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="386">
<emphasis id="6CA4F81F82334C60FF265BB3E632CA07" bold="true" box="[199,334,511,537]" pageId="6" pageNumber="386">Etymology.</emphasis>
The specific epithet
<taxonomicName id="99D05F8E82334C60FDA35BB3E5C6CA07" authorityName="Nakamine &amp; Yamamoto &amp; Takahashi" authorityYear="2022" box="[578,698,511,537]" class="Insecta" family="Hemerobiidae" genus="Archaeomegalomus" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="species" species="burmiticus">
<emphasis id="6CA4F81F82334C60FDA35BB3E5C6CA07" box="[578,698,511,537]" italics="true" pageId="6" pageNumber="386">burmiticus</emphasis>
</taxonomicName>
, noun in apposition, refers to the occurrence of the fossil in Burmese amber (burmite) from northern
<collectingCountry id="26C7649D82334C60FDD8586FE5D4CA23" box="[569,680,547,573]" name="Myanmar" pageId="6" pageNumber="386">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="16CA778682334C60FF26580BE5E0CAEF" pageId="6" pageNumber="386" type="discussion">
<paragraph id="5E6F240D82334C60FF26580BE5E0CAEF" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="386">
<emphasis id="6CA4F81F82334C60FF26580BE647CA7F" bold="true" box="[199,315,583,609]" pageId="6" pageNumber="386">Remarks.</emphasis>
The distal parts of Sc and RA in the forewing are separated in nearly all extant
<taxonomicName id="99D05F8E82334C60FB2A580BE211CA7F" authorityName="Latreille" authorityYear="1802" box="[1227,1389,583,609]" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
, except for
<taxonomicName id="99D05F8E82334C60FF175827E505CA9B" authority="Oswald, 1994" authorityName="Oswald" authorityYear="1994" box="[246,633,619,645]" class="Insecta" family="Hemerobiidae" genus="Adelphohemerobius" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="6CA4F81F82334C60FF175827E6A9CA9B" box="[246,469,619,645]" italics="true" pageId="6" pageNumber="386">Adelphohemerobius</emphasis>
<bibRefCitation id="3A4159FC82334C60FE3D5827E505CA9B" author="Oswald, J. D." box="[476,633,619,645]" pageId="6" pageNumber="386" pagination="295 - 302" refId="ref5547" refString="Oswald, J. D. (1994) A new phylogenetically basal subfamily of brown lacewings from Chile (Neuroptera: Hemerobiidae). Entomologica scandinavica, 25 (3), 295 - 302." type="journal article" year="1994">Oswald, 1994</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="3A4159FC82334C60FD545827E432CA9B" author="Oswald, J. D." box="[693,846,619,645]" pageId="6" pageNumber="386" pagination="295 - 302" refId="ref5547" refString="Oswald, J. D. (1994) A new phylogenetically basal subfamily of brown lacewings from Chile (Neuroptera: Hemerobiidae). Entomologica scandinavica, 25 (3), 295 - 302." type="journal article" year="1994">Oswald 1994</bibRefCitation>
: fig. 1;
<bibRefCitation id="3A4159FC82334C60FC445827E320CA9B" author="Monserrat, V. J." box="[933,1116,619,645]" pageId="6" pageNumber="386" pagination="123 - 206" refId="ref5411" refString="Monserrat, V. J. (1997) Revision del genero Megalomus de Latinoamerica (Neuroptera, Hemerobiidae). Fragmenta Entomologica, Roma, 29, 123 - 206." type="journal article" year="1997">Monserrat 1997</bibRefCitation>
: fig. 116). In contrast, in the Mesozoic
<taxonomicName id="99D05F8E82334C60FEED58C3E6D2CAB7" authorityName="Latreille" authorityYear="1802" box="[268,430,655,681]" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
,
<taxonomicName id="99D05F8E82334C60FE5B58C3E4E9CAB7" authority="Makarkin, 2020" authorityName="Makarkin &amp; Grohn" authorityYear="2020" box="[442,917,655,681]" class="Insecta" family="Hemerobiidae" genus="Hemeroberotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="species" species="sinefurca">
<emphasis id="6CA4F81F82334C60FE5B58C3E5A5CAB7" box="[442,729,655,681]" italics="true" pageId="6" pageNumber="386">Hemeroberotha sinefurca</emphasis>
Makarkin, 2020
</taxonomicName>
is the only species in which the distal part of Sc and RA in the forewing are definitely separated, while the others are fused (
<tableCitation id="135211B682334C60FBC658FFE300CAD3" box="[1063,1148,691,717]" captionStart="TABLE 1" captionStartId="5.[151,239,620,644]" captionTargetBox="[166,1419,661,1587]" captionTargetPageId="5" captionText="TABLE 1. Venation in Mesozoic Hemerobiidae." httpUri="http://table.plazi.org/id/0AAF748582304C63FF765820E5E2CA9A" pageId="6" pageNumber="386" tableUuid="0AAF748582304C63FF765820E5E2CA9A">Table 1</tableCitation>
). This trait is probably a plesiomorphic condition in the
<taxonomicName id="99D05F8E82334C60FE14589BE5EBCAEF" authorityName="Latreille" authorityYear="1802" box="[501,663,727,753]" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>