312 lines
38 KiB
XML
312 lines
38 KiB
XML
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<mods:title id="79F0C830D2F0EFEFCC9EFFA264C6AE38">New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:title id="0DACA19302319316EA9FBD0B651E0BE5">Russian Entomological Journal</mods:title>
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<mods:part id="05786158555FECEDCABC889A4B9079BB">
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<mods:date id="59B704A8FD2D8E7E15C27EE6DE9D8EEA">2020</mods:date>
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<treatment id="D6009B3AFFFC3B4FFC567EFB0FB0FBFA" ID-DOI="http://doi.org/10.5281/zenodo.11093721" ID-Zenodo-Dep="11093721" LSID="urn:lsid:plazi:treatment:D6009B3AFFFC3B4FFC567EFB0FB0FBFA" httpUri="http://treatment.plazi.org/id/D6009B3AFFFC3B4FFC567EFB0FB0FBFA" lastPageId="10" lastPageNumber="48" pageId="8" pageNumber="46">
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<subSubSection id="16B379A7FFFC3B4DFC567EFB0AA9FD3C" pageId="8" pageNumber="46" type="nomenclature">
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<paragraph id="5E162A2CFFFC3B4DFC567EFB0AA9FD3C" blockId="8.[812,1399,650,1051]" pageId="8" pageNumber="46">
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<taxonomicName id="99A951AFFFFC3B4DFC567EFB0AE6FD3C" ID-CoL="8MCK3" authority="Fedorenko" authorityName="Fedorenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="kuznetsovi" status="sp. nov." subGenus="Vietosteropus">
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<heading id="055E9D40FFFC3B4DFC567EFB0AAAFD5E" box="[951,1259,650,674]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="46" reason="8">
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<emphasis id="6CDDF63EFFFC3B4DFC567EFB0A7DFD5E" box="[951,1084,650,674]" italics="true" pageId="8" pageNumber="46">Pterostichus</emphasis>
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(
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<emphasis id="6CDDF63EFFFC3B4DFBAE7EFA0AA3FD5E" box="[1103,1250,651,674]" italics="true" pageId="8" pageNumber="46">Vietosteropus</emphasis>
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)
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</heading>
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<emphasis id="6CDDF63EFFFC3B4DFC5A7ED90A6BFD3C" box="[955,1066,680,704]" italics="true" pageId="8" pageNumber="46">kuznetsovi</emphasis>
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Fedorenko
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</taxonomicName>
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,
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<emphasis id="6CDDF63EFFFC3B4DFB527ED90AA9FD3C" bold="true" box="[1203,1256,680,704]" pageId="8" pageNumber="46">
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<taxonomicNameLabel id="77EE4B45FFFC3B4DFB527ED90AA9FD3C" box="[1203,1256,680,704]" pageId="8" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="16B379A7FFFC3B4DFC087EB40AFBFD21" box="[1001,1210,709,733]" pageId="8" pageNumber="46" type="description">
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<paragraph id="5E162A2CFFFC3B4DFC087EB40AFBFD21" blockId="8.[812,1399,650,1051]" box="[1001,1210,709,733]" pageId="8" pageNumber="46">
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<figureCitation id="C69236A9FFFC3B4DFC087EB40A25FD21" box="[1001,1124,709,733]" captionStart-0="Figs 23–29" captionStart-1="Figs 30–32" captionStartId-0="6.[226,263,1921,1940]" captionStartId-1="8.[226,263,1969,1988]" captionTargetBox-0="[192,1364,1003,1906]" captionTargetBox-1="[215,1385,1089,1939]" captionTargetId-0="figure-533@6.[189,1369,999,1909]" captionTargetId-1="figure-523@8.[189,1399,1085,1944]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="Figs 23–29. Aedeagus of Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (apical margin slightly damaged); 25, 28 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — median lobe, ventral aspect; 26–28 — inside of right paramere. Scale bars: 1 mm. Рис. 23–29. Эдеагус Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (апикальный край слегка обломан); 25, 28 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — среднЯЯ долЯ? вентрально; 26–28 — праваЯ парамера, иЗнутри. МасШтаб: 1 мм." captionText-1="Figs 30–32. Dorsal habitus, holotypes: 30 — Pterostichus (Vietosteropus) kuznetsovi sp.n.; 31 — Trigonaptus inaequalis gen. et sp.n.; 32 — T. longiscapus sp.n. Рис. 30–32. Габитус дорЗально, голотипы: 30 — Pterostichus (Vietosteropus) kuznetsovi sp.n.; 31 — Trigonaptus inaequalis gen. et sp.n.; 32 — T. longiscapus sp.n." figureDoi-0="http://doi.org/10.5281/zenodo.10978429" figureDoi-1="http://doi.org/10.5281/zenodo.10978431" httpUri-0="https://zenodo.org/record/10978429/files/figure.png" httpUri-1="https://zenodo.org/record/10978431/files/figure.png" pageId="8" pageNumber="46">Figs 29–30</figureCitation>
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,
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<figureCitation id="C69236A9FFFC3B4DFB917EB40AF6FD21" box="[1136,1207,709,733]" captionStart="Figs 33–38" captionStartId="9.[226,262,1921,1940]" captionTargetBox="[193,1394,960,1903]" captionTargetId="figure-543@9.[189,1399,956,1906]" captionTargetPageId="9" captionText="Figs 33–38. Aedeagus of Pterostichus (Vietosteropus) kuznetsovi sp.n.: 33–35 — median lobe; 36–38 — median lobe with everted and inflated internal sac; 33, 37 — left lateral aspect; 34, 38 — right lateral aspect; 35–36 — dorsal aspect; av1, av2+3 — apical vesicles; PBL — left preapical bulb; PBR — right preapical bulb. Scale bars: 1 mm. Рис. 33–38. Эдеагус Pterostichus (Vietosteropus) kuznetsovi sp.n.: 33–35 — среднЯЯ долЯ; 36–38 — среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 33, 37 — слева; 34, 38 — справа; 35–36 — дорЗально; av1, av2+3 — апикальные пуЗырьки; PBL — левый предверШинный пуЗырь; PBR — правый предверШинный пуЗырь. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978435" httpUri="https://zenodo.org/record/10978435/files/figure.png" pageId="8" pageNumber="46">33–38</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="16B379A7FFFC3B4DFCB37E9B0ADAFCB9" pageId="8" pageNumber="46" type="materials_examined">
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<paragraph id="5E162A2CFFFC3B4DFCB37E9B0ADAFCB9" blockId="8.[812,1399,650,1051]" pageId="8" pageNumber="46">
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<materialsCitation id="EEC12071FFFC3B4DFCB37E9B0ADAFCB9" collectingDate="2019-04-23" collectingDateMax="2019-05-11" collectingDateMin="2019-04-23" collectionCode="ZMMU" collectorName="D. Fedorenko'" country="Vietnam" county="Prov" elevation="1010" latitude="15.568611" location="Song Thanh Nat" longLatPrecision="21" longitude="107.38694" municipality="Nam Gian Distr" pageId="8" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Quang Nam" typeStatus="holotype">
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MATERIAL.
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<typeStatus id="8112948EFFFC3B4DFC317E9B0A61FD01" box="[976,1056,746,765]" pageId="8" pageNumber="46" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="38B8B2E9FFFC3B4DFBCE7E9B0A34FD01" box="[1071,1141,746,765]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="8" pageNumber="46" type="Museum">ZMMU</collectionCode>
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) and paratype (SIEE),
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<specimenCount id="48AFE1A5FFFC3B4DFAA87E960B30FD01" box="[1353,1393,743,765]" count="1" pageId="8" pageNumber="46" type="male">♂♂</specimenCount>
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, labelled: ‘
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<collectingCountry id="26BE6ABCFFFC3B4DFC607F730D90FCE9" box="[897,977,770,789]" name="Vietnam" pageId="8" pageNumber="46">Vietnam</collectingCountry>
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,
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<collectingRegion id="9C6DE4CEFFFC3B4DFC3D7F730A07FCE9" box="[988,1094,770,789]" country="Vietnam" name="Quang Nam" pageId="8" pageNumber="46">Quang Nam</collectingRegion>
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Prov[ince].,
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<collectingMunicipality id="BE72B056FFFC3B4DFB5B7F730B0CFCE9" box="[1210,1357,770,789]" pageId="8" pageNumber="46">Nam Gian Distr</collectingMunicipality>
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[ict].,
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||
<location id="5B767CF7FFFC3B4DFCCD7F6B0DFFFCD1" LSID="urn:lsid:plazi:treatment:D6009B3AFFFC3B4FFC567EFB0FB0FBFA:5B767CF7FFFC3B4DFCCD7F6B0DFFFCD1" box="[812,958,794,813]" country="Vietnam" county="Prov" latitude="15.568611" longLatPrecision="21" longitude="107.38694" municipality="Nam Gian Distr" name="Song Thanh Nat" pageId="8" pageNumber="46" stateProvince="Quang Nam">Song Thanh Nat</location>
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[io]n[al]. Park,
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<geoCoordinate id="3B9D4CEBFFFC3B4DFBDF7F6B0AE8FCD1" box="[1086,1193,794,813]" degrees="15" direction="north" minutes="34" orientation="latitude" pageId="8" pageNumber="46" precision="15" seconds="07" value="15.568611">15°34´07´´N</geoCoordinate>
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/
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<geoCoordinate id="3B9D4CEBFFFC3B4DFB547F6B0B6DFCD1" box="[1205,1324,794,813]" degrees="107" direction="east" minutes="23" orientation="longitude" pageId="8" pageNumber="46" precision="15" seconds="13" value="107.38694">107°23´13´´E</geoCoordinate>
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, h=
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<quantity id="995187C9FFFC3B4DFAAA7F6B0D7EFCB9" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="46" unit="m" value="1010.0">
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<elevation id="D584CD1FFFFC3B4DFAAA7F6B0D7EFCB9" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="46" unit="m" value="1010.0">1010 m</elevation>
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</quantity>
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,
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<date id="2A170CECFFFC3B4DFCA87F430DA2FCB9" box="[841,995,818,837]" pageId="8" pageNumber="46" value="2019-04-23" valueMax="2019-05-11" valueMin="2019-04-23">
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<collectingDate id="3A53F504FFFC3B4DFCA87F430DA2FCB9" box="[841,995,818,837]" pageId="8" pageNumber="46" value="2019-04-23" valueMax="2019-05-11" valueMin="2019-04-23">23.IV–11.V.2019</collectingDate>
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</date>
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, leg.
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<collectorName id="F35C4FFAFFFC3B4DFBF57F430AD7FCB9" box="[1044,1174,818,837]" pageId="8" pageNumber="46">D. Fedorenko’</collectorName>
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.
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||
</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="16B379A7FFFC3B4CFCB37F3B0F6CFDF7" lastPageId="9" lastPageNumber="47" pageId="8" pageNumber="46" type="diagnosis">
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<paragraph id="5E162A2CFFFC3B4CFCB37F3B0C9DFE7B" blockId="8.[812,1399,650,1051]" lastBlockId="9.[189,776,315,924]" lastPageId="9" lastPageNumber="47" pageId="8" pageNumber="46">
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DIAGNOSIS. With characters of the subgenus and of the formerly monobasic
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<taxonomicName id="99A951AFFFFC3B4DFC127F140A18FC86" authorityName="Fedorenko" authorityYear="2017" box="[1011,1113,869,890]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="gialaiensis">
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<emphasis id="6CDDF63EFFFC3B4DFC127F140A18FC86" box="[1011,1113,869,890]" italics="true" pageId="8" pageNumber="46">gialaiensis</emphasis>
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||
</taxonomicName>
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-group [
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||
<bibRefCitation id="3A3857DDFFFC3B4DFB447F140B0FFC86" author="Fedorenko D. N." box="[1189,1358,868,891]" pageId="8" pageNumber="46" pagination="211 - 238" refId="ref13316" refString="Fedorenko D. N. 2017. A new subgenus of Pterostichus (Coleoptera: Carabidae: Pterostichini) with 14 new species from Vietnam // Russian Entomol. J. Vol. 26. No. 3. P. 211 - 238." type="journal article" year="2017">
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||
<taxonomicName id="99A951AFFFFC3B4DFB447F140B4EFC87" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1189,1295,869,891]" pageId="8" pageNumber="39" rank="subGenus" status="subgen. nov." subGenus="Fedorenko">Fedorenko</taxonomicName>
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, 2017
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||
</bibRefCitation>
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||
]. Its nominal species,
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<taxonomicName id="99A951AFFFFC3B4DFC2F7FF10A0FFC68" authorityName="Fedorenko" authorityYear="2017" box="[974,1102,895,917]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="gialaiensis">
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<emphasis id="6CDDF63EFFFC3B4DFC2F7FF10A0FFC68" box="[974,1102,895,917]" italics="true" pageId="8" pageNumber="46">P. gialaiensis</emphasis>
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</taxonomicName>
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<bibRefCitation id="3A3857DDFFFC3B4DFBB57F0E0AB7FC68" author="Fedorenko D. N." box="[1108,1270,895,917]" pageId="8" pageNumber="46" pagination="211 - 238" refId="ref13316" refString="Fedorenko D. N. 2017. A new subgenus of Pterostichus (Coleoptera: Carabidae: Pterostichini) with 14 new species from Vietnam // Russian Entomol. J. Vol. 26. No. 3. P. 211 - 238." type="journal article" year="2017">Fedorenko, 2017</bibRefCitation>
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, is distinctive chiefly in having pronotal basal fovea and sulci impunctate and aedeagus distinctive. Additional differences are superficial: pronotal basal angles more rounded apically, explanate lateral margin barely narrower, inner basal sulcus deep. Aedeagus: median lobe with apex widely rounded and projecting laterad beyond everted and inflated internal sac; this latter with
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<collectionCode id="38B8B2E9FFFD3B4CFD387D4D0D47FEAD" box="[729,774,316,337]" pageId="9" pageNumber="47">PBR</collectionCode>
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much larger and directed apicad and
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<collectionCode id="38B8B2E9FFFD3B4CFDFC7D270C08FE97" box="[541,585,342,363]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12690" lsid="urn:lsid:biocol.org:col:12690" name="Botanical Survey of India, Andaman and Nicobar Circle" pageId="9" pageNumber="47" type="Herbarium">PBL</collectionCode>
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slightly different in shape due mainly to vesicles
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<emphasis id="6CDDF63EFFFD3B4CFE357D000FB0FE7A" box="[468,497,369,390]" italics="true" pageId="9" pageNumber="47">lv1</emphasis>
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||
and
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<emphasis id="6CDDF63EFFFD3B4CFDC17D000C7CFE7A" box="[544,573,369,390]" italics="true" pageId="9" pageNumber="47">lv2</emphasis>
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and
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<emphasis id="6CDDF63EFFFD3B4CFD8D7D000CCBFE7A" box="[620,650,369,390]" italics="true" pageId="9" pageNumber="47">lv3</emphasis>
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distinct.
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</paragraph>
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<caption id="0AD67AA4FFFC3B4DFF037BC00FB4F7EC" ID-DOI="http://doi.org/10.5281/zenodo.10978431" ID-Zenodo-Dep="10978431" httpUri="https://zenodo.org/record/10978431/files/figure.png" pageId="8" pageNumber="46" startId="8.[226,263,1969,1988]" targetBox="[215,1385,1089,1939]" targetPageId="8" targetType="figure">
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<paragraph id="5E162A2CFFFC3B4DFF037BC00FE6F820" blockId="8.[189,1398,1969,2064]" pageId="8" pageNumber="46">
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||
Figs 30–32. Dorsal habitus, holotypes: 30 —
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||
<taxonomicName id="99A951AFFFFC3B4DFD8C7BC00D83F838" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[621,962,1969,1988]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="kuznetsovi" status="sp. nov." subGenus="Vietosteropus">
|
||
<emphasis id="6CDDF63EFFFC3B4DFD8C7BC00C99F838" box="[621,728,1969,1988]" italics="true" pageId="8" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFFC3B4DFD047BC30D1AF838" box="[741,859,1970,1988]" italics="true" pageId="8" pageNumber="46">Vietosteropus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFFC3B4DFC887BC00D83F838" box="[873,962,1969,1988]" italics="true" pageId="8" pageNumber="46">kuznetsovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFFC3B4DFC297BC00DB2F838" bold="true" box="[968,1011,1969,1988]" pageId="8" pageNumber="46">
|
||
<taxonomicNameLabel id="77EE4B45FFFC3B4DFC297BC00DB2F838" box="[968,1011,1969,1988]" pageId="8" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 31 —
|
||
<taxonomicName id="99A951AFFFFC3B4DFBD77BC30ABEF838" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1078,1279,1969,1988]" class="Insecta" family="Carabidae" genus="Trigonaptus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="inaequalis" status="gen. et sp. n.">
|
||
<emphasis id="6CDDF63EFFFC3B4DFBD77BC30ABEF838" box="[1078,1279,1969,1988]" italics="true" pageId="8" pageNumber="46">Trigonaptus inaequalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="77EE4B45FFFC3B4DFAE57BC00B31F838" box="[1284,1392,1969,1989]" pageId="8" pageNumber="46" rank="species">
|
||
<emphasis id="6CDDF63EFFFC3B4DFAE57BC00B6BF838" bold="true" box="[1284,1322,1969,1988]" pageId="8" pageNumber="46">gen.</emphasis>
|
||
et
|
||
<emphasis id="6CDDF63EFFFC3B4DFAA47BC00B31F838" bold="true" box="[1349,1392,1969,1988]" pageId="8" pageNumber="46">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 32 —
|
||
<taxonomicName id="99A951AFFFFC3B4DFF177BBB0F34F820" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[246,373,1993,2012]" class="Insecta" family="Carabidae" genus="Trigonaptus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="longiscapus" status="sp.n.">
|
||
<emphasis id="6CDDF63EFFFC3B4DFF177BBB0F34F820" box="[246,373,1993,2012]" italics="true" pageId="8" pageNumber="46">T. longiscapus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFFC3B4DFE9D7BB80FE6F820" bold="true" box="[380,423,1993,2012]" pageId="8" pageNumber="46">
|
||
<taxonomicNameLabel id="77EE4B45FFFC3B4DFE9D7BB80FE6F820" box="[380,423,1993,2012]" pageId="8" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFC3B4DFF037B940FB4F7EC" blockId="8.[189,1398,1969,2064]" pageId="8" pageNumber="46">
|
||
Рис. 30–32. Габитус дорЗально, голотипы: 30 —
|
||
<taxonomicName id="99A951AFFFFC3B4DFD4C7B940A44F804" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[685,1029,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="kuznetsovi" status="sp. nov." subGenus="Vietosteropus">
|
||
<emphasis id="6CDDF63EFFFC3B4DFD4C7B940D59F804" box="[685,792,2021,2040]" italics="true" pageId="8" pageNumber="46">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFFC3B4DFCC67B970DDCF804" box="[807,925,2022,2040]" italics="true" pageId="8" pageNumber="46">Vietosteropus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFFC3B4DFC4D7B940A44F804" box="[940,1029,2021,2040]" italics="true" pageId="8" pageNumber="46">kuznetsovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFFC3B4DFBEC7B940A79F804" bold="true" box="[1037,1080,2021,2040]" pageId="8" pageNumber="46">
|
||
<taxonomicNameLabel id="77EE4B45FFFC3B4DFBEC7B940A79F804" box="[1037,1080,2021,2040]" pageId="8" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 31 —
|
||
<taxonomicName id="99A951AFFFFC3B4DFB9E7B970B0BF804" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1151,1354,2021,2040]" class="Insecta" family="Carabidae" genus="Trigonaptus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="inaequalis" status="gen. et sp. n.">
|
||
<emphasis id="6CDDF63EFFFC3B4DFB9E7B970B0BF804" box="[1151,1354,2021,2040]" italics="true" pageId="8" pageNumber="46">Trigonaptus inaequalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="77EE4B45FFFC3B4DFAB07B940EBFF7EC" pageId="8" pageNumber="46" rank="species">
|
||
<emphasis id="6CDDF63EFFFC3B4DFAB07B940B37F804" bold="true" box="[1361,1398,2021,2040]" pageId="8" pageNumber="46">gen.</emphasis>
|
||
et
|
||
<emphasis id="6CDDF63EFFFC3B4DFF327B8C0EBFF7EC" bold="true" box="[211,254,2045,2064]" pageId="8" pageNumber="46">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 32 —
|
||
<taxonomicName id="99A951AFFFFC3B4DFEA57B8F0F82F7EC" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[324,451,2045,2064]" class="Insecta" family="Carabidae" genus="Trigonaptus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="longiscapus" status="sp.n.">
|
||
<emphasis id="6CDDF63EFFFC3B4DFEA57B8F0F82F7EC" box="[324,451,2045,2064]" italics="true" pageId="8" pageNumber="46">T. longiscapus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="77EE4B45FFFC3B4DFE2B7B8C0FB4F7EC" box="[458,501,2045,2064]" pageId="8" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5E162A2CFFFD3B4CFF037DFA0F6CFDF7" blockId="9.[189,776,315,924]" pageId="9" pageNumber="47">
|
||
Besides, the submentum is bisetose in
|
||
<taxonomicName id="99A951AFFFFD3B4CFDB87DFD0C9FFE5C" authorityName="Fedorenko" authorityYear="2017" box="[601,734,395,417]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="47" phylum="Arthropoda" rank="species" species="gialaiensis">
|
||
<emphasis id="6CDDF63EFFFD3B4CFDB87DFD0C9FFE5C" box="[601,734,395,417]" italics="true" pageId="9" pageNumber="47">P. gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="6CDDF63EFFFD3B4CFD0D7DFD0D41FE5D" box="[748,768,396,417]" italics="true" pageId="9" pageNumber="47">vs</emphasis>
|
||
. quadrisetose in the new species). However, more material is necessary to decide whether this difference is species-specific or it may have come from individual variability of
|
||
<taxonomicName id="99A951AFFFFD3B4CFD137DAD0F66FDF7" authorityName="Fedorenko" authorityYear="2017" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="47" phylum="Arthropoda" rank="species" species="gialaiensis">
|
||
<emphasis id="6CDDF63EFFFD3B4CFD137DAD0F66FDF7" italics="true" pageId="9" pageNumber="47">P. gialaiensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFFD3B4FFF037E600FE9FC9A" lastPageId="10" lastPageNumber="48" pageId="9" pageNumber="47" type="description">
|
||
<paragraph id="5E162A2CFFFD3B4CFF037E600CA6FD1D" blockId="9.[189,776,315,924]" pageId="9" pageNumber="47">
|
||
DESCRIPTION. BL
|
||
<quantity id="995187C9FFFD3B4CFE507E600C6EFDDA" box="[433,559,529,550]" metricMagnitude="-2" metricUnit="m" metricValue="1.705" metricValueMax="1.71" metricValueMin="1.7" pageId="9" pageNumber="47" unit="mm" value="17.05" valueMax="17.1" valueMin="17.0">17–17.1 mm</quantity>
|
||
. Body (
|
||
<figureCitation id="C69236A9FFFD3B4CFD9C7E600C85FDDB" box="[637,708,529,551]" captionStart="Figs 30–32" captionStartId="8.[226,263,1969,1988]" captionTargetBox="[215,1385,1089,1939]" captionTargetId="figure-523@8.[189,1399,1085,1944]" captionTargetPageId="8" captionText="Figs 30–32. Dorsal habitus, holotypes: 30 — Pterostichus (Vietosteropus) kuznetsovi sp.n.; 31 — Trigonaptus inaequalis gen. et sp.n.; 32 — T. longiscapus sp.n. Рис. 30–32. Габитус дорЗально, голотипы: 30 — Pterostichus (Vietosteropus) kuznetsovi sp.n.; 31 — Trigonaptus inaequalis gen. et sp.n.; 32 — T. longiscapus sp.n." figureDoi="http://doi.org/10.5281/zenodo.10978431" httpUri="https://zenodo.org/record/10978431/files/figure.png" pageId="9" pageNumber="47">Fig. 30</figureCitation>
|
||
) shiny black. Antennae brownish toward apices. Dorsal microsculpture absent from head, very superficial on pronotum and elytra, consisting of very small meshes, pronotal meshes being rectangular and moderately transverse, elytral ones more transverse, fusiform, becoming dense transverse lines here and there. Explanate lateral margin of pronotum with hardly traceable microsculpture consisting of slightly longitudinal meshes.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFD3B4CFF037E970CDBFC9B" blockId="9.[189,776,315,924]" pageId="9" pageNumber="47">Head: Eyes convex, gena 0.37 times as long as eye. Frontal sulci very deep impunctate, diverging and slightly S-shaped, nearly parallel just before and behind, reaching the level of anterior supra-ocular seta. Neck constriction indistinct, hardly traceable only laterally. Submentum quadrisetose.</paragraph>
|
||
<paragraph id="5E162A2CFFFD3B4CFF037F1D0B57FD50" blockId="9.[189,776,315,924]" lastBlockId="9.[812,1399,315,924]" pageId="9" pageNumber="47">
|
||
Pronotum subquadrate,
|
||
<collectionCode id="38B8B2E9FFFD3B4CFE257F1A0FA6FC7C" box="[452,487,875,896]" country="Thailand" name="Paleontological Collections" pageId="9" pageNumber="47">PW</collectionCode>
|
||
/
|
||
<collectionCode id="38B8B2E9FFFD3B4CFE0F7F1A0C48FC7C" box="[494,521,875,896]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="9" pageNumber="47" type="Museum">PL</collectionCode>
|
||
1.32,
|
||
<collectionCode id="38B8B2E9FFFD3B4CFDA47F1A0C28FC7C" box="[581,617,875,896]" country="Thailand" name="Paleontological Collections" pageId="9" pageNumber="47">PW</collectionCode>
|
||
/HW 1.55, evenly rounded on sides, less so in basal two fifths, with basal angles obtuse and slightly blunt; basal margin gently concave at middle; apex evenly concave between narrowly rounded apical angles. Explanate lateral margin in form of a wide bead in front of anterolateral seta, otherwise gradually broadening basad and rather wide in basal half. Lateral grooves deep in apical three quarters and almost directly extended into outer basal sulci; these more shallow, slightly C-shaped (convexity inward), obliterate just basally. Inner basal sulci moderately deep, straight, running on and barely converging in basal third. Basal fovea finely and moderately densely punctate in and between basal sulci. Median line moderately deep, almost obliterate near base and apex. Both transverse impressions, basal and apical, imperceptible. Anterolateral seta inserted in lateral groove, basolateral seta close to basal angle.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFD3B4FFCB37EC00C73FE7A" blockId="9.[812,1399,315,924]" lastBlockId="10.[189,775,315,1031]" lastPageId="10" lastPageNumber="48" pageId="9" pageNumber="47">
|
||
Elytra elliptic, broadest behind middle, EW/EL 1.58– 1.62, EW/
|
||
<collectionCode id="38B8B2E9FFFD3B4CFC6E7EBA0DF2FD1C" box="[911,947,715,736]" country="Thailand" name="Paleontological Collections" pageId="9" pageNumber="47">PW</collectionCode>
|
||
1.14–1.15; sides poorly rounded in basal half, preapical sinuation and internal preapical plica distinct, apices truncate combined, with tips blunt. Humeral angle obtuse. Striae deep, densely punctate, punctures becoming increasingly large from very fine in stria 1 to medium-sized in stria 10; parascutellar striole missing. Intervals convex, subcostate to costate laterally and apically; 8
|
||
<superScript id="A9DC8764FFFD3B4CFB5A7F1A0A87FC8B" attach="left" box="[1211,1222,875,887]" fontSize="5" pageId="9" pageNumber="47">th</superScript>
|
||
as wide as 7
|
||
<superScript id="A9DC8764FFFD3B4CFAA07F1A0B0DFC8B" attach="none" box="[1345,1356,875,887]" fontSize="5" pageId="9" pageNumber="47">th</superScript>
|
||
; 7
|
||
<superScript id="A9DC8764FFFD3B4CFA847F1A0B31FC8B" attach="none" box="[1381,1392,875,887]" fontSize="5" pageId="9" pageNumber="47">th</superScript>
|
||
, 5
|
||
<superScript id="A9DC8764FFFD3B4CFCD97FF40D02FC6D" attach="left" box="[824,835,901,913]" fontSize="5" pageId="9" pageNumber="47">th</superScript>
|
||
and 3
|
||
<superScript id="A9DC8764FFFD3B4CFC647FF40DD0FC6D" attach="left" box="[901,913,901,913]" fontSize="5" pageId="9" pageNumber="47">rd</superScript>
|
||
confluent apicad in succession; 1
|
||
<superScript id="A9DC8764FFFD3B4CFB047FF40AAFFC6D" attach="left" box="[1253,1262,901,913]" fontSize="5" pageId="9" pageNumber="47">st</superScript>
|
||
merging into while 3rd separate from lateral bead apically; no costa or carina outside interval 9. Parascutellar seta just behind basal ridge. D2/EL 0.44–0.45. USS: 20–22.
|
||
</paragraph>
|
||
<caption id="0AD67AA4FFFD3B4CFF037BF00A58F7ED" ID-DOI="http://doi.org/10.5281/zenodo.10978435" ID-Zenodo-Dep="10978435" httpUri="https://zenodo.org/record/10978435/files/figure.png" pageId="9" pageNumber="47" startId="9.[226,262,1921,1940]" targetBox="[193,1394,960,1903]" targetPageId="9" targetType="figure">
|
||
<paragraph id="5E162A2CFFFD3B4CFF037BF00D4AF839" blockId="9.[189,1399,1921,2065]" pageId="9" pageNumber="47">
|
||
Figs 33–38. Aedeagus of
|
||
<taxonomicName id="99A951AFFFFD3B4CFE207BF00D52F868" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[449,787,1921,1940]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="47" phylum="Arthropoda" rank="species" species="kuznetsovi" status="sp. nov." subGenus="Vietosteropus">
|
||
<emphasis id="6CDDF63EFFFD3B4CFE207BF00C6BF868" box="[449,554,1921,1940]" italics="true" pageId="9" pageNumber="47">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFFD3B4CFDD87BF30CECF868" box="[569,685,1922,1940]" italics="true" pageId="9" pageNumber="47">Vietosteropus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFFD3B4CFD5A7BF00D52F868" box="[699,787,1921,1940]" italics="true" pageId="9" pageNumber="47">kuznetsovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFFD3B4CFCF87BF00D02F868" bold="true" box="[793,835,1921,1940]" pageId="9" pageNumber="47">
|
||
<taxonomicNameLabel id="77EE4B45FFFD3B4CFCF87BF00D02F868" box="[793,835,1921,1940]" pageId="9" pageNumber="47" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 33–35 — median lobe; 36–38 — median lobe with everted and inflated internal sac; 33, 37 — left lateral aspect; 34, 38 — right lateral aspect; 35–36 — dorsal aspect;
|
||
<emphasis id="6CDDF63EFFFD3B4CFBCC7BE80A0AF850" box="[1069,1099,1945,1964]" italics="true" pageId="9" pageNumber="47">av1</emphasis>
|
||
,
|
||
<emphasis id="6CDDF63EFFFD3B4CFBB67BE80ACFF850" box="[1111,1166,1945,1964]" italics="true" pageId="9" pageNumber="47">av2+3</emphasis>
|
||
— apical vesicles; PBL — left preapical bulb; PBR — right preapical bulb. Scale bars: 1 mm.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFD3B4CFF037BBF0A58F7ED" blockId="9.[189,1399,1921,2065]" pageId="9" pageNumber="47">
|
||
Рис. 33–38. Эдеагус
|
||
<taxonomicName id="99A951AFFFFD3B4CFE437BBC0CBBF81C" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[418,762,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="47" phylum="Arthropoda" rank="species" species="kuznetsovi" status="sp. nov." subGenus="Vietosteropus">
|
||
<emphasis id="6CDDF63EFFFD3B4CFE437BBC0C4FF81C" box="[418,526,1997,2016]" italics="true" pageId="9" pageNumber="47">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFFD3B4CFDFC7BBF0CD2F81C" box="[541,659,1998,2016]" italics="true" pageId="9" pageNumber="47">Vietosteropus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFFD3B4CFD407BBC0CBBF81C" box="[673,762,1997,2016]" italics="true" pageId="9" pageNumber="47">kuznetsovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFFD3B4CFCE37BBC0D6CF81C" bold="true" box="[770,813,1997,2016]" pageId="9" pageNumber="47">
|
||
<taxonomicNameLabel id="77EE4B45FFFD3B4CFCE37BBC0D6CF81C" box="[770,813,1997,2016]" pageId="9" pageNumber="47" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 33–35 — среднЯЯ долЯ; 36–38 — среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 33, 37 — слева; 34, 38 — справа; 35–36 — дорЗально;
|
||
<emphasis id="6CDDF63EFFFD3B4CFC127B940A53F804" box="[1011,1042,2021,2040]" italics="true" pageId="9" pageNumber="47">av1</emphasis>
|
||
,
|
||
<emphasis id="6CDDF63EFFFD3B4CFBFF7B940A17F804" box="[1054,1110,2021,2040]" italics="true" pageId="9" pageNumber="47">av2+3</emphasis>
|
||
— апикальные пуЗырьки; PBL — левый предверШинный пуЗырь; PBR — правый предверШинный пуЗырь. МасШтаб: 1 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5E162A2CFFFE3B4FFF037DFA0D44FDBD" blockId="10.[189,775,315,1031]" pageId="10" pageNumber="48">Underside. Prosternal process truncate apically, in lateral view rectangular or almost so; declivity flat, with lateral edges sharp and faintly beaded in ventral half. Abdomen entirely beaded on sides, except base of sternite II. Mes- and metepisternum moderately and densely punctate, sides of metaventrite with sparse punctures; abdominal sternites II– IV laterally with a few clusters of fine and dense punctures.</paragraph>
|
||
<paragraph id="5E162A2CFFFE3B4FFF037E370EB7FD50" blockId="10.[189,775,315,1031]" pageId="10" pageNumber="48">Legs: metatibia externally glabrous; mesotarsomeres 1–2 sulcate on each side of a conspicuous outer carina, metatarsomeres 1–3 similarly sculptured except that the carina is blunt.</paragraph>
|
||
<paragraph id="5E162A2CFFFE3B4FFF037EC00FE9FC9A" blockId="10.[189,775,315,1031]" pageId="10" pageNumber="48">
|
||
Aedeagus (
|
||
<figureCitation id="C69236A9FFFE3B4FFEAC7EC00FFCFD3B" box="[333,445,689,711]" captionStart="Figs 33–38" captionStartId="9.[226,262,1921,1940]" captionTargetBox="[193,1394,960,1903]" captionTargetId="figure-543@9.[189,1399,956,1906]" captionTargetPageId="9" captionText="Figs 33–38. Aedeagus of Pterostichus (Vietosteropus) kuznetsovi sp.n.: 33–35 — median lobe; 36–38 — median lobe with everted and inflated internal sac; 33, 37 — left lateral aspect; 34, 38 — right lateral aspect; 35–36 — dorsal aspect; av1, av2+3 — apical vesicles; PBL — left preapical bulb; PBR — right preapical bulb. Scale bars: 1 mm. Рис. 33–38. Эдеагус Pterostichus (Vietosteropus) kuznetsovi sp.n.: 33–35 — среднЯЯ долЯ; 36–38 — среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 33, 37 — слева; 34, 38 — справа; 35–36 — дорЗально; av1, av2+3 — апикальные пуЗырьки; PBL — левый предверШинный пуЗырь; PBR — правый предверШинный пуЗырь. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978435" httpUri="https://zenodo.org/record/10978435/files/figure.png" pageId="10" pageNumber="48">Figs 33–38</figureCitation>
|
||
): median lobe with apex triangular and slightly pointed, without right dorso-apical tooth. Right paramere crescent and fairly long. Everted and inflated internal sac: left preapical bulb (
|
||
<collectionCode id="38B8B2E9FFFE3B4FFE347F700C45FCEB" box="[469,516,769,791]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12690" lsid="urn:lsid:biocol.org:col:12690" name="Botanical Survey of India, Andaman and Nicobar Circle" pageId="10" pageNumber="48" type="Herbarium">PBL</collectionCode>
|
||
) entire,
|
||
<emphasis id="6CDDF63EFFFE3B4FFDB87F700C37FCEA" box="[601,630,769,790]" italics="true" pageId="10" pageNumber="48">lv3</emphasis>
|
||
missing; right preapical bulb (
|
||
<collectionCode id="38B8B2E9FFFE3B4FFEB97F6A0FC9FCCD" box="[344,392,795,817]" pageId="10" pageNumber="48">PBR</collectionCode>
|
||
) rather small and set perpendicular to median lobe in dorsal view. Two apical vesicles present, small
|
||
<emphasis id="6CDDF63EFFFE3B4FFF197F200F5AFC9A" box="[248,283,849,870]" italics="true" pageId="10" pageNumber="48">av1</emphasis>
|
||
and
|
||
<emphasis id="6CDDF63EFFFE3B4FFEAD7F200FE3FC9A" box="[332,418,849,870]" italics="true" pageId="10" pageNumber="48">av2+av3</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFFE3B4FFF037F1A0CA0FC4B" pageId="10" pageNumber="48" type="distribution">
|
||
<paragraph id="5E162A2CFFFE3B4FFF037F1A0CB3FC7D" blockId="10.[189,775,315,1031]" box="[226,754,875,897]" pageId="10" pageNumber="48">
|
||
DISTRIBUTION. Only known from the
|
||
<typeStatus id="8112948EFFFE3B4FFD907F1D0CDAFC7D" box="[625,667,876,897]" pageId="10" pageNumber="48">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFE3B4FFF037FF70CA0FC4B" blockId="10.[189,775,315,1031]" pageId="10" pageNumber="48">HABITATS AND HABITS. Both specimens were collected by pitfall traps in a monsoon broad-leaved forest.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFFE3B4FFF037FCA0FB0FBFA" pageId="10" pageNumber="48" type="description">
|
||
<paragraph id="5E162A2CFFFE3B4FFF037FCA0FB0FBFA" blockId="10.[189,775,315,1031]" pageId="10" pageNumber="48">
|
||
<collectionCode id="38B8B2E9FFFE3B4FFF037FCA0F6DFC2C" box="[226,300,955,976]" pageId="10" pageNumber="48">NAME</collectionCode>
|
||
. Patronymic, given after my good friend and colleague, chief of the Joint Russia-Vietnam Tropical Centre, botanist Dr. Andrei Kuznetsov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |