315 lines
30 KiB
XML
315 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="B4D84C038B1817B78379F0396B6234EE" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole ala Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="28B23A79-FBFE-41AB-A4FB-1029E6941339" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777026" LSID="urn:lsid:zoobank.org:act:28B23A79-FBFE-41AB-A4FB-1029E6941339" httpUri="http://treatment.plazi.org/id/B4D84C038B1817B78379F0396B6234EE" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/28B23A79-FBFE-41AB-A4FB-1029E6941339" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ala" status="sp. nov.">Pheidole ala</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">Figs 59A-F</figureCitation>
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84B</figureCitation>
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<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Antananarivo; 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe; -18.47333, 47.96; alt. 1300 m; 5 Dec 2000; B.L. Fisher et al. leg.; BLF02480, CASENT0413606, middle specimen (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •6w., 2s.; same data as for holotype; CASENT0427741, CASENT0427742, CASENT0872080, CASENT0872192-CASENT0872195 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antananarivo</emphasis>
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: •4w., 3s.; 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe; -18.47333, 47.96; alt. 1300 m; 5 Dec 2000; B.L. Fisher et al. leg.; CASENT0413601, CASENT0413608, CASENT0413609, CASENT0427739, CASENT0427740, CASENT0427745, CASENT0427747 (CASC). •3w., 8s.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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<normalizedToken originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
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,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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d Ambohitantely, 20.9 km 72°NE d Ankazobe;
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<geoCoordinate degrees="18.22528" direction="south" orientation="latitude" precision="1" value="-18.22528">-18.22528</geoCoordinate>
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,
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<geoCoordinate degrees="47.28683" direction="east" orientation="longitude" precision="1" value="47.28683">47.28683</geoCoordinate>
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; alt. 1410 m; 17 Apr 2001; B.L. Fisher et al. leg.; CASENT0437257, CASENT0437264, CASENT0437267, CASENT0479788, CASENT0479789, CASENT0479790, CASENT0480418, CASENT0480437 (CASC). •3w., 1q.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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Speciale
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<normalizedToken originalValue="d’Ambohitantely">d'Ambohitantely</normalizedToken>
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;
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<geoCoordinate degrees="18.18762" direction="south" orientation="latitude" precision="1" value="-18.18762">-18.18762</geoCoordinate>
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,
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<geoCoordinate degrees="47.28576" direction="east" orientation="longitude" precision="1" value="47.28576">47.28576</geoCoordinate>
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; alt. 1580 m; 8 Mar 2012; B.L. Fisher et al. leg.; CASENT0274613, CASENT0274614 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •1w., 1s., 1q.; 6.5 km SSW Befingotra,
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<normalizedToken originalValue="Rés">Res</normalizedToken>
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. Anjanaharibe-Sud; -14.75, 49.5; alt. 875 m; 28 Oct 1994; B.L. Fisher et al. leg.; CASENT0198550 (CASC). •1w., 1s., 1q.; 9.2 km WSW Befingotra,
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<normalizedToken originalValue="Rés">Res</normalizedToken>
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. Anjanaharibe-Sud; -14.75, 49.46667; alt. 1200 m; 8 Nov 1994; B.L. Fisher et al. leg.; CASENT0198551 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
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: •1s.; R.S. Ivohibe 8.0 km E Ivohibe;
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<geoCoordinate degrees="22.48333" direction="south" orientation="latitude" precision="1" value="-22.48333">-22.48333</geoCoordinate>
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,
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<geoCoordinate degrees="46.96833" direction="east" orientation="longitude" precision="1" value="46.96833">46.96833</geoCoordinate>
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; alt. 1200 m; 15 Oct 1997; B.L. Fisher et al. leg.; CASENT0198029 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •1s.; Montagne
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<normalizedToken originalValue="d’Anjanaharibe">d'Anjanaharibe</normalizedToken>
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, 19.5 km 27°NNE Ambinanitelo;
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<geoCoordinate degrees="15.17833" direction="south" orientation="latitude" precision="1" value="-15.17833">-15.17833</geoCoordinate>
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,
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<geoCoordinate degrees="49.635" direction="east" orientation="longitude" precision="55" value="49.635">49.635</geoCoordinate>
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; alt. 1100 m; 12 Mar 2003; B.L. Fisher et al. leg.; CASENT0038234 (CASC). •1w., 1s.; Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina;
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<geoCoordinate degrees="17.88667" direction="south" orientation="latitude" precision="1" value="-17.88667">-17.88667</geoCoordinate>
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,
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<geoCoordinate degrees="49.2025" direction="east" orientation="longitude" precision="5" value="49.2025">49.2025</geoCoordinate>
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; alt. 520 m; 2 Dec 2005; B.L. Fisher et al. leg.; CASENT0067738 (CASC). -Toliara: •2w., 1q.; Parc National
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<normalizedToken originalValue="d’Andohahela">d'Andohahela</normalizedToken>
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, Col du Sedro, 3.8 km 113°ESE Mahamavo, 37.6 km 341°NNW Tolagnaro;
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<geoCoordinate degrees="24.76389" direction="south" orientation="latitude" precision="1" value="-24.76389">-24.76389</geoCoordinate>
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,
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<geoCoordinate degrees="46.75167" direction="east" orientation="longitude" precision="1" value="46.75167">46.75167</geoCoordinate>
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; alt. 900 m; 21 Jan 2002; B.L. Fisher et al. leg.; CASENT0430763, CASENT0430764 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full-face view sub-rectangular, anterior and posterior sides of eyes slightly convex; sides of head with sparse, short, erect pilosity; scrobe surface shiny, foveolate with sparse, thick, irregular rugae; closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth slightly thinner and approximately as high as inner hypostomal teeth, with moderately narrow base, triangular; gaster smooth; body bright brown to dark brown.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head foveolate, genae smooth; promesonotum low, short, slightly convex, with posterior declivity smoothly declining towards propodeum; mesosoma foveolate, katepisternum, anepisternum, and mesosoma smooth; propodeal spines minute, triangular; body yellow.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 1.09-1.27 (1.18); HW: 1.03-1.21 (1.11); SL: 0.46-0.53 (0.49); EL: 0.12-0.17 (0.14); WL: 0.85-0.95 (0.9); PSL: 0.16-0.18 (0.17); MTL: 0.46-0.56 (0.51); PNW: 0.55-0.64 (0.59); PTW: 0.13-0.17 (0.15); PPW: 0.34-0.49 (0.44); CI: 91.3-96.0 (94.1); SI: 41.7-47.1 (44.1); PSLI: 13.6-16.1 (14.4); PPI: 31.0-40.8 (35.3); PNI: 51.0-54.6 (53.0); MTI: 44.2-48.0 (45.9).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view longer than wide, anterior and posterior sides of eyes slightly convex (Fig.
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<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming indistinct transverse depression between frons and occipital lobes; inner hypostomal teeth not visible. Sides of the head with sparse, short, erect pilosity; whole head with dense, moderately short, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, foveolate with sparse, thick, irregular rugae. Occipital lobes shiny, with indistinct, sparse, irregular rugae, sculpture weakening posteriorly; frons with dense, thick, and longitudinal rugae, interspaces distinctly to finely rugo-foveolate; genae shiny, with fine rugulae; malar area with dense, thin rugoreticulation. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae present. Scape, when laid back, slightly exceeding the midlength of head; pilosity suberect to erect (Fig.
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<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59B, D</figureCitation>
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). Inner hypostomal teeth distinct, closely spaced, triangular, with rounded apex, and wide base; outer hypostomal teeth slightly thinner and approximately as high as inner hypostomal teeth, and with moderately narrow base, triangular (Fig.
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84B</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, relatively low, and convex, dorsal mesonotum slightly concave, posterior mesonotum steep, with small, tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines small, triangular, with rounded apex; humeral area laterally well produced (Fig.
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<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59D</figureCitation>
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). Surface shiny, with fine but distinct foveolae or rugo-foveolae, and with additional sparse, fine, and irregular rugae; katepisternum smooth; lateral sides of pronotum and propodeum with fading sculpture. Pilosity moderately sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny and with fine foveolae; peduncle short, with indistinct horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shiny, finely shagreened; in dorsal view sides with moderately long, acute, and triangular projections; pilosity long, moderately long, and erect (Fig.
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<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny and smooth; pilosity dense, moderately long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59D, F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Bright brown to brown; legs yellow to bright brown (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1" start="Figure 59" startId="F59">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 59.</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ala</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) of paratype minor worker (CASENT0427742) and full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
) of holotype major worker (CASENT0413606).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.47-0.52 (0.49); HW: 0.42-0.46 (0.44); SL: 0.44-0.48 (0.45); EL: 0.09-0.11 (0.1); WL: 0.58-0.63 (0.6); PSL: 0.07-0.09 (0.08); MTL: 0.34-0.39 (0.37); PNW: 0.28-0.3 (0.28); PTW: 0.07-0.08 (0.07); PPW: 0.1-0.13 (0.12); CI: 86.0-90.2 (88.5); SI: 100.2-108.4 (104.4); PSLI: 14.7-18.2 (16.3); PPI: 53.1-66.1 (60.7); PNI: 62.8-67.9 (65.4); MTI: 80.6 -86.5 (83.8).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
||
<figureCitation captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Pheidole nemoralis Forel, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT 0430850) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT 0460136)." figureDoi="10.3897/zookeys.905.39592.figure58" httpUri="https://binary.pensoft.net/fig/373105" pageId="0" pageNumber="1">58A</figureCitation>
|
||
). Pilosity moderately sparse, moderately long, suberect to erect. Head foveolate; genae smooth. Clypeus with fine and sometimes reduced foveolae; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity erect (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59A, C</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, short, slightly convex, with posterior declivity smoothly declining toward propodeum; promesonotal groove absent; metanotal groove indistinct; propodeal spines minute, triangular, apex acute (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59C</figureCitation>
|
||
). Sculpture foveolate; katepisternum, anepisternum, and mesonotum smooth. Pilosity sparse, long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Peduncle short and thin with ventral face slightly convex; node low, bulge-like, and small; with few short, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Moderately short, low, and slightly convex; with few short, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
With moderately sparse, erect pilosity (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unicolourous, yellow (Fig.
|
||
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole ala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427742) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0413606)." figureDoi="10.3897/zookeys.905.39592.figure59" httpUri="https://binary.pensoft.net/fig/373106" pageId="0" pageNumber="1">59C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Malagasy for forest, in reference to habitat of the species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The species was collected between 520-1410 m in elevation, in rainforest and montane rainforest. Nests were located in rotten logs, and in rotten twigs on the ground.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ala</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName lsidName="P. nemoralis" pageId="0" pageNumber="1" rank="species" species="nemoralis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. nemoralis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ala</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName lsidName="P. nemoralis" pageId="0" pageNumber="1" rank="species" species="nemoralis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. nemoralis</emphasis>
|
||
</taxonomicName>
|
||
in frons with rugo-foveolate sculpture between rugae and never shagreened gaster.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ala</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName lsidName="P. nemoralis" pageId="0" pageNumber="1" rank="species" species="nemoralis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. nemoralis</emphasis>
|
||
</taxonomicName>
|
||
in mesosoma with posterior declivity smoothly declining towards propodeum, smooth genae, and minute propodeal spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |