222 lines
27 KiB
XML
222 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="B1BA5EB37B3D5113BF9C991F6618A2D2" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole sava Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="B7516043-C09C-4573-9A1B-9B649C329C67" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>949</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="165090168" LSID="urn:lsid:zoobank.org:act:B7516043-C09C-4573-9A1B-9B649C329C67" httpUri="http://treatment.plazi.org/id/B1BA5EB37B3D5113BF9C991F6618A2D2" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="B1BA5EB3-7B3D-5113-BF9C-991F6618A2D2" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava" status="sp. nov.">Pheidole sava</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">Figs 52A-F</figureCitation>
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<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64J</figureCitation>
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<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Distribution. Pheidole renirano sp. nov. (A). Pheidole sava sp. nov. (B). Pheidole sikorae Forel (C). Pheidole sofia sp. nov. (D). Pheidole sparsa sp. nov. (E). Pheidole tampony sp. nov. (F). Pheidole trichotos sp. nov. (G). Pheidole tsaravoniana sp. nov. (H). Pheidole vadum sp. nov. (I). Pheidole veteratrix Forel (J). Pheidole volontany sp. nov. (K). Pheidole vony sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure67" httpUri="https://binary.pensoft.net/fig/432236" pageId="0" pageNumber="1">, 67B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Antsiranana; Sava Region: Parc National de Marojejy, Manantenina River, 21.3 km 27.0°NE Andapa;
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<geoCoordinate degrees="14.43686" direction="south" orientation="latitude" precision="1" value="-14.43686">-14.43686</geoCoordinate>
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,
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<geoCoordinate degrees="49.74291" direction="east" orientation="longitude" precision="1" value="49.74291">49.74291</geoCoordinate>
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; alt. 1350 m; 12 Feb 2018; B. L. Fisher et al. leg.; montane rainforest, ex rotten log; BLF41084; CASENT0808425 (CASC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype; CASENT0923296 (CASC).
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</paragraph>
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<caption doi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sava</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923296) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0808425).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: • 1w., 1s.; Sava Region: Parc National de Marojejy, Manantenina River, 21.3 km 27.0°NE Andapa;
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<geoCoordinate degrees="14.43686" direction="south" orientation="latitude" precision="1" value="-14.43686">-14.43686</geoCoordinate>
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,
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<geoCoordinate degrees="49.74291" direction="east" orientation="longitude" precision="1" value="49.74291">49.74291</geoCoordinate>
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; alt. 1350 m; 12 Feb 2018; B. L. Fisher et al. leg.; montane rainforest, ex root mat; BLF41089 (CASC). • 1w., 1s.; Sava Region: Parc National de Marojejy, Manantenina River, 21.3 km 27.0°NE Andapa;
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<geoCoordinate degrees="14.43686" direction="south" orientation="latitude" precision="1" value="-14.43686">-14.43686</geoCoordinate>
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,
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<geoCoordinate degrees="49.74291" direction="east" orientation="longitude" precision="1" value="49.74291">49.74291</geoCoordinate>
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; alt. 1350 m; 12 Feb 2018; B. L. Fisher et al. leg.; montane rainforest, ex rotten log; BLF41097 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with dense, short, suberect pilosity; anterior and medial parts of frons with moderately dense, thick, and longitudinal rugae sometimes interrupted in the posteromedial part, interspaces with sparse and distinct rugulae; posterolateral sides with longitudinal to irregular and thick rugae with distinctly rugulate interspaces; area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by approximately one-fifth of its length; inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity; mesosoma with fine rugofoveolae, dorsal and lateral sides of pronotum with additional transverse and thin rugae and smooth notches; gaster smooth; body brownish orange.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate with additional indistinct longitudinal rugulae on medial frons, area posterolateral from eyes smooth; scape, when laid back, surpassing the posterior head margin by one-fifth of its length; promesonotum moderately high and long, arched; promesonotal groove present; propodeal spines indistinct, triangular; mesosoma smooth and shiny, only promesonotal dorsum with arched rugae and indistinct foveolae, mesonotum and lateral sides of propodeum with indistinct and sparse foveolae; body yellowish brown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 3): HL: 1.14-1.24 (1.2); HW: 1.16-1.23 (1.2); SL: 0.66-0.69 (0.68); EL: 0.13-0.15 (0.14); WL: 1.07-1.1 (1.09); PSL: 0.19-0.21 (0.2); MTL: 0.68-0.73 (0.7); PNW: 0.52-0.53 (0.52); PTW: 0.14-0.17 (0.16); PPW: 0.28-0.34 (0.31); CI: 98.2-101.0 (99.7); SI: 55.5-57.1 (56.4); PSLI: 16.2-16.7 (16.5); PPI: 44.8-61.3 (51.3); PNI: 42.7-45.3 (43.7); MTI: 57.5-59.1 (58.4).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth invisible. Sides of the head with dense, short, suberect pilosity; whole head with relatively dense, short, decumbent to erect pilosity. Anterior and medial parts of frons with moderately dense and thick, longitudinal, and sometimes interrupted rugae in posteromedial part, interspaces with sparse and distinct rugulae; posterolateral sides with longitudinal to irregular and thick rugae with distinctly rugulate interspaces. Occipital lobes with thick, sparse, irregular rugae, interspaces smooth or with fine, indistinct rugulae. Gena with sparse, thick, and longitudinal rugae and smooth interspaces. Area posterolateral from eyes with dense longitudinal to irregular rugofoveolae. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by approximately one-fifth of its length; pilosity decumbent to suberect (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52B, D</figureCitation>
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). Inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by indistinct concavity (Fig.
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<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64J</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderate, thin, with acute apex; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D</figureCitation>
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). Surface shiny with fine rugofoveolae, dorsal and lateral sides of pronotum with additional transverse and thin rugae and smooth notches. Pilosity sparse, moderately long, and erect (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with fine and sparse rugofoveolae; peduncle; node smooth to indistinctly rugulose, low, triangular, with rounded apex, in rear view node dorsoventrally convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny, with fine and sparse rugulae; in dorsal view postpetiole oval, lateral margins medially with two indistinct dentate projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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Shiny and smooth; pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Brownish orange; legs dark yellow, gaster bright brown (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52D, F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 3): HL: 0.58-0.61 (0.59); HW: 0.48-0.49 (0.49); SL: 0.6-0.63 (0.61); EL: 0.11-0.12 (0.11); WL: 0.73-0.74 (0.73); PSL: 0.07-0.09 (0.08); MTL: 0.5-0.52 (0.51); PNW: 0.3-0.34 (0.32); PTW: 0.09-0.1 (0.09); PPW: 0.13-0.16 (0.15); CI: 117.1-127.1 (121.6); SI: 122.0-129.4 (126.2); PSLI: 11.3-14.2 (12.8); PPI: 60.4-65.7 (63.0); PNI: 62.7-70.4 (66.4); MTI: 103.3-106.8 (104.7).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Cephalic margin indistinctly convex (Fig.
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52A</figureCitation>
|
||
). Pilosity relatively sparse, long, decumbent to subdecumbent. Head foveolate with additional indistinct longitudinal rugulae on medial frons, area posterolateral from eyes smooth; antennal sockets with few thick, curved outward rugae and foveolae interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52A, C</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and long, arched; promesonotal groove present but indistinct; metanotal groove present and distinct; propodeal spines indistinct, lobe-like (Fig.
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C</figureCitation>
|
||
). Sculpture smooth and shiny, only promesonotal dorsum with arched rugae and indistinct foveolae, mesonotum and lateral sides of propodeum with indistinct and sparse foveolae. Pilosity sparse, moderately long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Peduncle short and thin with ventral face relatively straight (Fig.
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Brown; legs and lateral sides of pronotum yellowish brown (Fig.
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Pheidole sava sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923296) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0808425)." figureDoi="10.3897/zookeys.949.51269.figure52" httpUri="https://binary.pensoft.net/fig/432221" pageId="0" pageNumber="1">52C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The species was collected at 1350 m in elevation, in montane rainforest. Nests were located in rotten logs and root mat.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sava</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from Parc National de Marojejy in Antsiranana, is most similar to
|
||
<taxonomicName lsidName="P. itremo" pageId="0" pageNumber="1" rank="species" species="itremo">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. itremo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken originalValue="d’Atsirakambiaty">d'Atsirakambiaty</normalizedToken>
|
||
in Fianarantsoa and
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
d Ambohitantely in Antananarivo. Major workers of
|
||
<taxonomicName lsidName="P. sava" pageId="0" pageNumber="1" rank="species" species="sava">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. sava</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be separated based on brighter body colouration, presence of dense, short, suberect pilosity sides of the head, medial frons with sparse and distinctly rugulate interspaces, finely rugofoveolae mesosoma with dorsal and lateral sides of pronotum with additional transverse and thin rugae and smooth notches; minor workers can be separated based on foveolate head with additional indistinct longitudinal rugulae on medial frons, smooth area posterolateral from eyes, predominantly smooth mesosoma with promesonotal dorsum with arched rugae and indistinct foveolae, and indistinct and sparse foveolae on mesonotum and lateral sides of propodeum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |