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<document ID-DOI="http://dx.doi.org/10.3897/aiep.52.91098" ID-Pensoft-Pub="1734-1515-4-267" ID-Pensoft-UUID="7899994C19435AFDAAE6057F1E9466FB" ID-ZooBank="062CD70A849F407F977A451428E8E19B" ModsDocID="1734-1515-52-4-267" checkinTime="1671200936426" checkinUser="pensoft" docAuthor="Imamura, Hisashi, Koeda, Keita &amp; Ho, Hsuan-Ching" docDate="2022" docId="AD1374DABB1053829C5A03EE81E3E7A4" docLanguage="en" docName="ActIchthPisc 52(4): 267-271" docOrigin="Acta Ichthyologica et Piscatoria 52 (4)" docPubDate="2022-12-15" docSource="http://dx.doi.org/10.3897/aiep.52.91098" docTitle="Thysanophrys papillaris Imamura et Knapp 1999" docType="treatment" docVersion="2" id="7899994C19435AFDAAE6057F1E9466FB" lastPageNumber="267" masterDocId="7899994C19435AFDAAE6057F1E9466FB" masterDocTitle="A northward range extension of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) to Taiwan" masterLastPageNumber="271" masterPageNumber="267" pageNumber="267" updateTime="1671201236919" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A northward range extension of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) to Taiwan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Imamura, Hisashi</mods:namePart>
<mods:affiliation>Laboratory of Marine Biology and Biodiversity (Systematic Ichthyology), Faculty of Fisheries Sciences, Hokkaido University, Hakodate, Hokkaido, Japan &amp; Fisheries Science Center, Hokkaido University Museum, Hakodate, Hokkaido, Japan</mods:affiliation>
<mods:nameIdentifier type="email">imamura@fish.hokudai.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Koeda, Keita</mods:namePart>
<mods:affiliation>Faculty of Science, University of the Ryukyus, Nishihara, Okinawa, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ho, Hsuan-Ching</mods:namePart>
<mods:affiliation>National Museum of Marine Biology &amp; Aquarium, Pingtung, Taiwan &amp; Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan &amp; Australian Museum, Sydney, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acta Ichthyologica et Piscatoria</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>52</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>267</mods:start>
<mods:end>271</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/aiep.52.91098</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/aiep.52.91098</mods:identifier>
<mods:identifier type="Pensoft-Pub">1734-1515-4-267</mods:identifier>
<mods:identifier type="ZooBank">062CD70A849F407F977A451428E8E19B</mods:identifier>
<mods:identifier type="Pensoft-UUID">7899994C19435AFDAAE6057F1E9466FB</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4" httpUri="http://treatment.plazi.org/id/AD1374DABB1053829C5A03EE81E3E7A4" lastPageNumber="267" pageId="0" pageNumber="267">
<subSubSection pageId="0" pageNumber="267" type="nomenclature">
<paragraph pageId="0" pageNumber="267">
<taxonomicName LSID="AD1374DA-BB10-5382-9C5A-03EE81E3E7A4" authority="Imamura et Knapp, 1999" authorityName="Imamura et Knapp" authorityYear="1999" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">Thysanophrys papillaris Imamura et Knapp, 1999</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="description">
<paragraph pageId="0" pageNumber="267">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lateral (A), dorsal (B), and ventral (C) views of Thysanophrys papillaris (fresh condition), NMMB-P 28534, 137.3 mm SL, Ke-tzu-liao, Kaohsiung, southern Taiwan." figureDoi="10.3897/aiep.52.91098.figure1" httpUri="https://binary.pensoft.net/fig/782177" pageId="0" pageNumber="267">
English common name: smallknob flathead New Chinese common name:
<normalizedToken originalValue="皮瓣多棘牛尾魚">皮瓣多棘牛尾魚</normalizedToken>
Fig. 1
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="reference_group">
<paragraph pageId="0" pageNumber="267">
<taxonomicName baseAuthorityName="Schultz" baseAuthorityYear="1966" class="Actinopterygii" family="Platycephalidae" genus="Sunagocia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunagocia arenicola" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="arenicola">Sunagocia arenicola</taxonomicName>
(not Schultz, 1966):
<bibRefCitation author="Ho, HC" pageId="0" pageNumber="267" refId="B2" refString="Ho, HC, Chong, LH, 2020. Platycephalidae. Pp. 573-590. In: Koeda K, Ho HC (Eds.) Fishes of southern Taiwan. Volumes I-II. 2nd edn. National Museum of Marine Biology and Aquarium, Pingtung, Taiwan." title="Platycephalidae. Pp. 573 - 590. In: Koeda K, Ho HC (Eds.) Fishes of southern Taiwan. Volumes I-II. 2 nd edn. National Museum of Marine Biology and Aquarium, Pingtung, Taiwan." year="2020">Ho and Chong 2020</bibRefCitation>
: 589, unnumbered fig. (description, Ke-tzu-liao, Kaohsiung, southern Taiwan) (in part).
</paragraph>
<paragraph pageId="0" pageNumber="267">
<taxonomicName authorityName="Imamura et Knapp" authorityYear="1999" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">Thysanophrys papillaris</taxonomicName>
Imamura et Knapp, 1999: 180, figs. 1-4, 5A (original description, type locality: southwest of Flat Top Bank, Timor Sea, Western Australia;
<bibRefCitation author="Hoese, DF" pageId="0" pageNumber="267" refId="B3" refString="Hoese, DF, Bray, DJ, Paxton, JR, Allen, GR, 2006. Platycephalidae. Pp. 933-948. In: Beesley PL, Wellas A (Eds.) Zoological catalogue of Australia. Volume 35. Part 2. Fishes. ABRS and CSIRO Publishing, Collingwood, Vic, Australia." title="Platycephalidae. Pp. 933 - 948. In: Beesley PL, Wellas A (Eds.) Zoological catalogue of Australia. Volume 35. Part 2. Fishes. ABRS and CSIRO Publishing, Collingwood, Vic, Australia." year="2006">Hoese et al. (2006</bibRefCitation>
: 947) (list, northern Australia);
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3696.1.1" author="Larson, HK" journalOrPublisher="Zootaxa" pageId="0" pageNumber="267" pagination="1 - 293" refId="B10" refString="Larson, HK, Williams, RS, Hammer, MP, 2013. An annotated checklist of the fishes of the Northern Territory, Australia. Zootaxa 3696 (1): 1 - 293, DOI: https://doi.org/10.11646/zootaxa.3696.1.1" title="An annotated checklist of the fishes of the Northern Territory, Australia." url="https://doi.org/10.11646/zootaxa.3696.1.1" volume="3696" year="2013">Larson et al. (2013</bibRefCitation>
: 93) (list, northern Australia);
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.24.17" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="17 - 22" refId="B7" refString="Imamura, H, Kimura, K, Quan, NV, 2019. First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific. Species Diversity 24 (1): 17 - 22, DOI: https://doi.org/10.12782/specdiv.24.17" title="First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific." url="https://doi.org/10.12782/specdiv.24.17" volume="24" year="2019">Imamura et al. (2019</bibRefCitation>
: 17, figs. 1, 2) (description, Nha Trang, Vietnam, western Pacific Ocean).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="materials_examined">
<paragraph pageId="0" pageNumber="267">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="267">
<materialsCitation collectingDate="2017-03-11" collectingDateMax="2018-02-18" collectingDateMin="2017-03-11" country="Taiwan" location="Dong-gang" specimenCount="5" stateProvince="Pingtung">
Six specimens: NMMB-P26824 (
<specimenCount type="generic">5 specimens</specimenCount>
,
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.2375" metricValueMax="1.4819999999999998" metricValueMin="0.993" unit="mm" value="123.75" valueMax="148.2" valueMin="99.3">99.3-148.2 mm</quantity>
SL), off
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:DC1654FB4A22BDDA4ECBA73C1BB77996" country="Taiwan" name="Dong-gang" stateProvince="Pingtung">Dong-gang</location>
,
<collectingRegion country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, southern
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
(ca. 22°28ʹN, 120°20ʹE),
<collectingDate value="2017-03-11">11 March 2017</collectingDate>
, bottom trawl; NMMB-P28534 (
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.373" unit="mm" value="137.3">137.3 mm</quantity>
SL), off Ke-tzu-liao,
<collectingRegion country="Taiwan" name="Kaohsiung">Kaohsiung</collectingRegion>
, southern
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
(ca. 22°43ʹN, 120°15ʹE),
<collectingDate value="2018-02-18">18 February 2018</collectingDate>
, bottom trawl
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="materials_examined">
<paragraph pageId="0" pageNumber="267">Comparative material.</paragraph>
<paragraph pageId="0" pageNumber="267">
<materialsCitation collectingDate="1990-12-08" elevation="98" location="Timor Sea" municipality="Flat Top Bank" specimenCount="1" stateProvince="Western Australia" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
: NTM S. 13325-001 (
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.217" unit="mm" value="121.7">121.7 mm</quantity>
SL), southwest of
<collectingMunicipality>Flat Top Bank</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:8BD7CE2373F2B8BB378F16F63FDAFA59" municipality="Flat Top Bank" name="Timor Sea" stateProvince="Western Australia">Timor Sea</location>
,
<collectingRegion country="Australia" name="Western Australia">Western Australia</collectingRegion>
(12°28ʹS, 128°37ʹE),
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.8" unit="m" value="98.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.8" unit="m" value="98.0">98 m</quantity>
depth
</elevation>
,
<collectingDate value="1990-12-08">8 December 1990</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="1990-11-10" country="Australia" elevation="64" location="Arafura Sea" specimenCount="5" stateProvince="Northern Territory" typeStatus="Paratypes">
<typeStatus>Paratypes</typeStatus>
(
<specimenCount type="generic">5 specimens</specimenCount>
): NTM S. 12912-008 (
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.013" unit="mm" value="101.3">101.3 mm</quantity>
SL),
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:93DEBD0F8773B6F0B56B9B00D7E0F28D" country="Australia" name="Arafura Sea" stateProvince="Northern Territory">Arafura Sea</location>
,
<collectingRegion country="Australia" name="Northern Territory">Northern Territory</collectingRegion>
,
<collectingCountry name="Australia">Australia</collectingCountry>
(10°15ʹS, 134°42ʹE),
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.4" unit="m" value="64.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.4" unit="m" value="64.0">64 m</quantity>
depth
</elevation>
,
<collectingDate value="1990-11-10">10 Nov. 1990</collectingDate>
</materialsCitation>
;
<materialsCitation collectingDate="1963-03-30" country="Australia" elevation="77" location="Indian Ocean" municipality="Andaman Sea" specimenCount="2" stateProvince="Northern Territory" typeStatus="paratype">
USNM 344833 (
<specimenCount type="generic">2 specimens</specimenCount>
,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="8.86" metricValueMax="9.94" metricValueMin="7.779999999999999" unit="mm" value="88.6" valueMax="99.4" valueMin="77.8">77.8-99.4 mm</quantity>
SL),
<collectingMunicipality>Andaman Sea</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:75B9CD8ADB71C5A79D6628FE44FCCAF4" country="Australia" municipality="Andaman Sea" name="Indian Ocean" stateProvince="Northern Territory">Indian Ocean</location>
(13°28ʹN, 97°19ʹE),
<elevation metricMagnitude="1" metricUnit="m" metricValue="7.7" metricValueMax="8.1" metricValueMin="7.3" unit="m" value="77.0" valueMax="81.0" valueMin="73.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="7.7" metricValueMax="8.1" metricValueMin="7.3" unit="m" value="77.0" valueMax="81.0" valueMin="73.0">73-81 m</quantity>
depth
</elevation>
,
<collectingDate value="1963-03-30">30 March 1963</collectingDate>
</materialsCitation>
;
<materialsCitation collectingDate="1963-03-30" country="Australia" elevation="71" location="Indian Ocean" municipality="Andaman Sea" specimenCount="2" stateProvince="Northern Territory" typeStatus="paratype">
USNM 344834 (
<specimenCount type="generic">2 specimens</specimenCount>
,
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.063" metricValueMax="1.133" metricValueMin="0.993" unit="mm" value="106.3" valueMax="113.3" valueMin="99.3">99.3-113.3 mm</quantity>
SL),
<collectingMunicipality>Andaman Sea</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:F4FB9E84ACDE118E4C87B274AB3D33C4" country="Australia" municipality="Andaman Sea" name="Indian Ocean" stateProvince="Northern Territory">Indian Ocean</location>
(14°07ʹN, 97°05ʹE),
<elevation metricMagnitude="1" metricUnit="m" metricValue="7.1" metricValueMax="7.3" metricValueMin="6.9" unit="m" value="71.0" valueMax="73.0" valueMin="69.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="7.1" metricValueMax="7.3" metricValueMin="6.9" unit="m" value="71.0" valueMax="73.0" valueMin="69.0">69-73 m</quantity>
depth
</elevation>
,
<collectingDate value="1963-03-30">30 March 1963</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="1987-09-27" collectorName="All, V Anton Bruun." country="Australia" elevation="94" location="Soela" municipality="Port Hedland" specimenCount="7" stateProvince="Western Australia" typeStatus="paratypes">
<collectorName>All</collectorName>
USNM
<typeStatus>paratypes</typeStatus>
were trawled by R/
<collectorName>
V
<emphasis italics="true" pageId="0" pageNumber="267">Anton Bruun</emphasis>
.
</collectorName>
Additional non-types (
<specimenCount type="generic">5 specimens</specimenCount>
): CSIRO H1037-1 (
<specimenCount type="generic">2 specimens</specimenCount>
,
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.3705000000000003" metricValueMax="1.4969999999999999" metricValueMin="1.2440000000000002" unit="mm" value="137.05" valueMax="149.7" valueMin="124.4">124.4-149.7 mm</quantity>
SL), north-northwest of
<collectingMunicipality>Port Hedland</collectingMunicipality>
,
<collectingRegion country="Australia" name="Western Australia">Western Australia</collectingRegion>
(19°03ʹS, 117°59ʹE - 19°04ʹS, 118°01ʹE),
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.35" metricValueMax="9.7" metricValueMin="9.0" unit="m" value="93.5" valueMax="97.0" valueMin="90.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.35" metricValueMax="9.7" metricValueMin="9.0" unit="m" value="93.5" valueMax="97.0" valueMin="90.0">90-97 m</quantity>
depth
</elevation>
, FRV
<emphasis italics="true" pageId="0" pageNumber="267">
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:6B5D130F9D678768C8EF9B4A2B23935C" country="Australia" municipality="Port Hedland" name="Soela" stateProvince="Western Australia">Soela</location>
</emphasis>
, demersal trawl,
<collectingDate value="1987-09-27">27 September 1987</collectingDate>
</materialsCitation>
;
<materialsCitation collectingDate="1988-10-03" country="Australia" elevation="86" location="Soela" municipality="Port Hedland" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
CSIRO H1500-01 (
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.247" unit="mm" value="124.7">124.7 mm</quantity>
SL), north-northwest of
<collectingMunicipality>Port Hedland</collectingMunicipality>
,
<collectingRegion country="Australia" name="Western Australia">Western Australia</collectingRegion>
(19°06ʹS, 118°12ʹE - 19°05ʹS, 118°10ʹE),
<elevation metricMagnitude="1" metricUnit="m" metricValue="8.55" metricValueMax="8.7" metricValueMin="8.4" unit="m" value="85.5" valueMax="87.0" valueMin="84.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="8.55" metricValueMax="8.7" metricValueMin="8.4" unit="m" value="85.5" valueMax="87.0" valueMin="84.0">84-87 m</quantity>
depth
</elevation>
, FRV
<emphasis italics="true" pageId="0" pageNumber="267">
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:A1E7A54E8D9F85EDE7C48DDF5C896038" country="Australia" municipality="Port Hedland" name="Soela" stateProvince="Western Australia">Soela</location>
</emphasis>
, demersal trawl,
<collectingDate value="1988-10-03">3 October 1988</collectingDate>
</materialsCitation>
;
<materialsCitation collectingDate="2018-07-25" country="Vietnam" location="Nha Trang" municipality="Cua Be" specimenCount="1" typeStatus="paratype">
HUMZ
<quantity metricMagnitude="5" metricUnit="m" metricValue="2.297211703" unit="mm" value="2.297211703E8">229721, 170.3 mm</quantity>
SL, fish landing port,
<collectingMunicipality>Cua Be</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:10D3B5546293AC8BB2182EBF5579D0B3" country="Vietnam" municipality="Cua Be" name="Nha Trang">Nha Trang</location>
,
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
(12°11ʹN, 109°12ʹE),
<collectingDate value="2018-07-25">25 July 2018</collectingDate>
</materialsCitation>
,
<materialsCitation collectingDate="2016-10-30" collectorName="N. V. Quan" country="Vietnam" elevation="64" location="Lynher Bank" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
coll. by
<collectorName>N. V. Quan</collectorName>
and party; WAM P.34804-005 (
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" unit="mm" value="112.0">112 mm</quantity>
SL),
<location LSID="urn:lsid:plazi:treatment:AD1374DABB1053829C5A03EE81E3E7A4:D4F7E9218CE24258A397EB22338B37EB" country="Vietnam" name="Lynher Bank" stateProvince="Western Australia">Lynher Bank</location>
,
<collectingRegion inferredBySuffix="true">Western Australia</collectingRegion>
(15°32.69ʹS, 122°12.43ʹE - 15°32.65ʹS, 122°12.45ʹE),
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.375" metricValueMax="6.43" metricValueMin="6.32" unit="m" value="63.75" valueMax="64.3" valueMin="63.2">
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.375" metricValueMax="6.43" metricValueMin="6.32" unit="m" value="63.75" valueMax="64.3" valueMin="63.2">63.2-64.3 m</quantity>
depth
</elevation>
,
<collectingDate value="2016-10-30">30 October 2016</collectingDate>
</materialsCitation>
, coll. by M.G. Allen.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="description">
<paragraph pageId="0" pageNumber="267">Description.</paragraph>
<paragraph pageId="0" pageNumber="267">
Counts and proportional measurements are given in Tables
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Meristic characters of Thysanophrys papillaris." httpUri="http://table.plazi.org/id/5E5D7055ACB1F562ADDB877B95886A62" pageId="0" pageNumber="267" tableUuid="5E5D7055ACB1F562ADDB877B95886A62">1</tableCitation>
and
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Absolute and relative morphometric characters of Thysanophrys papillaris." httpUri="http://table.plazi.org/id/B135845AE3A8A3787F75E4F7F4A6A07A" pageId="0" pageNumber="267" tableUuid="B135845AE3A8A3787F75E4F7F4A6A07A">2</tableCitation>
, respectively. The following description was based on the six Taiwanese specimens. Body moderately depressed; caudal peduncle somewhat cylindrical. Head strongly depressed. Snout longer than orbit diameter; ratio of snout length / orbit diameter 1.1-1.2. Posterior end of upper jaw nearly or just reaching below anterior margin of pupil. Villiform teeth in bands on jaws and palatine, in 2 separate patches on vomer, except for anteromedial portion of upper jaw with short slender conical teeth. Notch absent on posterior margin of upper jaw tooth band. Anterior and posterior nostrils tubular; former with single flap on posterior margin. Single papilla on upper surface of eye; additional papilla present posteriorly and posteromedially to former on right side eye in 99.3 mm SL specimen of NMMB-P26824 and right side eye in NMMB-P28534, respectively. Upper iris lappet with short branches; lower iris lappet simple in 5 specimens, bilobed in left eye in 99.3 mm SL specimen of NMMB-P26824. Interorbital space narrow, concave. Dorsal surface of head with many spines but lacking tubercles. Two antrorse lachrymal spines present. Nasal with 1 small spine. Ethmoid spines absent in 5 specimens, 1 spine present on left side in 147.4 mm SL specimen of NMMB-P26824. Single preorbital spine present. Single preocular spine present, lacking small spines on base. Supraorbital ridge finely serrated. Single postocular spine, followed by 2 or 3 pterotic spines. One or two stout parietal spines present. Two to six frontal spines present between posteriormost supraorbital spine and parietal spine. One or two nuchal spines present posterior to parietal spine. Suborbital ridge with single preorbital spine anteriorly, followed by 4-6 stout suborbital spines. Three preopercular spines present; uppermost longest, not reaching posterior margin of opercle, with supplemental spine on base. Two stout spines on opercle, each with short supporting ridge and without serrations or tubercles. Supratemporal, posttemporal and supracleithrum each with single stout spine. Well-developed fleshy sensory tubes completely covering cheek region. Interopercular flap absent. Small ctenoid scales on postorbital area and opercle. Scales on body ctenoid dorsally and laterally, cycloid ventrally. Lateral-line scales each with 2 sensory ducts directed upward and downward, respectively; each duct terminating with single external opening. First dorsal fin originating nearly level with opercular margin, narrowly separated from second dorsal fin. Posterior margin of pectoral fin rounded. Tip of pelvic fin reaching anus in 147.4 mm SL specimen of NMMB-P26824, to base of second ray of anal fin in remaining specimens. Caudal fin slightly rounded posteriorly.
</paragraph>
<caption ID-Table-UUID="5E5D7055ACB1F562ADDB877B95886A62" httpUri="http://table.plazi.org/id/5E5D7055ACB1F562ADDB877B95886A62" pageId="0" pageNumber="267" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="267">
<emphasis bold="true" pageId="0" pageNumber="267">Table 1.</emphasis>
Meristic characters of
<taxonomicName authorityName="Imamura et Knapp" authorityYear="1999" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys papillaris</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="267">
<table pageId="0" pageNumber="267">
<tr pageId="0" pageNumber="267">
<th colspan="1" pageId="0" pageNumber="267" rowspan="2">Character</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Present specimens</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Holotype</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Paratypes</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Additional non-types</th>
</tr>
<tr pageId="0" pageNumber="267">
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">
(
<emphasis italics="true" pageId="0" pageNumber="267">n</emphasis>
= 6)
</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">NTM S. 13325-001</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">
(
<emphasis italics="true" pageId="0" pageNumber="267">n</emphasis>
= 5)
</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">
(
<emphasis italics="true" pageId="0" pageNumber="267">n</emphasis>
= 5)
</th>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">First dorsal-fin rays</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">I + VII-VIII</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">I + VIII</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">I + VI-VII</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">I + VII-VIII</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Second dorsal-fin rays</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">11</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Anal-fin rays</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">12</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Pectoral-fin rays</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">2 + 12-14 + 4-6 = 19-20</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">2 + 12 + 6 = 20</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1-2 + 10-13 + 5-8 = 19-20 (4)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">2 + 12-15 + 3-6 = 20</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Branched caudal-fin rays</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">10 (2)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">10 (4)</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Pored lateral-line scales (LLS)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52-54</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">51</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52-54</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52-53</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Anterior LLS with spine</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">2-3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">2-3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">2-3</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">SADB</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52-54</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">51</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">54 (1)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52-53</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">SADF</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52-55</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">51</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52 (1)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52-54 (4)</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">SBDF</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">59-74 (5)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">69</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">58-71 (4)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">58-75 (3)</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Gill rakers</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1 + 5-6 = 6-7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1 + 4 = 5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1 + 5-6 = 6-7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1 + 5 = 6</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="267">
<tableNote pageId="0" pageNumber="267">Pectoral fin rays are expressed as &quot;upper unbranched rays + middle branched rays + lower unbranched rays = total rays&quot;; SADB = Scale rows above lateral line slanting downward and backward; SADF = Scale rows above lateral line slanting downward and forward; SBDF = Scale rows below lateral line slanting downward and forward. Numbers in parentheses indicate the number of specimens examined.</tableNote>
</paragraph>
<caption ID-Table-UUID="B135845AE3A8A3787F75E4F7F4A6A07A" httpUri="http://table.plazi.org/id/B135845AE3A8A3787F75E4F7F4A6A07A" pageId="0" pageNumber="267" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="267">
<emphasis bold="true" pageId="0" pageNumber="267">Table 2.</emphasis>
Absolute and relative morphometric characters of
<taxonomicName authorityName="Imamura et Knapp" authorityYear="1999" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys papillaris</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="267">
<table pageId="0" pageNumber="267">
<tr pageId="0" pageNumber="267">
<th colspan="1" pageId="0" pageNumber="267" rowspan="2">Character</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Present specimens</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Holotype</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Paratypes</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">Additional non-types</th>
</tr>
<tr pageId="0" pageNumber="267">
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">
(
<emphasis italics="true" pageId="0" pageNumber="267">n</emphasis>
= 6)
</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">NTM S. 13325-001</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">
(
<emphasis italics="true" pageId="0" pageNumber="267">n</emphasis>
= 5)
</th>
<th colspan="1" pageId="0" pageNumber="267" rowspan="1">
(
<emphasis italics="true" pageId="0" pageNumber="267">n</emphasis>
= 5)
</th>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="5" pageId="0" pageNumber="267" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="267">Absolute values [mm</emphasis>
]
</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Standard length (SL)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">99.3-148.2</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">120.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">77.8-113.3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">111.9-170.3</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="5" pageId="0" pageNumber="267" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="267">Relative values [%SL</emphasis>
]
</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Head length (HL)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.3-37.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">38.4</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.0-39.4</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.2-38.5</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Predorsal length</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.6-37.4</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">38.4</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.0-39.3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.5-39.5</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Length of first dorsal-fin base</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">16.5-19.8</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">17.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">13.6-16.6</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">16.2-18.6</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Length of second dorsal-fin base</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">25.1-26.0</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">24.3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">24.4-26.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">25.3-28.0</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Length of anal-fin base</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">29.7-31.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">28.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">30.0-32.8</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">29.5-31.9</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Snout length</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">10.9-11.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">11.4</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">11.3-12.2</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">10.5-12.2</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Orbital diameter</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">9.4-9.9</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">9.9</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">8.6-10.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">9.4-10.1</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Upper jaw</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">13.1-14.0</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">14.0</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">12.9-15.1</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">13.5-13.7 (3)</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Lower jaw</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">20.4-20.9</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">20.9</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">20.9-22.0 (4)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">20.1-20.6</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Interorbital width</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.3-1.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.4-1.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.4-1.9</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Pectoral-fin length</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">14.8-16.1</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">15.6</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">14.1-16.8</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">14.9-16.4</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Pelvic-fin length</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">21.3-24.6</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">24.0</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">22.9-24.6</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">22.2-26.0</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Caudal-fin length</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">20.4-22.3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">21.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">22.1-22.6 (3)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">20.6-22.7 (4)</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="5" pageId="0" pageNumber="267" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="267">Relative values [%HL</emphasis>
]
</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Snout length</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">29.8-31.8</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">29.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">29.4-31.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">28.7-31.8</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Orbital diameter</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">25.6-26.8</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">25.6</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">23.8-27.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">25.1-26.8</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Upper jaw</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">35.9-37.8</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.4</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">35.9-38.4</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">36.2-36.8 (3)</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Lower jaw</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">55.3-56.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">54.3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">53.1-57.2 (4)</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">52.3-55.5</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Interorbital width</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">3.5-4.7</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">4.5</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">3.7-4.0</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">3.8-5.1</td>
</tr>
<tr pageId="0" pageNumber="267">
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">Ratio of snout length / orbital diameter</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.1-1.2</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.1-1.3</td>
<td colspan="1" pageId="0" pageNumber="267" rowspan="1">1.1-1.3</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="267">
<tableNote pageId="0" pageNumber="267">Numbers in parentheses indicate the number of specimens examined.</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="color when fresh">
<paragraph pageId="0" pageNumber="267">Color when fresh</paragraph>
<paragraph pageId="0" pageNumber="267">
[based on color photographs of NMMB-P28534 (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lateral (A), dorsal (B), and ventral (C) views of Thysanophrys papillaris (fresh condition), NMMB-P 28534, 137.3 mm SL, Ke-tzu-liao, Kaohsiung, southern Taiwan." figureDoi="10.3897/aiep.52.91098.figure1" httpUri="https://binary.pensoft.net/fig/782177" pageId="0" pageNumber="267">1</figureCitation>
)]. Dorsal surface of body and head pale brown, ventral surface white. Two brownish bands crossing interorbit and both eyes; several small brown or dark brown blotches laterally on head, including single dark brown blotch posteriorly on preopercle. Dorsal surface of body with four irregular dark brown bands, level with posterior portion of first dorsal fin, middle and posterior portions of second dorsal fin, and caudal peduncle, respectively. First dorsal fin with broad blackish band distally; spines with several brown spots. Second dorsal fin with brown spots along rays. Anal fin pale (coloration details unclear). Pectoral and pelvic fins pale brown with small brown spots tending to form irregular narrow bands. Anteroventral portion of pectoral fin with elongate dark brown spot. Basal portion of caudal fin dark brown; remaining area of caudal fin with four irregular dark brown bands.
</paragraph>
<caption doi="10.3897/aiep.52.91098.figure1" httpUri="https://binary.pensoft.net/fig/782177" pageId="0" pageNumber="267" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="267">
<emphasis bold="true" pageId="0" pageNumber="267">Figure 1.</emphasis>
Lateral (
<emphasis bold="true" pageId="0" pageNumber="267">A</emphasis>
), dorsal (
<emphasis bold="true" pageId="0" pageNumber="267">B</emphasis>
), and ventral (
<emphasis bold="true" pageId="0" pageNumber="267">C</emphasis>
) views of
<taxonomicName authorityName="Imamura et Knapp" authorityYear="1999" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys papillaris</emphasis>
</taxonomicName>
(fresh condition), NMMB-P28534, 137.3 mm SL, Ke-tzu-liao, Kaohsiung, southern Taiwan.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="color in alcohol">
<paragraph pageId="0" pageNumber="267">Color in alcohol.</paragraph>
<paragraph pageId="0" pageNumber="267">Dorsal surface of body and head pale brown, ventral surface pale yellow. One or two brownish bands crossing interorbit and both eyes; several small brown or dark brown blotches laterally on head, including a single dark brown blotch posteriorly on preopercle. Dorsal surface of body with four irregular brown or dark brown bands, level with posterior portion of first dorsal fin, middle and posterior portions of second dorsal fin, and caudal peduncle, respectively. First dorsal fin with broad blackish band distally; spines with several brown spots. Second dorsal fin with brown or dark brown spots along rays. Anal fin pale; posterior 2-10 rays with a brown spot distally. Pectoral and pelvic fins pale brown with small brown spots tending to form irregular narrow bands. Anteroventral portion of pectoral fin with elongate dark brown spot. Basal portion of caudal fin dark brown; remaining area of caudal fin with several irregular brown to dark brown bands.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="distribution">
<paragraph pageId="0" pageNumber="267">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="267">
Recorded from the Andaman, Timor, and Arafura seas (eastern Indian Ocean) (
<bibRefCitation DOI="https://doi.org/10.1007/BF02675436" author="Imamura, H" journalOrPublisher="Ichthyological Research" pageId="0" pageNumber="267" pagination="179 - 183" refId="B6" refString="Imamura, H, Knapp, LW, 1999. Thysanophrys papillaris, a new species of flathead from the Andaman Sea and northern Australia (Scorpaeniformes: Platycephalidae). Ichthyological Research 46 (2): 179 - 183, DOI: https://doi.org/10.1007/BF02675436" title="Thysanophrys papillaris, a new species of flathead from the Andaman Sea and northern Australia (Scorpaeniformes: Platycephalidae)." url="https://doi.org/10.1007/BF02675436" volume="46" year="1999">Imamura and Knapp 1999</bibRefCitation>
), and Nha Trang, Vietnam (
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.24.17" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="17 - 22" refId="B7" refString="Imamura, H, Kimura, K, Quan, NV, 2019. First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific. Species Diversity 24 (1): 17 - 22, DOI: https://doi.org/10.12782/specdiv.24.17" title="First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific." url="https://doi.org/10.12782/specdiv.24.17" volume="24" year="2019">Imamura et al. 2019</bibRefCitation>
) and southern Taiwan (western Pacific Ocean) (this study).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="267" type="remarks">
<paragraph pageId="0" pageNumber="267">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="267">
The presently reported six specimens, collected from southern Taiwan, conformed to the genus
<taxonomicName authorityName="J.D.Ogilby" authorityYear="1898" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys</emphasis>
</taxonomicName>
Ogilby, 1898 in having the upper iris lappet with short branches, many spines on the dorsal surface of the head (but lacking tubercles), the suborbital ridge with 5 or 6 spines (including 1 preorbital spine), the lateral-line scales each with 2 sensory ducts (directed upward and downward, respectively, and terminating in a single external opening), and well-developed fleshy sensory tubes completely covering the cheek region, and lacking an interopercular flap (
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.1.123" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="123 - 233" refId="B4" refString="Imamura, H, 1996. Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes). Species Diversity 1 (2): 123 - 233, DOI: https://doi.org/10.12782/specdiv.1.123" title="Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes)." url="https://doi.org/10.12782/specdiv.1.123" volume="1" year="1996">Imamura 1996</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.24.17" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="17 - 22" refId="B7" refString="Imamura, H, Kimura, K, Quan, NV, 2019. First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific. Species Diversity 24 (1): 17 - 22, DOI: https://doi.org/10.12782/specdiv.24.17" title="First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific." url="https://doi.org/10.12782/specdiv.24.17" volume="24" year="2019">Imamura et al. 2019</bibRefCitation>
). To date, the genus includes the following 10 valid species:
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1938" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys armata" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="armata">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys armata</emphasis>
</taxonomicName>
(Fowler, 1938);
<taxonomicName baseAuthorityName="Bleeker" baseAuthorityYear="1855" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys celebica" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="celebica">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys celebica</emphasis>
</taxonomicName>
(Bleeker, 1855);
<taxonomicName authorityName="Schultz" authorityYear="1966" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys chiltonae" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="chiltonae">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys chiltonae</emphasis>
</taxonomicName>
Schultz, 1966;
<taxonomicName baseAuthorityName="Richardson" baseAuthorityYear="1848" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys cirronasa" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="cirronasa">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys cirronasa</emphasis>
</taxonomicName>
(Richardson, 1848);
<taxonomicName baseAuthorityName="Shao &amp; Chen" baseAuthorityYear="1987" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys longirostris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="longirostris">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys longirostris</emphasis>
</taxonomicName>
Shao et Chen, 1987;
<taxonomicName class="Pisces" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">T. papillaris</emphasis>
</taxonomicName>
;
<taxonomicName authorityName="Knapp" authorityYear="2013" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys randalli" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="randalli">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys randalli</emphasis>
</taxonomicName>
Knapp, 2013;
<taxonomicName authorityName="Knapp" authorityYear="2013" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys rarita" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="rarita">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys rarita</emphasis>
</taxonomicName>
Knapp, 2013;
<taxonomicName authorityName="Knapp" authorityYear="2013" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys springeri" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="springeri">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys springeri</emphasis>
</taxonomicName>
Knapp, 2013; and
<taxonomicName authorityName="Knapp" authorityYear="2013" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys tricaudata" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="tricaudata">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys tricaudata</emphasis>
</taxonomicName>
Knapp, 2013 (see
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.1.123" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="123 - 233" refId="B4" refString="Imamura, H, 1996. Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes). Species Diversity 1 (2): 123 - 233, DOI: https://doi.org/10.12782/specdiv.1.123" title="Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes)." url="https://doi.org/10.12782/specdiv.1.123" volume="1" year="1996">Imamura 1996</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/BF02675436" author="Imamura, H" journalOrPublisher="Ichthyological Research" pageId="0" pageNumber="267" pagination="179 - 183" refId="B6" refString="Imamura, H, Knapp, LW, 1999. Thysanophrys papillaris, a new species of flathead from the Andaman Sea and northern Australia (Scorpaeniformes: Platycephalidae). Ichthyological Research 46 (2): 179 - 183, DOI: https://doi.org/10.1007/BF02675436" title="Thysanophrys papillaris, a new species of flathead from the Andaman Sea and northern Australia (Scorpaeniformes: Platycephalidae)." url="https://doi.org/10.1007/BF02675436" volume="46" year="1999">Imamura and Knapp 1999</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3608.2.3" author="Knapp, LW" journalOrPublisher="Zootaxa" pageId="0" pageNumber="267" pagination="127 - 136" refId="B8" refString="Knapp, LW, 2013. Descriptions of four new species of Thysanophrys (Scorpaeniformes: Platycephalidae) from the western Indian Ocean. Zootaxa 3608 (2): 127 - 136, DOI: https://doi.org/10.11646/zootaxa.3608.2.3" title="Descriptions of four new species of Thysanophrys (Scorpaeniformes: Platycephalidae) from the western Indian Ocean." url="https://doi.org/10.11646/zootaxa.3608.2.3" volume="3608" year="2013">Knapp 2013</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.24.17" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="17 - 22" refId="B7" refString="Imamura, H, Kimura, K, Quan, NV, 2019. First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific. Species Diversity 24 (1): 17 - 22, DOI: https://doi.org/10.12782/specdiv.24.17" title="First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific." url="https://doi.org/10.12782/specdiv.24.17" volume="24" year="2019">Imamura et al. 2019</bibRefCitation>
).
<taxonomicName authorityName="Prokofiev" authorityYear="2017" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys haploblepharis" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="haploblepharis">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys haploblepharis</emphasis>
</taxonomicName>
Prokofiev, 2017 was regarded as a junior synonym of
<taxonomicName baseAuthorityName="Regan" baseAuthorityYear="1908" class="Actinopterygii" family="Platycephalidae" genus="Insidiator" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Insidiator cooperi" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="cooperi">
<emphasis italics="true" pageId="0" pageNumber="267">Insidiator cooperi</emphasis>
</taxonomicName>
(Regan, 1908) by
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.24.17" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="17 - 22" refId="B7" refString="Imamura, H, Kimura, K, Quan, NV, 2019. First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific. Species Diversity 24 (1): 17 - 22, DOI: https://doi.org/10.12782/specdiv.24.17" title="First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific." url="https://doi.org/10.12782/specdiv.24.17" volume="24" year="2019">Imamura et al. (2019)</bibRefCitation>
. The presently reported specimens from southern Taiwan conformed to
<taxonomicName class="Pisces" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">T. papillaris</emphasis>
</taxonomicName>
, differing from all congeners in having the following combination of characters: 11 second dorsal-fin rays, 12 anal-fin rays, 59-74 scale rows below the lateral line slanting downward and forward, a longer snout (snout length / orbit diameter ratio 1.1-1.2), 1 or 2 small papillae on the eye, the upper iris lappet with short branches, a single preorbital spine and 3-5 suborbital spines (see
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.24.17" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="17 - 22" refId="B7" refString="Imamura, H, Kimura, K, Quan, NV, 2019. First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific. Species Diversity 24 (1): 17 - 22, DOI: https://doi.org/10.12782/specdiv.24.17" title="First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific." url="https://doi.org/10.12782/specdiv.24.17" volume="24" year="2019">Imamura et al. 2019</bibRefCitation>
for a detailed comparison of all 10 species of
<taxonomicName authorityName="J.D.Ogilby" authorityYear="1898" class="Actinopterygii" family="Platycephalidae" genus="Thysanophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thysanophrys" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="267">Thysanophrys</emphasis>
</taxonomicName>
, and materials of six of the nine species deposited in the collections of AMS, BMNH, CSIRO, NMV, NSMT, NTM, QM, USNM, and WAM). Accordingly, this study identified the six Taiwanese specimens as
<taxonomicName class="Pisces" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">T. papillaris</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.1007/BF02675436" author="Imamura, H" journalOrPublisher="Ichthyological Research" pageId="0" pageNumber="267" pagination="179 - 183" refId="B6" refString="Imamura, H, Knapp, LW, 1999. Thysanophrys papillaris, a new species of flathead from the Andaman Sea and northern Australia (Scorpaeniformes: Platycephalidae). Ichthyological Research 46 (2): 179 - 183, DOI: https://doi.org/10.1007/BF02675436" title="Thysanophrys papillaris, a new species of flathead from the Andaman Sea and northern Australia (Scorpaeniformes: Platycephalidae)." url="https://doi.org/10.1007/BF02675436" volume="46" year="1999">Imamura and Knapp (1999)</bibRefCitation>
originally described
<taxonomicName class="Pisces" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">T. papillaris</emphasis>
</taxonomicName>
as having a single small papilla on the eye (based on six specimens). Recently, however,
<bibRefCitation DOI="https://doi.org/10.12782/specdiv.24.17" author="Imamura, H" journalOrPublisher="Species Diversity" pageId="0" pageNumber="267" pagination="17 - 22" refId="B7" refString="Imamura, H, Kimura, K, Quan, NV, 2019. First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific. Species Diversity 24 (1): 17 - 22, DOI: https://doi.org/10.12782/specdiv.24.17" title="First record of Thysanophrys papillaris (Actinopterygii: Scorpaeniformes: Platycephalidae) from the western Pacific." url="https://doi.org/10.12782/specdiv.24.17" volume="24" year="2019">Imamura et al. (2019)</bibRefCitation>
recorded a single specimen with an additional papilla on the right eye from Nha Trang, Vietnam, and noted the potential for intraspecific variation in the number of eye papillae. Recognition herein of two specimens of
<taxonomicName class="Pisces" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">T. papillaris</emphasis>
</taxonomicName>
with an additional papilla on the eye confirms such intraspecific variation. The presently reported specimens represent the first record of
<taxonomicName class="Pisces" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. papillaris" order="Scorpaeniformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="papillaris">
<emphasis italics="true" pageId="0" pageNumber="267">T. papillaris</emphasis>
</taxonomicName>
from Taiwan and the northernmost distributional record of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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