584 lines
55 KiB
XML
584 lines
55 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.97.57202" ID-GBIF-Dataset="88a8354f-7333-461b-ab84-6d742b9a7500" ID-GBIF-Taxon="176584340" ID-Pensoft-Pub="1860-0743-1-55" ID-Pensoft-UUID="8E2883E1FFEE5B6884EADB662AB420A1" ID-ZooBank="A125EC81050E449A890427674F5265EF" LSID="urn:lsid:zoobank.org:act:B2939E10-5F5D-45D0-9CBF-352D1D8A6B9A" ModsDocID="1860-0743-1-55" checkinTime="1610662682717" checkinUser="pensoft" docAuthor="Branch, William R., Schmitz, Andreas, Lobon-Rovira, Javier, Baptista, Ninda L., Antonio, Telmo & Conradie, Werner" docDate="2021" docId="A5ABF1097A475B11905E00252EDBB003" docLanguage="en" docName="ZoosystEvol 97(1): 55-82" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-01-12" docSource="http://dx.doi.org/10.3897/zse.97.57202" docTitle="Afroedura donveae Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie 2021, sp. nov." docType="treatment" docUuid="B2939E10-5F5D-45D0-9CBF-352D1D8A6B9A" docUuidSource="ZooBank" docVersion="4" id="8E2883E1FFEE5B6884EADB662AB420A1" lastPageNumber="55" masterDocId="8E2883E1FFEE5B6884EADB662AB420A1" masterDocTitle="Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola" masterLastPageNumber="82" masterPageNumber="55" pageId="0" pageNumber="55" updateTime="1635948359121" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Branch, William R.</mods:namePart>
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<mods:affiliation>Port Elizabeth Museum, P. O. Box 13147, Humewood, 6013, Port Elizabeth, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela University, Port Elizabeth 6031, South Africa</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schmitz, Andreas</mods:namePart>
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<mods:affiliation>Natural History Museum of Geneva, UREC - Herpetology and Ichthyology, C. P. 6434, 1211 Geneva 6, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lobon-Rovira, Javier</mods:namePart>
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<mods:affiliation>CIBIO Centro de Investigacao em Biodiversidade e Recursos Geneticos, Universidade do Porto, Rua Padre Armando Quintas, Campus de Vairao, 4485 - 661 Vairao, Portugal & Faculdade de Ciencias, Universidade do Porto, 4169 - 007 Porto, Portugal</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baptista, Ninda L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2859-6606</mods:nameIdentifier>
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<mods:affiliation>CIBIO Centro de Investigacao em Biodiversidade e Recursos Geneticos, Universidade do Porto, Rua Padre Armando Quintas, Campus de Vairao, 4485 - 661 Vairao, Portugal & Faculdade de Ciencias, Universidade do Porto, 4169 - 007 Porto, Portugal & Instituto Superior de Ciencias da Educacao da Huila (ISCED), Rua Sarmento Rodrigues s / n, Lubango, Angola</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Antonio, Telmo</mods:namePart>
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<mods:affiliation>Instituto Superior de Ciencias da Educacao da Huila (ISCED), Rua Sarmento Rodrigues s / n, Lubango, Angola & Faculty of Natural Resources and Spatial Sciences, Namibia University of Science and Technology, Private Bag 13388, Windhoek, Namibia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Conradie, Werner</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0805-9683</mods:nameIdentifier>
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<mods:affiliation>Port Elizabeth Museum, P. O. Box 13147, Humewood, 6013, Port Elizabeth, South Africa & School of Natural Resource Management, George Campus, Nelson Mandela University, George 6530, South Africa</mods:affiliation>
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<mods:nameIdentifier type="email">werner@bayworld.co.za</mods:nameIdentifier>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-01-12</mods:number>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:start>55</mods:start>
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<mods:end>82</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.97.57202</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.57202</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-55</mods:identifier>
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<mods:identifier type="Pensoft-UUID">8E2883E1FFEE5B6884EADB662AB420A1</mods:identifier>
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<treatment ID-GBIF-Taxon="176584340" LSID="urn:lsid:zoobank.org:act:B2939E10-5F5D-45D0-9CBF-352D1D8A6B9A" httpUri="http://treatment.plazi.org/id/A5ABF1097A475B11905E00252EDBB003" lastPageNumber="55" pageId="0" pageNumber="55">
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<subSubSection pageId="0" pageNumber="55" type="nomenclature">
|
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<paragraph pageId="0" pageNumber="55">
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<taxonomicName LSID="http://zoobank.org/B2939E10-5F5D-45D0-9CBF-352D1D8A6B9A" authority="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie, 2021" authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura donveae" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="donveae" status="sp. nov.">Afroedura donveae</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="55">sp. nov.</taxonomicNameLabel>
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Live photos of Angolan flat geckos: A. Afroedura bogerti (PEM R 24187); B. Afroedura wulfhaackei sp. nov. (paratype PEM R 24232); C. Afroedura praedicta sp. nov. (holotype NB 854); D. Afroedura donveae sp. nov. (holotype PEM R 17937); E. Afroedura vazpintorum sp. nov. (holotype PEM R 24118); F. Afroedura vazpintorum sp. nov. (NB 0745); Photos: A, B, D, E. William R. Branch; C, F. Pedro Vaz Pinto." figureDoi="10.3897/zse.97.57202.figure5" httpUri="https://binary.pensoft.net/fig/496674" pageId="0" pageNumber="55">Iona Flat Gecko Osga-achatada do Iona Figures 5D</figureCitation>
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Holotype of Afroedura donveae sp. nov. (PEM R 17937) from Omauha Lodge, 15 km south of Tambor, Namibe Province, Angola. Photos: Werner Conradie." figureDoi="10.3897/zse.97.57202.figure9" httpUri="https://binary.pensoft.net/fig/496678" pageId="0" pageNumber="55">, 9</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Summary of morphological data for Afroedura bogerti, A. wulfhaackei sp. nov. (including the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades), A. donveae sp. nov., A. vazpintorum sp. nov. and A. praedicta sp. nov. Values are given as a range with mean values in parenthesis. M = male, F = female, n = sample size." pageId="0" pageNumber="55">Tables 3</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm) and scale counts for the type series of Afroedura donveae sp. nov." pageId="0" pageNumber="55">7</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="synonym">
|
||
<paragraph pageId="0" pageNumber="55">Synonym.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura bogerti</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<bibRefCitation DOI="https://doi.org/10.1080/21564574.2008.9635571" author="Haacke, WD" journalOrPublisher="African Journal Herpetology" pageId="0" pageNumber="55" pagination="85 - 92" refId="B33" refString="Haacke, WD, 2008. A new leaf-toed gecko (Reptilia: Gekkonidae) from south-western Angola. African Journal Herpetology 57 (2): 85 - 92, DOI: https://doi.org/10.1080/21564574.2008.9635571" title="A new leaf-toed gecko (Reptilia: Gekkonidae) from south-western Angola." url="https://doi.org/10.1080/21564574.2008.9635571" volume="57" year="2008">Haacke 2008</bibRefCitation>
|
||
:6,
|
||
<bibRefCitation author="Huntley, BJ" journalOrPublisher="Science and Conservation: A Modern Synthesis. Editors, Springer Open, Cham" pageId="0" pageNumber="55" refId="B38" refString="Huntley, BJ, 2009. SANBI/ISCED/UAN Angolan biodiversity assessment capacity building project. Report on pilot project. Unpublished Report to Ministry of Environment, Luanda 97 pp." title="SANBI / ISCED / UAN Angolan biodiversity assessment capacity building project. Report on pilot project. Unpublished Report to Ministry of Environment, Luanda 97 pp." year="2009">Huntley 2009</bibRefCitation>
|
||
:84,
|
||
<bibRefCitation author="Roesler, H" journalOrPublisher="Gekkota" pageId="0" pageNumber="55" pagination="28 - 153" refId="B59" refString="Roesler, H, 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28 - 153" title="Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha)." volume="2" year="2000">
|
||
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
||
2000
|
||
</bibRefCitation>
|
||
:57, Barts and Haacke 2010:39,
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3846.4.1" author="Jacobsen, NHG" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="451 - 501" refId="B41" refString="Jacobsen, NHG, Kuhn, AL, Jackman, TR, Bauer, AM, 2014. A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa. Zootaxa 3846 (4): 451 - 501, DOI: https://doi.org/10.11646/zootaxa.3846.4.1" title="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa." url="https://doi.org/10.11646/zootaxa.3846.4.1" volume="3846" year="2014">Jacobsen et al. 2014</bibRefCitation>
|
||
:456 & 468 (part),
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zse.93.10915" author="Branch, WR" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="55" pagination="157 - 166" refId="B13" refString="Branch, WR, Haacke, W, Vaz Pinto, P, Conradie, W, Baptista, N, Verburgt, L, Verissimo, L, 2017b. Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status. Zoosystematics and Evolution 93 (1): 157 - 166, DOI: https://doi.org/10.3897/zse.93.10915" title="Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status." url="https://doi.org/10.3897/zse.93.10915" volume="93" year="2017 b">Branch et al. 2017b</bibRefCitation>
|
||
:157 (part),
|
||
<bibRefCitation author="Marques, MP" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" refId="B49" refString="Marques, MP, Ceriaco, LMP, Blackburn, DC, Bauer, AM, 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola. Atlas of Historical and Bibliographic Records (1840-2017). Proceedings of the California Academy of Science, Series 4 65(Supplement II): 1-501." title="Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola. Atlas of Historical and Bibliographic Records (1840 - 2017). Proceedings of the California Academy of Science, Series 4 65 (Supplement II): 1 - 501." year="2018">Marques et al. 2018</bibRefCitation>
|
||
: 177 (part),
|
||
<bibRefCitation DOI="https://doi.org/10.1007/978-3-030-03083-4_13" author="Branch, WR" editor="Huntley, BJ" journalOrPublisher="Science and Conservation: A Modern Synthesis. Springer Open, Cham" pageId="0" pageNumber="55" pagination="283 - 326" refId="B9" refString="Branch, WR, Baptista, N, Vaz Pinto, P, Conradie, W, 2019a. The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research. In: Huntley, BJ, Ferrand, N, Eds., Biodiversity of Angola. Science and Conservation: A Modern Synthesis. Springer Open, Cham: 283 - 326, DOI: https://doi.org/10.1007/978-3-030-03083-4_13" title="The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research." url="https://doi.org/10.1007/978-3-030-03083-4_13" volumeTitle="Biodiversity of Angola." year="2019 a">Branch et al. 2019a</bibRefCitation>
|
||
: 287 (part);
|
||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="bogerti">Afroedura cf. bogerti</taxonomicName>
|
||
-
|
||
<bibRefCitation author="Agarwal, I" journalOrPublisher="Herpetological Review" pageId="0" pageNumber="55" pagination="649 - 650" refId="B1" refString="Agarwal, I, Ceriaco, LMP, Bauer, AM, Bandeira, SA, 2017. Kolekanos plumicaudus (Slender Feather-tailed Gecko) Habitat use. Herpetological Review 48: 649 - 650" title="Kolekanos plumicaudus (Slender Feather-tailed Gecko) Habitat use." volume="48" year="2017">Agarwal et al. 2017</bibRefCitation>
|
||
:649;
|
||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura bogerti</emphasis>
|
||
</taxonomicName>
|
||
(clade 1) -
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4324.1.8" author="Branch, WR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="142 - 160" refId="B12" refString="Branch, WR, Guyton, JA, Schmitz, A, Barej, MF, Naskrecki, P, Farooq, H, Verburgt, L, Roedel, M-O, 2017a. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142 - 160, DOI: https://doi.org/10.11646/zootaxa.4324.1.8" title="Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique." url="https://doi.org/10.11646/zootaxa.4324.1.8" volume="4324" year="2017 a">Branch et al. 2017a</bibRefCitation>
|
||
:146.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="holotype">
|
||
<paragraph pageId="0" pageNumber="55">Holotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
PEM R17937, adult female, collected from Omauha Lodge, 15 km south of Tambor (
|
||
<geoCoordinate degrees="16.20061" direction="south" orientation="latitude" precision="1" value="-16.20061">-16.20061</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.40183" direction="east" orientation="longitude" precision="1" value="12.40183">12.40183</geoCoordinate>
|
||
, 341 m a.s.l.), Namibe Province, Angola, by William R. Branch, Werner Conradie, Krystal Tolley and John Measey on 18 January 2009.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="paratype">
|
||
<paragraph pageId="0" pageNumber="55">Paratype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
PEM R17936, collected from Omauha Lodge, 15 km South Tambor (
|
||
<geoCoordinate degrees="16.20061" direction="south" orientation="latitude" precision="1" value="-16.20061">-16.20061</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.40183" direction="east" orientation="longitude" precision="1" value="12.40183">12.40183</geoCoordinate>
|
||
, 341 m a.s.l), Namibe Province, Angola, by William R. Branch, Werner Conradie, Krystal Tolley and John Measey on 18 January 2009.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="55">Additional material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Males</emphasis>
|
||
: TM 40508, TM 40512, TM 40516, TM 40518, collected from Tambor (
|
||
<geoCoordinate degrees="16.06667" direction="south" orientation="latitude" precision="1" value="-16.06667">-16.06667</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.43333" direction="east" orientation="longitude" precision="1" value="12.43333">12.43333</geoCoordinate>
|
||
, 355 m a.s.l.), Namibe Province, Angola, by Wulf Haacke on 1 April 1971.
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Females</emphasis>
|
||
: PEM R18041-2, collected 0.5 km south of Tambor (
|
||
<geoCoordinate degrees="16.07414" direction="south" orientation="latitude" precision="1" value="-16.07414">-16.07414</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.43328" direction="east" orientation="longitude" precision="1" value="12.43328">12.43328</geoCoordinate>
|
||
, 352 m a.s.l.), Namibe Province, Angola, by William R. Branch, Krystal Tolley and John Measey on 23 January 2009; TM 40509-11, TM 40513-5, TM 40517, TM 40536-8, collected from Tambor (
|
||
<geoCoordinate degrees="16.06667" direction="south" orientation="latitude" precision="1" value="-16.06667">-16.06667</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.43333" direction="east" orientation="longitude" precision="1" value="12.43333">12.43333</geoCoordinate>
|
||
, 355 m a.s.l.), Namibe Province, Angola, by Wulf Haacke on 1 April 1971.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="55">Additional referred material</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">(not examined).</emphasis>
|
||
TM 40519-20, collected from Tambor (
|
||
<geoCoordinate degrees="16.06667" direction="south" orientation="latitude" precision="1" value="-16.06667">-16.06667</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.43333" direction="east" orientation="longitude" precision="1" value="12.43333">12.43333</geoCoordinate>
|
||
, 355 m a.s.l.), Namibe Province, Angola, by Wulf Haacke on 1 April 1971; FKH 0341-2, collected from Omauha Lodge (
|
||
<geoCoordinate degrees="16.20061" direction="south" orientation="latitude" precision="1" value="-16.20061">-16.20061</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.40183" direction="east" orientation="longitude" precision="1" value="12.40183">12.40183</geoCoordinate>
|
||
, 338 m a.s.l.), Namibe Province, Angola, by Pedro Vaz Pinto on 3 October 2019; CAS:HERP 263012-3, collected from Omauha (
|
||
<geoCoordinate degrees="16.19858" direction="south" orientation="latitude" precision="1" value="-16.19858">-16.19858</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.40073" direction="east" orientation="longitude" precision="1" value="12.40073">12.40073</geoCoordinate>
|
||
, 338 m a.s.l.), Namibe Province, Angola, by Luis M.P. Ceriaco, Suzana Bandeira and Isham Agarwal on 25 and 27 November 2017; CAS:HERP 248780-1, collected 0.5 km south of Tambor (
|
||
<geoCoordinate degrees="16.07414" direction="south" orientation="latitude" precision="1" value="-16.07414">-16.07414</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12.43328" direction="east" orientation="longitude" precision="1" value="12.43328">12.43328</geoCoordinate>
|
||
, 352 m a.s.l.), Namibe Province, Angola, by William R. Branch, Krystal Tolley and John Measey on 23 January 2009.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="etymology">
|
||
<paragraph pageId="0" pageNumber="55">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
This gecko is named after
|
||
<normalizedToken originalValue="Donvé">Donve</normalizedToken>
|
||
Branch,
|
||
<normalizedToken originalValue="WRB’s">WRB's</normalizedToken>
|
||
wife, with the following personal quote: "This, the most beautiful of all the Angolan flat geckos, is named for my wife,
|
||
<normalizedToken originalValue="Donvé">Donve</normalizedToken>
|
||
Branch (
|
||
<normalizedToken originalValue="‘Dove’">'Dove'</normalizedToken>
|
||
) who bore the long periods I was away on fieldwork, and to whose nest I returned, and surrounded me with love until the end". The name is constructed in the feminine singular genitive.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="55">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
A member of the greater '
|
||
<taxonomicName baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="transvaalica">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura transvaalica</emphasis>
|
||
</taxonomicName>
|
||
' group as it possesses two pairs of enlarged scansors per digit and a strongly verticillate and flattened tail (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3846.4.1" author="Jacobsen, NHG" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="451 - 501" refId="B41" refString="Jacobsen, NHG, Kuhn, AL, Jackman, TR, Bauer, AM, 2014. A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa. Zootaxa 3846 (4): 451 - 501, DOI: https://doi.org/10.11646/zootaxa.3846.4.1" title="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa." url="https://doi.org/10.11646/zootaxa.3846.4.1" volume="3846" year="2014">Jacobsen et al. 2014</bibRefCitation>
|
||
). Part of the
|
||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
-group which differs from other members of the '
|
||
<taxonomicName baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="transvaalica">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura transvaalica</emphasis>
|
||
</taxonomicName>
|
||
' group by having less than 78 mid-body scale rows (vs. 97-102 in
|
||
<taxonomicName genus="A." lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
|
||
</taxonomicName>
|
||
, 113-120 in
|
||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||
</taxonomicName>
|
||
, 102-119 in
|
||
<taxonomicName genus="A." lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
|
||
</taxonomicName>
|
||
); by the rostral bordering the nostril (nostril excluded from rostral in
|
||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||
</taxonomicName>
|
||
); by the anterior nasals always in contact (separated by 1-3 granules in
|
||
<taxonomicName genus="A." lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
|
||
</taxonomicName>
|
||
; always in broad contact in
|
||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||
</taxonomicName>
|
||
; usually in broad contact in
|
||
<taxonomicName genus="A." lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
|
||
</taxonomicName>
|
||
~ 3-18%); and in having 11-14 scales between the anterior borders of the eyes (19-22 in
|
||
<taxonomicName genus="A." lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
|
||
</taxonomicName>
|
||
; 15-19 in
|
||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||
</taxonomicName>
|
||
; 15-20 in
|
||
<taxonomicName genus="A." lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
|
||
</taxonomicName>
|
||
) (comparative data
|
||
<emphasis italics="true" pageId="0" pageNumber="55">fide</emphasis>
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4324.1.8" author="Branch, WR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="142 - 160" refId="B12" refString="Branch, WR, Guyton, JA, Schmitz, A, Barej, MF, Naskrecki, P, Farooq, H, Verburgt, L, Roedel, M-O, 2017a. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142 - 160, DOI: https://doi.org/10.11646/zootaxa.4324.1.8" title="Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique." url="https://doi.org/10.11646/zootaxa.4324.1.8" volume="4324" year="2017 a">Branch et al. 2017a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura donveae" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura donveae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from other members of the
|
||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
-group by a combination of the following characters (see Tables
|
||
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Summary of morphological data for Afroedura bogerti, A. wulfhaackei sp. nov. (including the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades), A. donveae sp. nov., A. vazpintorum sp. nov. and A. praedicta sp. nov. Values are given as a range with mean values in parenthesis. M = male, F = female, n = sample size." httpUri="http://table.plazi.org/id/742172291B68E9AFACEE2244550D3FEF" pageId="0" pageNumber="55" tableUuid="742172291B68E9AFACEE2244550D3FEF">3</tableCitation>
|
||
,
|
||
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Tukey multiple comparisons of means of the four species combinations of Angolan Afroedura (values for A. wulfhaackei sp. nov. include the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades). Bold values indicate significant pairwise comparisons. SVL - snout-vent length; HL - head length; HW - head width; ED - eye-diameter; EE - ear-eye length; IN - internostril distance; MSR - mid-body scale rows." httpUri="http://table.plazi.org/id/C30887A74A39988C01C45C883AEB5E57" pageId="0" pageNumber="55" tableUuid="C30887A74A39988C01C45C883AEB5E57">4</tableCitation>
|
||
): 64-78 (mean 72.8) mid-body scale rows (69-77 [mean 73.5] in
|
||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
, 76-88 [mean 79.3] in
|
||
<taxonomicName genus="A." lsidName="A. wulfhaackei" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., 73-86 [mean 80.3] in
|
||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., 73-78 [mean 74.8] in
|
||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.); by the anterior nasals always in contact (similar to
|
||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; in contact in ~ 33% of
|
||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
; in contact in ~ 68% of
|
||
<taxonomicName genus="A." lsidName="A. wulfhaackei" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.); in each verticil having 5-6 (mean 5.5) ventral and 6-7 (mean 6.6) dorsal rows of scales (5-6 [mean 5.0] and 6-7 [mean 6.1] to
|
||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; 4 and 5 in
|
||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., 4-5 [mean 4.0] and 5-6 [mean 5.1] in
|
||
<taxonomicName genus="A." lsidName="A. wulfhaackei" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.); ventral surfaces immaculate (similar to
|
||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; greyish with black spots in
|
||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="A." lsidName="A. wulfhaackei" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.); larger average adult size 57.6 mm SVL (versus 50.0 mm in
|
||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||
</taxonomicName>
|
||
, 51.7 mm in
|
||
<taxonomicName genus="A." lsidName="A. wulfhaackei" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. wulfhaackei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., 51.3 mm in
|
||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; 49.9 mm
|
||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.).
|
||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura donveae" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura donveae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs more specifically from its sister lowland species
|
||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in being larger (57.6 mm versus 51.3 mm average SVL) and having lower mid-body scale counts (64-78 [mean 72.8] versus 73-86 [mean 80.3]), higher numbers of precloacal pores (11-12 [mean 11.5] versus 9-11 [mean 10.2]), bolder colouration and distinct tail banding (versus duller colouration and less distinct tail banding).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="description">
|
||
<paragraph pageId="0" pageNumber="55">Holotype description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
Adult female: SVL 61.0 mm; tail 59.0 mm (full original tail), with a small mid-ventral incision for the removal of liver sample. Measurements and meristic characters of holotype presented in Table
|
||
<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm) and scale counts for the type series of Afroedura donveae sp. nov." httpUri="http://table.plazi.org/id/EAE8B9D45A4B8F42893D9A2DD5FC3759" pageId="0" pageNumber="55" tableUuid="EAE8B9D45A4B8F42893D9A2DD5FC3759">7</tableCitation>
|
||
. Head and body dorsoventrally compressed; HL 13.6 mm, HW 11.4 mm, broadest at posterior level of eye and 1.19 times longer than wide. Eyes large (3.4 mm wide), pupil vertical with indented margins; circumorbital scales small and smooth, elongated at upper anterior margin, upper three posterior scales with small upward pointing spines. Snout rounded, 5.6 mm long, longer than distance between eye and ear openings (4.5 mm). Scales on top of snout smooth, rounded, equal in size, with no intervening minute granules. Scales on snout slightly larger than those on back of head or nape. Scales on eyelids larger than those on the crown, 5 scales deep from circumorbital scale to crown. Nostril pierced between rostral, three nasal scales; 1st supralabial narrowly excluded from nostril; the supranasals are much larger than the subequal smaller postnasals, in broad contact with each other. Nostrils slightly elevated. Rostral roughly rectangular, but with its upper edges elongated due to extensions to the nostril. Nine supralabials on each side, the labial margin flexing upwards at the rictus (approx. mid-orbital position), with 1-2 minute scales proximal to the flexure. Ten infralabials on either side, with a small scale proximal to the flexure. At the lip, mental slightly narrower than adjacent infralabial, 63% of rostral and in contact with three distinctly elongated postmental scales. Scales on throat much smaller than those on belly, scales touching infralabials larger. Thirteen scales across the crown at level of front of eyes; 11 scales between nostril and front of eye; 13 scales from ear to eye; 74 scales around mid-body. Ear opening deep, oblique and roughly oval, only half as high as wide (0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
0.9 mm). Scales on dorsum smooth, non-overlapping, largest at mid-body, smaller on nape and tail base. Scales on ventrum flattened, not overlapping, more-or-less ovate at mid-ventrum, twice the size of lateral granules and 1.5 times the size of scales along the dorsal mid-line. Original tail slightly dorsoventrally flattened and distinctly verticillate (16 whorls in total), with obvious lateral constrictions; each verticil comprising 7 rows of imbricate scales dorsally and 6 rows of imbricate scales ventrally, with ventral scales approximately twice the size of those on the dorsal surface. Limbs well-developed, hindlimbs slightly longer than forelimbs, both without obvious mite pockets at posterior margin of limb insertions; mite pockets present on anterior margin of hindlimbs. All digits with a large pair of distal scansors, separated by a large, curved claw and followed after a large gap (twice length of terminal scansor) by a smaller pair of scansors; infero-median row of digital scales enlarged transversely, particularly towards the scansors, where the terminal scale adjoining the first pair of scansors may be medially constricted, swollen and scansor-like; 8 enlarged scale rows under 4th toe.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="EAE8B9D45A4B8F42893D9A2DD5FC3759" httpUri="http://table.plazi.org/id/EAE8B9D45A4B8F42893D9A2DD5FC3759" pageId="0" pageNumber="55" start="Table 7" startId="T7">
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">Table 7.</emphasis>
|
||
Measurements (in mm) and scale counts for the type series of
|
||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura donveae" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura donveae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<table pageId="0" pageNumber="55">
|
||
<tr pageId="0" pageNumber="55">
|
||
<th colspan="1" pageId="0" pageNumber="55" rowspan="1">Catalogue number</th>
|
||
<th colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R17937</th>
|
||
<th colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R17936</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Type Status</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Holotype</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Paratype</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Sex</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Female</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Female</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Snout vent length</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">61.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">64.0</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Tail length</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">59.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">38.0</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Tail condition</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Original</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Regenerated</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Head length</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">16.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">13.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Head width</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">11.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">11.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Snout length</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">5.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">5.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Eye distance</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">3.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">3.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Eye-Ear distance</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Dorsal rows per tail verticil</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">6</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Ventral rows per tail verticil</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales below 4th toe</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Mid-body scale rows</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">74</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">68</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales between eyes</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">12</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales: nostril to eye</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">11</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">10</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales: ear to eye</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">14</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Anterior nasals in contact</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Yes</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Yes</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Supralabials</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="55">
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Infralabials</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">10</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="type material">
|
||
<paragraph pageId="0" pageNumber="55">Paratypes and additional examined material variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
SVL varied from 49.1-55.7 mm; original tail length 26.5-59.0 mm, 0.78 times SVL; head length 1.13-1.46 times head width; snout 1.99 times diameter of eye (see Table
|
||
<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm) and scale counts for the type series of Afroedura donveae sp. nov." httpUri="http://table.plazi.org/id/EAE8B9D45A4B8F42893D9A2DD5FC3759" pageId="0" pageNumber="55" tableUuid="EAE8B9D45A4B8F42893D9A2DD5FC3759">7</tableCitation>
|
||
for more measurements and scale counts of type series). Supranasals always in contact; the first upper labial and rostral always enters the nostril and the width of the rostral at the lip margin is always wider than that of the mental; 2-3 postmental scales (except PEM R18401 with only one); supralabials 8-10, infralabials 8-11; scales between anterior edges of eyes 10-13; scales between nostril and anterior edge of orbit 8-12; scales from ear to eye 11-14; scales around mid-body 64-78; subdigital lamellae on 4th toe 6-8; dorsal scales per tail verticil 6-7 (mean 6.6); ventral scales per tail verticil 5-6 (mean 5.5); precloacal pores 11-12.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="colouration">
|
||
<paragraph pageId="0" pageNumber="55">Colouration.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">In life</emphasis>
|
||
(holotype PEM R17937, Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Live photos of Angolan flat geckos: A. Afroedura bogerti (PEM R 24187); B. Afroedura wulfhaackei sp. nov. (paratype PEM R 24232); C. Afroedura praedicta sp. nov. (holotype NB 854); D. Afroedura donveae sp. nov. (holotype PEM R 17937); E. Afroedura vazpintorum sp. nov. (holotype PEM R 24118); F. Afroedura vazpintorum sp. nov. (NB 0745); Photos: A, B, D, E. William R. Branch; C, F. Pedro Vaz Pinto." figureDoi="10.3897/zse.97.57202.figure5" httpUri="https://binary.pensoft.net/fig/496674" pageId="0" pageNumber="55">5D</figureCitation>
|
||
): Yellowish above, brighter anteriorly, with nine irregularly-shaped darker brown to black crossbars from the occiput to the sacrum, each separated by thinner (3-4 scales deep), light yellow crossbars; head with dark brown-black blotches on the crown with intervening pale yellow colouration; dark brown bar from nostril across the upper margins of the ear opening, connecting with dark brown lateral bar on the neck; a thin pale yellow canthal stripe extends on both sides from the nasal region to anterior margins of eye, continuing posteriorly from the eye on to the nape; upper and lower labials light grey with diffuse brown edges; lateral sides of the body with a mix of dark grey and yellow blotches; limbs yellowish above with scattered darker grey markings; tail (original) with irregular dark brown bars, separated by yellow bars anteriorly, posteriorly with black and white bars; iris dark black with irregular golden spots, a black, narrow, elliptical, crenulate-edged pupil with black reticulation; ventrum uniform beige with scattered brown specks on lateral edges only; ventrally, limbs with scattered brown spots.
|
||
<emphasis italics="true" pageId="0" pageNumber="55">In preservative</emphasis>
|
||
(holotype PEM R17937, Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Holotype of Afroedura donveae sp. nov. (PEM R 17937) from Omauha Lodge, 15 km south of Tambor, Namibe Province, Angola. Photos: Werner Conradie." figureDoi="10.3897/zse.97.57202.figure9" httpUri="https://binary.pensoft.net/fig/496678" pageId="0" pageNumber="55">9</figureCitation>
|
||
): dorsum with nine irregularly spaced dark brown crossbars from the occiput to the sacrum with beige intervening blotches; dorsally, arms and legs darkly barred; tail with nine dark brown bars, bolder towards the tip; dorsally, head with mottled dark brown scales; a light beige canthal stripe from nostril to anterior corner of eyes, continuing from the posterior part of the eye to above the ear opening; dark brown bar running from the nasals above the ear opening to the neck; light beige stripe (one-scale-wide) from above the supralabials to the ear opening (two-scales-wide); supralabials ventrally dark brown-edged; infralabials with scattered dark brown markings dorsally; ventrum is mostly immaculate, with dark brown spots ventrally on the arms and legs; ventrally, tail with nine dark brown bars, bolder posteriorly.
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Variation</emphasis>
|
||
. Similar colouration and patterning as to the holotype. Dorsal crossbars are often fused to form 2-3 dark brown, X-shaped crossbars. Original tails with 6-7 broad, dark brown to black bars, separated by light beige to white bars; some specimens with fine one-scale-wide black bar separating the lighter bars. Regenerated tails with fine dark brown mottling. Juveniles with more sharply-defined patterns.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.57202.figure9" httpUri="https://binary.pensoft.net/fig/496678" pageId="0" pageNumber="55" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">Figure 9.</emphasis>
|
||
Holotype of
|
||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura donveae" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="donveae">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura donveae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (PEM R17937) from Omauha Lodge, 15 km south of Tambor, Namibe Province, Angola. Photos: Werner Conradie.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="natural history">
|
||
<paragraph pageId="0" pageNumber="55">Natural history and habitat</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
<emphasis bold="true" pageId="0" pageNumber="55">
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Typical habitat of Angolan Afroedura. A. Namba area (A. bogerti); B. Mt Sandula (Afroedura sp. 5); C. Serra da Neve (A. praedicta sp. nov.); D. Omauha Lodge (A. donveae sp. nov.); E. Farm Mucungo (A. vazpintorum sp. nov.); F. Bimbe (A. vazpintorum sp. nov.). Photos: A. Javier Lobon-Rovira; B, D, E. William R. Branch; C. Pedro Vaz Pinto; F. Ninda L. Baptista." figureDoi="10.3897/zse.97.57202.figure4" httpUri="https://binary.pensoft.net/fig/496673" pageId="0" pageNumber="55">4D</figureCitation>
|
||
).
|
||
</emphasis>
|
||
One or two elongated white eggs were laid on limestone rock, to which they adhere in captivity; eggs measure 12.9-14.7 mm (mean 13.8 mm)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
10.5-12.1 mm (mean 11.6 mm) (n = 5) and hatch after 88-92 days, with hatchlings measuring 30.2-30.9 mm SVL and 26.5-27.9 mm tail length (n = 5) (Barts and Haacke 2010). A rupicolous species living in crevices between rocks and under flakes of exfoliating rock amongst larger granite boulders, at elevations of 340-355 m a.s.l. in the arid Namib Desert. It appears to be associated with large granite outcrops in semi-desert shrublands, along the drier fringes of the Pro-Namib. Vegetation includes
|
||
<taxonomicName authorityName="Rafinesque" authorityYear="1838" class="Equisetopsida" family="Fabaceae" genus="Senegalia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Senegalia" order="Fabales" pageId="0" pageNumber="55" phylum="Magnoliophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Senegalia</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName class="Dicotyledoneae" family="Fabaceae" genus="Acacia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acacia" order="Rosales" pageId="0" pageNumber="55" phylum="Angiospermae" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Acacia</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<emphasis italics="true" pageId="0" pageNumber="55">mellifera</emphasis>
|
||
,
|
||
<taxonomicName authorityName="Rafinesque" authorityYear="1838" class="Equisetopsida" family="Fabaceae" genus="Senegalia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Senegalia" order="Fabales" pageId="0" pageNumber="55" phylum="Magnoliophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Senegalia</emphasis>
|
||
</taxonomicName>
|
||
spp.,
|
||
<taxonomicName authorityName="N.J.Jacquin" authorityYear="1797" class="Magnoliopsida" family="Burseraceae" genus="Commiphora" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Commiphora" order="Sapindales" pageId="0" pageNumber="55" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Commiphora</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName authorityName="Schinz" authorityYear="1888" class="Magnoliopsida" family="Capparaceae" genus="Boscia" higherTaxonomySource="GBIF,IPNI" kingdom="Plantae" lsidName="Boscia foetida" order="Brassicales" pageId="0" pageNumber="55" phylum="Tracheophyta" rank="species" species="foetida">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Boscia foetida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Reptilia" family="Colubridae" genus="Salvadora" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Salvadora persica" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="persica">
|
||
<emphasis italics="true" pageId="0" pageNumber="55">Salvadora persica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="55" type="distribution">
|
||
<paragraph pageId="0" pageNumber="55">Distribution and conservation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="55">
|
||
Currently known only from the south-western parts of Namibe Province in Angola, in granite formations around Tambor and on the right bank of the mid-Curoca River (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Digital elevation map showing the predicted distribution of the five newly-recognised species of Afroedura occurring in Angola (data for A. wulfhaackei sp. nov. also contain the morphologically-indistinguishable Afroedura sp. 5 - 7 clades). Confirmed locality records are indicated by solid circles, with type localities indicated by stars. Coloured polygons indicate potential distribution areas with more than 90 % of climatic habitat suitability." figureDoi="10.3897/zse.97.57202.figure1" httpUri="https://binary.pensoft.net/fig/496670" pageId="0" pageNumber="55">1</figureCitation>
|
||
). The species remains poorly known, but it is probably stable in numbers as the local habitat is currently not threatened. It likely occurs in Iona National Park.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |