300 lines
42 KiB
XML
300 lines
42 KiB
XML
<document id="7C772D65564E779F5B16BFDF9F79763B" ID-DOI="10.5281/zenodo.210542" ID-GBIF-Dataset="c9e5cc54-2a89-4b75-91aa-a4fce79f17fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="210542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460500319311" checkinUser="plazi" docAuthor="Ramírez-Ponce, Andrés & Morón, Miguel Angel" docDate="2012" docId="A567C22F5851FF8BFF64A648532B8143" docLanguage="en" docName="zt03394p024.pdf" docOrigin="Zootaxa 3394" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pachystethus Blanchard 1851" docType="treatment" docVersion="9" lastPageNumber="4" masterDocId="595EBA575850FF88FFF3A25B52478434" masterDocTitle="Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="2" updateTime="1698311598655" updateUser="plazi">
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<mods:title id="B4E95EB241447597080C8406F9620D6D">Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)</mods:title>
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<mods:namePart id="26A065DF3864533466A2A9FBD556BC8B">Ramírez-Ponce, Andrés</mods:namePart>
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<mods:namePart id="22D60B62A81DA348597D4BDA22D8CFD1">Morón, Miguel Angel</mods:namePart>
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<mods:typeOfResource id="138C51B860BB26F6015540320592CC15">text</mods:typeOfResource>
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<mods:title id="530EF0000CE69444B1D6E5F3363BAC40">Zootaxa</mods:title>
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<mods:date id="B9DB8D2F02AD12C1AA301AE2ECE8BBDA">2012</mods:date>
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<mods:identifier id="5E9C5036853D74796010CFB6DC4C1F17" type="DOI">10.5281/zenodo.210542</mods:identifier>
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<mods:identifier id="BD6EFC1E72EC070864D5470E5F36AF81" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A567C22F5851FF8BFF64A648532B8143" ID-DOI="http://doi.org/10.5281/zenodo.6179190" ID-GBIF-Taxon="119521049" ID-Zenodo-Dep="6179190" LSID="urn:lsid:plazi:treatment:A567C22F5851FF8BFF64A648532B8143" httpUri="http://treatment.plazi.org/id/A567C22F5851FF8BFF64A648532B8143" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="65D420B25851FF89FF64A6485358807A" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="2D7173395851FF89FF64A64850268019" blockId="1.[151,609,1043,1102]" box="[151,609,1043,1070]" pageId="1" pageNumber="2">
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<heading id="7639C4555851FF89FF64A64850268019" bold="true" box="[151,609,1043,1070]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="1FBAAF2B5851FF89FF64A64850268019" bold="true" box="[151,609,1043,1070]" pageId="1" pageNumber="2">
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Genus
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<taxonomicName id="EACE08BA5851FF89FF1EA64F50268019" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[237,609,1043,1070]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="1FBAAF2B5851FF89FF1EA64F53CA801A" bold="true" box="[237,397,1044,1070]" italics="true" pageId="1" pageNumber="2">Pachystethus</emphasis>
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<bibRefCitation id="495F0EC85851FF89FE66A64850268019" author="Blanchard" box="[405,609,1043,1069]" pageId="1" pageNumber="2" refString="Blanchard, C. E. (1851) Catalogue de la collection entomologique. Class des insects. Ordre des coleopteres. Volume 2. Museum d´Historie Naturelle de Paris, Paris" type="book" year="1851">Blanchard, 1851</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2D7173395851FF89FF64A66D5358807A" blockId="1.[151,609,1043,1102]" box="[151,287,1078,1102]" pageId="1" pageNumber="2">
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(
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<figureCitation id="B5F56FBC5851FF89FF6CA66D5350807A" box="[159,279,1078,1102]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 7" captionStart-2="FIGURES 8 – 13" captionStart-3="FIGURES 14 – 19" captionStart-4="FIGURES 20 – 25" captionStart-5="FIGURES 26 – 35" captionStart-6="FIGURES 36 – 41" captionStart-7="FIGURES 42 – 47" captionStart-8="FIGURES 48 – 53" captionStart-9="FIGURE 54" captionStartId-0="2.[151,250,1996,2019]" captionStartId-1="5.[151,264,1786,1809]" captionStartId-2="7.[151,264,1669,1692]" captionStartId-3="11.[151,264,1976,1999]" captionStartId-4="14.[151,264,1651,1674]" captionStartId-5="15.[151,264,1938,1961]" captionStartId-6="18.[151,264,1619,1642]" captionStartId-7="19.[151,264,1909,1932]" captionStartId-8="20.[151,264,1259,1282]" captionStartId-9="21.[151,250,1234,1257]" captionTargetBox-0="[183,1336,544,1995]" captionTargetBox-1="[183,1412,244,1709]" captionTargetBox-2="[185,1401,251,1668]" captionTargetBox-3="[167,1348,573,1945]" captionTargetBox-4="[205,1355,262,1613]" captionTargetBox-5="[238,1394,484,1899]" captionTargetBox-6="[189,1372,224,1565]" captionTargetBox-7="[214,1377,207,1832]" captionTargetBox-8="[200,1411,240,1184]" captionTargetBox-9="[157,1436,382,1193]" captionTargetId-0="figure@2.[182,1387,509,1998]" captionTargetId-1="figure@5.[151,1436,193,1765]" captionTargetId-2="figure@7.[151,1436,225,1677]" captionTargetId-3="figure@11.[151,1436,553,1965]" captionTargetId-4="figure@14.[151,1436,239,1645]" captionTargetId-5="figure@15.[151,1436,445,1918]" captionTargetId-6="figure@18.[151,1436,193,1598]" captionTargetId-7="figure@19.[198,1392,193,1847]" captionTargetId-8="figure@20.[151,1436,193,1238]" captionTargetId-9="figure@21.[157,1442,373,1193]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="11" captionTargetPageId-4="14" captionTargetPageId-5="15" captionTargetPageId-6="18" captionTargetPageId-7="19" captionTargetPageId-8="20" captionTargetPageId-9="21" captionText-0="FIGURE 1. Dorsal habitus of Pachystethus matzapatlecus new species, male." captionText-1="FIGURES 2 – 7. Diagnostic structures in P. viduus: 2) right proleg; 3) right antenna, 4) male genitalia in dorsal view, 5) male genitalia in lateral view, 6) spiculum gastrale, 7) female genital plates. Scale lines 1 mm." captionText-2="FIGURES 8 – 13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm." captionText-3="FIGURES 14 – 19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm." captionText-4="FIGURES 20 – 25. Diagnostic structures in P. nectoctenus: 20) right proleg, 21) right antenna, 22) male genitalia in dorsal view, 23) male genitalia in lateral view, 24) spiculum gastrale, 25) female genital plates. Scale lines 1 mm." captionText-5="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." captionText-6="FIGURES 36 – 41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm." captionText-7="FIGURES 42 – 47. Dorsal view of Pachystethus species: 42 a, b) P. viduus males; 42 c) P. viduus female; 42 d) P. viduus female, lectotype; 43 a) P. nectoctenus male; 43 b) P. nectoctenus female; 44 a) P. crassesculptus male; 44 b) P. crassesculptus female; 45 a, b) P. nutans males; 45 c) P. nu t an s female; 46 a, b) P. matzapatlecus males; 46 c) P. matzapatlecus female; 47 a) P. ixtacomitanus male; 47 b) P. ixtacomitanus female. Scale lines 1 mm." captionText-8="FIGURES 48 – 53. Lateral view of Pachystethus species: 48) P. viduus a) male, b) female; 49) P. nectoctenus a) male, b) female; 50) P. crassesculptus male; 51) P. n u t a n s male; 52) P. matzapatlecus male; 53) P. ixtacomitanus male. Scale lines 1 mm." captionText-9="FIGURE 54. Distribution of species of Pachystethus." httpUri-0="https://zenodo.org/record/210543/files/figure.png" httpUri-1="https://zenodo.org/record/210544/files/figure.png" httpUri-2="https://zenodo.org/record/210545/files/figure.png" httpUri-3="https://zenodo.org/record/210546/files/figure.png" httpUri-4="https://zenodo.org/record/210547/files/figure.png" httpUri-5="https://zenodo.org/record/210548/files/figure.png" httpUri-6="https://zenodo.org/record/210549/files/figure.png" httpUri-7="https://zenodo.org/record/210550/files/figure.png" httpUri-8="https://zenodo.org/record/210551/files/figure.png" httpUri-9="https://zenodo.org/record/210552/files/figure.png" pageId="1" pageNumber="2">Figs. 1–54</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="65D420B25851FF89FF64A62353EB8080" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="2D7173395851FF89FF64A623516B80A4" blockId="1.[151,1437,1143,2032]" box="[151,812,1143,1168]" pageId="1" pageNumber="2">
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<emphasis id="1FBAAF2B5851FF89FF64A623536C80A4" bold="true" box="[151,299,1143,1168]" pageId="1" pageNumber="2">
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<typeStatus id="F275CD9B5851FF89FF64A623529580A4" box="[151,210,1144,1168]" pageId="1" pageNumber="2">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="EACE08BA5851FF89FECAA62350CD80BB" authority="Newman, 1838" authorityName="Newman" authorityYear="1838" box="[313,650,1143,1168]" class="Insecta" family="Rutelidae" genus="Popillia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="vidua">
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<emphasis id="1FBAAF2B5851FF89FECAA623539F80A4" box="[313,472,1144,1168]" italics="true" pageId="1" pageNumber="2">Popillia vidua</emphasis>
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<bibRefCitation id="495F0EC85851FF89FE2CA62350CD80BB" author="Newman" box="[479,650,1143,1167]" pageId="1" pageNumber="2" refString="Newman, E. F. L. S. (1838) Descriptions of new species of Popillia. The Magazine of Natural History, 2, 336 - 392." type="journal article" year="1838">Newman, 1838</bibRefCitation>
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</taxonomicName>
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by monotypy.
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</paragraph>
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<paragraph id="2D7173395851FF89FF3AA6C053EB8080" blockId="1.[151,1437,1143,2032]" box="[201,428,1179,1204]" pageId="1" pageNumber="2">
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<emphasis id="1FBAAF2B5851FF89FF3AA6C053658080" bold="true" box="[201,290,1179,1204]" pageId="1" pageNumber="2">Gender</emphasis>
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. Masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="65D420B25851FF8AFF3AA6E453D885C0" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
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<paragraph id="2D7173395851FF8AFF3AA6E453D885C0" blockId="1.[151,1437,1143,2032]" lastBlockId="2.[151,1437,151,500]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<emphasis id="1FBAAF2B5851FF89FF3AA6E4531380EC" bold="true" box="[201,340,1215,1240]" pageId="1" pageNumber="2">Description</emphasis>
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. Total body length
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<quantity id="EA36DEDC5851FF89FDDCA6E450DB80E3" box="[559,668,1215,1239]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="1" pageNumber="2" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">7–12 mm</quantity>
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, humeral width 4.0–
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<quantity id="EA36DEDC5851FF89FC77A6E4519B80E3" box="[900,988,1215,1239]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="2" unit="mm" value="4.5">4.5 mm</quantity>
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. Body robust, slightly depressed, dorsal surface shiny and glabrous. Head: clypeus subtrapezoidal or subrectangular, borders slightly upturned, disc surface rugopunctate, coarse and dense punctures. Frons with weak, wide central protuberance and small lateral depressions between extremes of frontoclypeal suture and the base of ocular canthus; frontoclypeal suture straight (best seen in species with contrasting coloration, but hidden under irregular texture and dark color in other species). Eyes small, ovate, circumocular depression accentuated. Ocular canthus wide and glabrous. Mouthparts (
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<figureCitation id="B5F56FBC5851FF89FA90A72F52A5819B" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="1" pageNumber="2">Figs. 32–35</figureCitation>
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): fully developed; mandibles with three teeth, wide molar area; maxilla with six teeth, long, straight, sharply pointed, not fused. Antennal club long (as long as the length of all preceding antennomeres combined) with lamella straight or distally curved inward. Pronotum subtrapezoidal, weakly convex; one small, lateral pit at each side; lateral borders slightly angled or rounded; posterior border without margin; posterior angles straight; anterior border with membranous rim, anterior angles rounded; basal section wider than mesial, as wide as elytral base; disc surface with scattered minute punctures or with irregular, large, deep punctures. Distance between mesocoxae wide, meso-metasternal process well developed, wide, rounded at apex, directed forward or slightly directed downward; overhanging the bases of mesofemora; meso-metasternal suture not evident. Elytral surface weakly marked by discontinuous, small punctures or deeply marked by striae; elytral suture flattened or partially raised, spine-like apex; preapical slope shortened; humeral and apical calli well developed; epipleural border as long as 2/ 3 of total length of elytral border (epipleural border in females thickened and expanded laterally).
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<taxonomicName id="EACE08BA5851FF89FAD9A55B57DB832C" box="[1322,1436,1792,1816]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
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flattened, glabrous, acute apex. Abdomen depressed, anal plate in males with apical margin entire. Propygidium striate, completely covered with elytra. Pygidium uniformly convex, surface striate, partially covered by apex of elytra, apical border acute, with long setae. Protibiae bidentate in both sexes. Protarsomeres 1–4 shortened; protarsomere 5 with ventral denticle not evident; protarsomeres 3–5 with ventrolateral longitudinal keels; protarsal empodia with two long setae, onychium partially hidden; apex of protarsal inner claw deeply cleft, lower denticle more than two times wider than upper denticle; apex of protarsal outer claw entire, sharply acute but with basal part swollen. Mesotibiae and metatibiae slightly swollen at middle. Apex of mesotarsal outer claw narrowly cleft, apex of mesotarsal inner claw entire. Apex of both metatarsal claws entire. Male genitalia: parameres much shorter than basal piece, sagitally separated and each one divided in an upper part more or less upturned apically; lower part fused with ventral plate. Tectum with prominent basal calli. Spiculum gastrale “Y” shaped, basal part straight and shorter than lateral sclerites; associated plates proportionally larger. Sexual dimorphism: males with protarsomere 1 reduced, as long as the length of protarsomeres 2–4 combined. Protibia widened with lateroapical denticle shortened. Antennal club longer than length of all other antennomeres combined. Females: protarsomere 1 long, longer than the length of protarsomeres 2–4 combined. Protibia not widened with lateroapical denticle enlarged. Antennal club as long as length of the preceding five antennomeres combined. Frequently each sex displays different color patterns.
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</paragraph>
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</subSubSection>
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<caption id="79B123B15852FF8AFF64A59751E883D6" box="[151,943,1996,2019]" httpUri="https://zenodo.org/record/210543/files/figure.png" pageId="2" pageNumber="3" targetBox="[183,1336,544,1995]" targetPageId="2">
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<paragraph id="2D7173395852FF8AFF64A59751E883D6" blockId="2.[151,943,1996,2019]" box="[151,943,1996,2019]" pageId="2" pageNumber="3">
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<emphasis id="1FBAAF2B5852FF8AFF64A597534B83D5" bold="true" box="[151,268,1996,2019]" pageId="2" pageNumber="3">FIGURE 1</emphasis>
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. Dorsal habitus of
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<taxonomicName id="EACE08BA5852FF8AFE3EA59750AD83D6" box="[461,746,1996,2018]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="matzapatlecus" status="sp. nov.">
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<emphasis id="1FBAAF2B5852FF8AFE3EA59750AD83D6" box="[461,746,1996,2018]" italics="true" pageId="2" pageNumber="3">Pachystethus matzapatlecus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="048912505852FF8AFD02A596512A83D6" box="[753,877,1996,2018]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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, male.
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</paragraph>
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</caption>
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<subSubSection id="65D420B25853FF8BFF3AA2CC56088623" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="2D7173395853FF8BFF3AA2CC56088623" blockId="3.[151,1436,151,1400]" pageId="3" pageNumber="4">
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<emphasis id="1FBAAF2B5853FF8BFF3AA2CC53048484" bold="true" box="[201,323,151,176]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
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<taxonomicName id="EACE08BA5853FF8BFEA3A2CC538F849B" box="[336,456,151,175]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
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with body robust, slightly depressed, dorsal surface shiny and glabrous; protarsus shortened and swollen; protarsomere 5 without or with very reduced ventral denticle; meso-metasternal intercoxal region wide and increased projection; male antennal club as long as the length of all preceding antennomeres combined; epipleural border widened; and parameres sagittally separated. Species of
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<taxonomicName id="EACE08BA5853FF8BFB99A35F56B98528" box="[1130,1278,260,284]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="1FBAAF2B5853FF8BFB99A35F56B98528" box="[1130,1278,260,284]" italics="true" pageId="3" pageNumber="4">Pachystethus</emphasis>
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</taxonomicName>
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are separated from species of
|
||
<taxonomicName id="EACE08BA5853FF8BFEBAA37353888574" box="[329,463,296,320]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFEBAA37353888574" box="[329,463,296,320]" italics="true" pageId="3" pageNumber="4">Callistethus</emphasis>
|
||
</taxonomicName>
|
||
based on the meso-metasternal process that does not reach the procoxae and its apex is not acute and frequently directed upwards. Species of
|
||
<taxonomicName id="EACE08BA5853FF8BFCE1A31751E18550" box="[786,934,332,356]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFCE1A31751E18550" box="[786,934,332,356]" italics="true" pageId="3" pageNumber="4">Pachystethus</emphasis>
|
||
</taxonomicName>
|
||
are separated from species of
|
||
<taxonomicName id="EACE08BA5853FF8BFAE3A31757DB8550" box="[1296,1436,332,356]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFAE3A31757DB8550" box="[1296,1436,332,356]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
based on the absence of posterior pronotal margin (present in
|
||
<taxonomicName id="EACE08BA5853FF8BFCB9A32B519185BC" box="[842,982,368,392]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFCB9A32B519185BC" box="[842,982,368,392]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
), protarsomeres shortened and widened, the length of first to fourth protarsomeres is similar to fifth (protarsomeres not shortened nor widened in
|
||
<taxonomicName id="EACE08BA5853FF8BFF64A3E3536485E4" box="[151,291,440,464]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFF64A3E3536485E4" box="[151,291,440,464]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
, the length of the first to fourth protarsomere clearly longer than fifth), apex of elitral suture spiniform (apex of elitral suture rouded in
|
||
<taxonomicName id="EACE08BA5853FF8BFDF2A38750CA85C0" box="[513,653,476,500]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFDF2A38750CA85C0" box="[513,653,476,500]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
), flattened body (body not flattened in
|
||
<taxonomicName id="EACE08BA5853FF8BFBB5A387569585C0" box="[1094,1234,476,500]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFBB5A387569585C0" box="[1094,1234,476,500]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
) and male genital capsule markedly angled (male genital capsule not markedly angled in
|
||
<taxonomicName id="EACE08BA5853FF8BFC40A05B5678862C" box="[947,1087,512,536]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFC40A05B5678862C" box="[947,1087,512,536]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B25853FF8BFF3AA07851938608" box="[201,980,547,572]" pageId="3" pageNumber="4" type="distribution">
|
||
<paragraph id="2D7173395853FF8BFF3AA07851938608" blockId="3.[151,1436,151,1400]" box="[201,980,547,572]" pageId="3" pageNumber="4">
|
||
<emphasis id="1FBAAF2B5853FF8BFF3AA07853258608" bold="true" box="[201,354,547,572]" pageId="3" pageNumber="4">Distribution.</emphasis>
|
||
Eastern and southeastern
|
||
<collectingCountry id="55D933A95853FF8BFD74A07F509B8608" box="[647,732,548,572]" name="Mexico" pageId="3" pageNumber="4">Mexico</collectingCountry>
|
||
to central
|
||
<collectingCountry id="55D933A95853FF8BFCA0A07F51978608" box="[851,976,548,572]" name="Guatemala" pageId="3" pageNumber="4">Guatemala</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B25853FF8BFF3AA013532B8143" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="2D7173395853FF8BFF3AA013510087D8" blockId="3.[151,1436,151,1400]" pageId="3" pageNumber="4">
|
||
<emphasis id="1FBAAF2B5853FF8BFF3AA01353718654" bold="true" box="[201,310,584,608]" pageId="3" pageNumber="4">Remarks</emphasis>
|
||
. The unusual and complicated history of these species and genera is mainly derived from the misidentification of sexes that exhibit a marked dimorphism in coloration, which caused nomenclatural complications as stated by
|
||
<bibRefCitation id="495F0EC85853FF8BFE3FA0CB502E8693" author="Arrow" box="[460,617,655,680]" pageId="3" pageNumber="4" refString="Arrow, G. J. (1899) On sexual dimorphism in beetles of the family Rutelidae. Transactions of the Entomological Society of London, 1899, 255 - 269." type="journal article" year="1899">Arrow (1899)</bibRefCitation>
|
||
and that will be discussed and resolved in next sections. Besides being synonymized with
|
||
<taxonomicName id="EACE08BA5853FF8BFE98A0EF53B086F8" box="[363,503,692,716]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFE98A0EF53B086F8" box="[363,503,692,716]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="495F0EC85853FF8BFDF7A0EF508786F8" author="Lacordaire" box="[516,704,692,716]" pageId="3" pageNumber="4" refString="Lacordaire, T. (1856) Histoire Naturelle des Insectes. Genera des Coleopteres ou Expos Methodique et Critique de Tous les Genres, Proposes Jusqu'ici dans cet Ordre d'Insectes. Tome 3. Librairie Enciclopedique de Roret, Paris." type="book chapter" year="1856">Lacordaire 1856</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFD3FA0EE510C86F8" author="Bates" box="[716,843,692,716]" pageId="3" pageNumber="4" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin & F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates 1888</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFCA5A0EF519886F8" author="Ohaus" box="[854,991,692,716]" pageId="3" pageNumber="4" refString="Ohaus, F. (1918) Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus, pars. 66. W. Junk, Berlin." type="book chapter" year="1918">Ohaus 1918</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFC1AA0EF56FF86F8" author="Blackwelder" box="[1001,1208,692,716]" pageId="3" pageNumber="4" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America, Part 2. Bulletin of the United States National Museum, 185, 189 - 341." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFB30A0EF57DF86F8" author="Machatschke" box="[1219,1432,692,716]" pageId="3" pageNumber="4" refString="Machatschke, J. W. (1972) Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa, 66, 363 - 429." type="journal article" year="1972">Machatschke 1972</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFF64A083534886DB" author="Potts" box="[151,271,727,751]" pageId="3" pageNumber="4" refString="Potts, R. L. (1974) Revision of the Scarabaeidae: Anomalinae 1. The genera occurring in the United States and Canada. The Pan-Pacific Entomologist, 50, 148 - 154." type="journal article" year="1974">Potts 1974</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="EACE08BA5853FF8BFEBCA083539286C4" box="[335,469,728,752]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFEBCA083539286C4" box="[335,469,728,752]" italics="true" pageId="3" pageNumber="4">Callistethus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="495F0EC85853FF8BFE17A08C50FB86DB" author="Machatschke" box="[484,700,727,751]" pageId="3" pageNumber="4" refString="Machatschke, J. W. (1957) Coleoptera Lamellicornia, Scarabaeidae, Rutelinae, Anomalini. Genera Insectorum, 199 B, 1 - 219." type="journal article" year="1957">Machatschke 1957</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFD34A08C511186DB" author="Moron" box="[711,854,727,751]" pageId="3" pageNumber="4" refString="Moron, M. A. (1997) Rutelinae, pp. 9 - 49. In: M. A. Moron, B. C. Ratcliffe, & C. Deloya. Atlas de los escarabajos de Mexico. Coleoptera Lamelicornia. Vol. I. Familia Melolonthidae. CONABIO y Sociedad Mexicana de Entomologia, A. C. Mexico." type="book chapter" year="1997">Morón 1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFC91A08C566786DB" author="Mico" box="[866,1056,727,752]" pageId="3" pageNumber="4" refString="Mico, E. B., Moron, M. A., & Galante, E. (2003) New larval descriptions and biology of some New World Anomalini beetles (Scarabaeidae: Rutelinae). Annals of the Entomological Society of America, 96, 597 - 614." type="journal article" year="2003">
|
||
Micó
|
||
<emphasis id="1FBAAF2B5853FF8BFC57A082519186C4" box="[932,982,728,752]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
), the genus
|
||
<taxonomicName id="EACE08BA5853FF8BFB54A083577C86C4" box="[1191,1339,728,752]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFB54A083577C86C4" box="[1191,1339,728,752]" italics="true" pageId="3" pageNumber="4">Pachystethus</emphasis>
|
||
</taxonomicName>
|
||
has also been synonymized with
|
||
<taxonomicName id="EACE08BA5853FF8BFE54A0A750D18720" authority="Casey 1915" authorityName="Casey" authorityYear="1915" box="[423,662,764,788]" class="Insecta" family="Scarabaeidae" genus="Spilota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFE54A0A753B18720" box="[423,502,764,788]" italics="true" pageId="3" pageNumber="4">Spilota</emphasis>
|
||
(
|
||
<bibRefCitation id="495F0EC85853FF8BFDF6A0A750C98720" author="Casey" box="[517,654,764,788]" pageId="3" pageNumber="4" refString="Casey T. L. (1915) A review of the American species of Rutelinae, Dynastinae and Cetoniinae. Memoirs on the Coleoptera, 6, 1 - 460." type="journal article" year="1915">Casey 1915</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, placed as a subgenus of
|
||
<taxonomicName id="EACE08BA5853FF8BFC42A0A756E38720" authority="Frey 1968" authorityName="Frey" authorityYear="1968" box="[945,1188,764,788]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFC42A0A756518720" box="[945,1046,764,788]" italics="true" pageId="3" pageNumber="4">Anomala</emphasis>
|
||
(
|
||
<bibRefCitation id="495F0EC85853FF8BFBD7A0A756DC8720" author="Frey" box="[1060,1179,764,788]" pageId="3" pageNumber="4" refString="Frey, G. (1968) Bestimmungstabelle der Zentral- und Sudamerikanischen arten der untergattung Spilota der gattung Anomala nebst einigen neubeschreibungen Entomologische Arbeiten aus dem museum G. Frey, 19, 281 - 295." type="journal article" year="1968">Frey 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, or synonymized with
|
||
<taxonomicName id="EACE08BA5853FF8BFF64A17B535A870C" box="[151,285,800,824]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFF64A17B535A870C" box="[151,285,800,824]" italics="true" pageId="3" pageNumber="4">Callistethus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="495F0EC85853FF8BFED8A14450458703" author="Machatschke" box="[299,514,799,823]" pageId="3" pageNumber="4" refString="Machatschke, J. W. (1957) Coleoptera Lamellicornia, Scarabaeidae, Rutelinae, Anomalini. Genera Insectorum, 199 B, 1 - 219." type="journal article" year="1957">Machatschke 1957</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFDFEA14450DD8703" author="Moron" box="[525,666,799,823]" pageId="3" pageNumber="4" refString="Moron, M. A. (1997) Rutelinae, pp. 9 - 49. In: M. A. Moron, B. C. Ratcliffe, & C. Deloya. Atlas de los escarabajos de Mexico. Coleoptera Lamelicornia. Vol. I. Familia Melolonthidae. CONABIO y Sociedad Mexicana de Entomologia, A. C. Mexico." type="book chapter" year="1997">Morón 1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="495F0EC85853FF8BFD56A14451258703" author="Mico" box="[677,866,799,824]" pageId="3" pageNumber="4" refString="Mico, E. B., Moron, M. A., & Galante, E. (2003) New larval descriptions and biology of some New World Anomalini beetles (Scarabaeidae: Rutelinae). Annals of the Entomological Society of America, 96, 597 - 614." type="journal article" year="2003">
|
||
Micó
|
||
<emphasis id="1FBAAF2B5853FF8BFD15A17A5150870C" box="[742,791,800,824]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
). Among
|
||
<taxonomicName id="EACE08BA5853FF8BFC3DA17B5613870C" box="[974,1108,800,824]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFC3DA17B5613870C" box="[974,1108,800,824]" italics="true" pageId="3" pageNumber="4">Callistethus</emphasis>
|
||
</taxonomicName>
|
||
and some species considered as
|
||
<taxonomicName id="EACE08BA5853FF8BFF45A11F53428768" box="[182,261,836,860]" class="Insecta" family="Scarabaeidae" genus="Spilota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFF45A11F53428768" box="[182,261,836,860]" italics="true" pageId="3" pageNumber="4">Spilota</emphasis>
|
||
</taxonomicName>
|
||
(as well as with other genera like
|
||
<taxonomicName id="EACE08BA5853FF8BFD7CA11F515D8768" box="[655,794,836,860]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFD7CA11F515D8768" box="[655,794,836,860]" italics="true" pageId="3" pageNumber="4">Strigoderma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EACE08BA5853FF8BFCD9A11E51FE8768" box="[810,953,837,860]" class="Insecta" family="Rutelidae" genus="Epectinaspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFCD9A11E51FE8768" box="[810,953,837,860]" italics="true" pageId="3" pageNumber="4">Epectinaspis</emphasis>
|
||
</taxonomicName>
|
||
, or
|
||
<taxonomicName id="EACE08BA5853FF8BFC14A11F56218768" box="[999,1126,836,860]" class="Insecta" family="Rutelidae" genus="Callirhinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFC14A11F56218768" box="[999,1126,836,860]" italics="true" pageId="3" pageNumber="4">Callirhinus</emphasis>
|
||
</taxonomicName>
|
||
), morphological characters that
|
||
<taxonomicName id="EACE08BA5853FF8BFF39A133531987B4" box="[202,350,872,896]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFF39A133531987B4" box="[202,350,872,896]" italics="true" pageId="3" pageNumber="4">Pachystethus</emphasis>
|
||
</taxonomicName>
|
||
shares with these genera are the prominent and wide meso-metasternal process, depressed and robust body, elytra reduced towards apex, apex of elytral suture spiniform, pronotum without posterior margin, and evident sexual dimorphism. Therefore, it might be more appropriate to assume that
|
||
<taxonomicName id="EACE08BA5853FF8BFBC4A1EB568C87FC" box="[1079,1227,944,968]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFBC4A1EB568C87FC" box="[1079,1227,944,968]" italics="true" pageId="3" pageNumber="4">Pachystethus</emphasis>
|
||
</taxonomicName>
|
||
would have closer affinity with some of the latter genera than with
|
||
<taxonomicName id="EACE08BA5853FF8BFD47A18F510787D8" box="[692,832,980,1004]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFD47A18F510787D8" box="[692,832,980,1004]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2D7173395853FF8BFF3AA1A3532B8143" blockId="3.[151,1436,151,1400]" pageId="3" pageNumber="4">
|
||
<emphasis id="1FBAAF2B5853FF8BFF3AA1A353588024" bold="true" box="[201,287,1016,1040]" pageId="3" pageNumber="4">Larvae</emphasis>
|
||
. According to the description provided by
|
||
<bibRefCitation id="495F0EC85853FF8BFCF0A1AC5188803B" box="[771,975,1015,1040]" pageId="3" pageNumber="4" refString="Mico, E. B., Moron, M. A., & Galante, E. (2003) New larval descriptions and biology of some New World Anomalini beetles (Scarabaeidae: Rutelinae). Annals of the Entomological Society of America, 96, 597 - 614." type="journal article">
|
||
Micó
|
||
<emphasis id="1FBAAF2B5853FF8BFCB6A1A251318024" box="[837,886,1016,1040]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
|
||
. (2003)
|
||
</bibRefCitation>
|
||
the diagnostic characters for the larva of
|
||
<taxonomicName id="EACE08BA5853FF8BFF64A64753F28000" box="[151,437,1052,1076]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nectoctenus">
|
||
<emphasis id="1FBAAF2B5853FF8BFF64A64753F28000" box="[151,437,1052,1076]" italics="true" pageId="3" pageNumber="4">Pachystethus nectoctenus</emphasis>
|
||
</taxonomicName>
|
||
(cited as
|
||
<taxonomicName id="EACE08BA5853FF8BFDD3A64750AD8000" box="[544,746,1052,1076]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vidua">
|
||
<emphasis id="1FBAAF2B5853FF8BFDD3A64750AD8000" box="[544,746,1052,1076]" italics="true" pageId="3" pageNumber="4">Callistethus vidua</emphasis>
|
||
</taxonomicName>
|
||
) are: head with two posterior frontal setae on each side; dorsa of abdominal segments VII–IX each with poorly-defined transverse rows of slender long setae, alternated with medium-sized setae; semicircular furrow on dorsum of last abdominal segment nearly absent; lower anal lip with 20–21 setae; raster with 9–12 acute, slender pali on each palidium, palidia diverging posteriorly, and with 25 preseptular hamate setae. It is possible that the development of semicircular furrow on abdominal segment X may be an important generic character for
|
||
<taxonomicName id="EACE08BA5853FF8BFDA5A68B509680DC" box="[598,721,1232,1256]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Anomalini">Anomalini</taxonomicName>
|
||
. Main differences with larvae of
|
||
<emphasis id="1FBAAF2B5853FF8BFBADA68B572680DC" box="[1118,1377,1232,1256]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="EACE08BA5853FF8BFBADA68B56AA80DC" box="[1118,1261,1232,1256]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Paranomala</taxonomicName>
|
||
,
|
||
<taxonomicName id="EACE08BA5853FF8BFB0FA68B572680DC" box="[1276,1377,1232,1256]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="EACE08BA5853FF8BFF64A6AF53658138" box="[151,290,1268,1292]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFF64A6AF53658138" box="[151,290,1268,1292]" italics="true" pageId="3" pageNumber="4">Strigoderma</emphasis>
|
||
</taxonomicName>
|
||
are in the distribution of setae on the dorsa of abdominal segments VII–IX, the presence of dorsal furrow on last abdominal segment, and composition and form of palidia. But larval descriptions of additional species of
|
||
<emphasis id="1FBAAF2B5853FF8BFEE2A76750918160" box="[273,726,1340,1364]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="EACE08BA5853FF8BFEE2A76753E08160" box="[273,423,1340,1364]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Pachystethus</taxonomicName>
|
||
,
|
||
<taxonomicName id="EACE08BA5853FF8BFE46A76750048160" box="[437,579,1340,1364]" class="Insecta" family="Rutelidae" genus="Strigoderma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Strigoderma</taxonomicName>
|
||
,
|
||
<taxonomicName id="EACE08BA5853FF8BFDA3A76750918160" box="[592,726,1340,1364]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Callistethus</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="EACE08BA5853FF8BFCEAA76751E28160" box="[793,933,1340,1364]" class="Insecta" family="Rutelidae" genus="Paranomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B5853FF8BFCEAA76751E28160" box="[793,933,1340,1364]" italics="true" pageId="3" pageNumber="4">Paranomala</emphasis>
|
||
</taxonomicName>
|
||
are required before precise larval diagnosis may be completed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |