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<document id="8DD68669AD1D9A8419C59996A204DE59" ID-DOI="10.5281/zenodo.210542" ID-GBIF-Dataset="c9e5cc54-2a89-4b75-91aa-a4fce79f17fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="210542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460500319311" checkinUser="plazi" docAuthor="Ramírez-Ponce, Andrés &amp; Morón, Miguel Angel" docDate="2012" docId="A567C22F5841FF9CFF64A2CC57608297" docLanguage="en" docName="zt03394p024.pdf" docOrigin="Zootaxa 3394" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pachystethus ixtacomitanus Ramirez-Ponce &amp; Moron, new species" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="595EBA575850FF88FFF3A25B52478434" masterDocTitle="Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="18" updateTime="1698311598655" updateUser="plazi">
<mods:mods id="45BA12F62180B565F713902F8FB26F0A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3646E520D44A7E520691FDD615236B0F">
<mods:title id="1DC7937B59DF1E893233EA827FCBAB36">Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)</mods:title>
</mods:titleInfo>
<mods:name id="9345AB6957E03DC3162C5A972CB6B4F1" type="personal">
<mods:role id="F6B32D70D7D5E9B0876A3742BCDA06FE">
<mods:roleTerm id="CB541025D87D33E28C36568BE948E7A6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7CFE72A91A9B532E13D544B887F188A1">Ramírez-Ponce, Andrés</mods:namePart>
</mods:name>
<mods:name id="35C65D3D195346729260C4B249E13A9B" type="personal">
<mods:role id="C2FF6ED2399310CF534A41574AD9E8A8">
<mods:roleTerm id="C9991E8C7F24C9D56A51F5F3C1B21A22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B34148AC8154E621A7D76853FFFC47AD">Morón, Miguel Angel</mods:namePart>
</mods:name>
<mods:typeOfResource id="CEDE9657EC614F6CDACFEB1676B16673">text</mods:typeOfResource>
<mods:relatedItem id="1839F01398161D20F7D04341A5002AC8" type="host">
<mods:titleInfo id="FBF893191F3500D071F3383CAD75A9C9">
<mods:title id="364691BEEEA8C4482F79C9AFCD1AAFA6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9BAB7F8AF69388FA097F02E8173E56A9">
<mods:date id="3513C9D182C8D30AB333E7CFA80F2937">2012</mods:date>
<mods:detail id="98FFC5E64329CC453A39C380F7325F46" type="volume">
<mods:number id="D544CE2537FC95A60F51351A12F19612">3394</mods:number>
</mods:detail>
<mods:extent id="8DFDDF84094EF05EAB64B35753D0B711" unit="page">
<mods:start id="2F7609604BD301657276E3FC8784A169">1</mods:start>
<mods:end id="3239A9AB58E775116B48103AA99A8F66">24</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7997DFC8AAA5D23EBF843386FF651FB4">journal article</mods:classification>
<mods:identifier id="C634D8A320365EFBBDD29E87D3E96062" type="DOI">10.5281/zenodo.210542</mods:identifier>
<mods:identifier id="0F9396DE29C292D6442AEC1F1FDD37F0" type="GBIF-Dataset">c9e5cc54-2a89-4b75-91aa-a4fce79f17fc</mods:identifier>
<mods:identifier id="5890A21B9EADB5230E4412B821010EDC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3C5198C0EC12EFEF60FCB03E4365AA8D" type="Zenodo-Dep">210542</mods:identifier>
</mods:mods>
<treatment id="A567C22F5841FF9CFF64A2CC57608297" ID-DOI="http://doi.org/10.5281/zenodo.6179204" ID-GBIF-Taxon="119521056" ID-Zenodo-Dep="6179204" LSID="urn:lsid:plazi:treatment:A567C22F5841FF9CFF64A2CC57608297" httpUri="http://treatment.plazi.org/id/A567C22F5841FF9CFF64A2CC57608297" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18">
<subSubSection id="65D420B25841FF99FF64A2CC53ED84E6" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="2D7173395841FF99FF64A2CC518D8485" blockId="17.[151,970,151,210]" box="[151,970,151,177]" pageId="17" pageNumber="18">
<heading id="7639C4555841FF99FF64A2CC518D8485" bold="true" box="[151,970,151,177]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="1FBAAF2B5841FF99FF64A2CC518D8485" bold="true" box="[151,970,151,177]" pageId="17" pageNumber="18">
<taxonomicName id="EACE08BA5841FF99FF64A2CC51698485" authority="Ramirez-Ponce &amp; Moron" authorityName="Ramirez-Ponce &amp; Moron" box="[151,814,151,177]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ixtacomitanus" status="sp. nov.">
<emphasis id="1FBAAF2B5841FF99FF64A2CC53AF8485" bold="true" box="[151,488,151,177]" italics="true" pageId="17" pageNumber="18">Pachystethus ixtacomitanus</emphasis>
Ramírez-Ponce &amp; Morón
</taxonomicName>
,
<taxonomicNameLabel id="048912505841FF99FCC8A2CC518D8485" box="[827,970,151,177]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2D7173395841FF99FF64A2E153ED84E6" blockId="17.[151,970,151,210]" box="[151,426,186,210]" pageId="17" pageNumber="18">
(
<figureCitation id="B5F56FBC5841FF99FF6CA2E1536084E6" box="[159,295,186,210]" captionStart="FIGURES 36 – 41" captionStartId="18.[151,264,1619,1642]" captionTargetBox="[189,1372,224,1565]" captionTargetId="figure@18.[151,1436,193,1598]" captionTargetPageId="18" captionText="FIGURES 36 – 41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210549/files/figure.png" pageId="17" pageNumber="18">Figs. 36–41</figureCitation>
,
<figureCitation id="B5F56FBC5841FF99FEC1A2E1530884E6" box="[306,335,186,210]" captionStart="FIGURES 42 – 47" captionStartId="19.[151,264,1909,1932]" captionTargetBox="[214,1377,207,1832]" captionTargetId="figure@19.[198,1392,193,1847]" captionTargetPageId="19" captionText="FIGURES 42 – 47. Dorsal view of Pachystethus species: 42 a, b) P. viduus males; 42 c) P. viduus female; 42 d) P. viduus female, lectotype; 43 a) P. nectoctenus male; 43 b) P. nectoctenus female; 44 a) P. crassesculptus male; 44 b) P. crassesculptus female; 45 a, b) P. nutans males; 45 c) P. nu t an s female; 46 a, b) P. matzapatlecus males; 46 c) P. matzapatlecus female; 47 a) P. ixtacomitanus male; 47 b) P. ixtacomitanus female. Scale lines 1 mm." httpUri="https://zenodo.org/record/210550/files/figure.png" pageId="17" pageNumber="18">47</figureCitation>
a–b, 53)
</paragraph>
</subSubSection>
<subSubSection id="65D420B25841FF99FF64A2A652BD86DD" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="2D7173395841FF99FF64A2A652BD86DD" blockId="17.[151,1437,253,2022]" pageId="17" pageNumber="18">
<emphasis id="1FBAAF2B5841FF99FF64A2A653078522" bold="true" box="[151,320,253,278]" pageId="17" pageNumber="18">
<typeStatus id="F275CD9B5841FF99FF64A2A652958521" box="[151,210,253,277]" pageId="17" pageNumber="18">Type</typeStatus>
material
</emphasis>
(
<specimenCount id="3BC8B8B05841FF99FEA0A2A653FA8521" box="[339,445,253,277]" pageId="17" pageNumber="18" type="male">12 males</specimenCount>
,
<specimenCount id="3BC8B8B05841FF99FE38A2A6500A8521" box="[459,589,253,277]" pageId="17" pageNumber="18" type="female">10 females</specimenCount>
).
<emphasis id="1FBAAF2B5841FF99FD90A2A650978521" bold="true" box="[611,720,253,277]" pageId="17" pageNumber="18">
<typeStatus id="F275CD9B5841FF99FD90A2A650978521" box="[611,720,253,277]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
male housed at
<collectionCode id="4BDFEBFC5841FF99FC62A2A651928521" box="[913,981,253,277]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">CNIN</collectionCode>
with following label data: a) “
<collectingCountry id="55D933A95841FF99FACEA2A657DE8521" box="[1341,1433,253,277]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
: Chiapas, Ixtacomitán, Río Blanco,
<date id="597055F95841FF99FDEDA37A50D4850D" box="[542,659,289,313]" pageId="17" pageNumber="18" value="1985-12-10">10-XII-85</date>
. Col. F. Arias, L. Cervantes y R. Barba” (typeface), b) “Colección del Instituto de Biología,
<collectionCode id="4BDFEBFC5841FF99FE66A31D53AA856A" box="[405,493,326,350]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">UNAM</collectionCode>
,
<collectingCountry id="55D933A95841FF99FE0FA31E50138569" box="[508,596,325,349]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
, D. F.” (typeface), c) our
<typeStatus id="F275CD9B5841FF99FC75A31E51AF8569" box="[902,1000,325,349]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
red label.
<emphasis id="1FBAAF2B5841FF99FB9BA31E568A856A" bold="true" box="[1128,1229,325,350]" pageId="17" pageNumber="18">Allotype</emphasis>
female housed at
<collectionCode id="4BDFEBFC5841FF99FF64A332529C85B5" box="[151,219,361,385]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">CNIN</collectionCode>
with following label data: a) “
<collectingCountry id="55D933A95841FF99FDB0A33250D885B5" box="[579,671,361,385]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
: Chiapas, Ixtacomitán, Río Blanco,
<date id="597055F95841FF99FBB3A332569685B5" box="[1088,1233,361,385]" pageId="17" pageNumber="18" value="1985-12-10">10-XII-1985</date>
. Col. F. Arias, L. Cervantes y R. Barba” (typeface), b) Instituto de Biología,
<collectionCode id="4BDFEBFC5841FF99FCBDA3D551E18592" box="[846,934,398,422]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">UNAM</collectionCode>
,
<collectingCountry id="55D933A95841FF99FC46A3D6564A8591" box="[949,1037,397,421]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
, D. F.” (typeface), c) our allotype yellow label.
<emphasis id="1FBAAF2B5841FF99FEDCA3EA53E085FD" bold="true" box="[303,423,433,457]" pageId="17" pageNumber="18">
<typeStatus id="F275CD9B5841FF99FEDCA3EA53E085FD" box="[303,423,433,457]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="3BC8B8B05841FF99FE45A3EA505A85FD" box="[438,541,433,457]" pageId="17" pageNumber="18" type="male">11 males</specimenCount>
,
<specimenCount id="3BC8B8B05841FF99FDDAA3EA50D085FD" box="[553,663,433,457]" pageId="17" pageNumber="18" type="female">9 females</specimenCount>
) with our
<typeStatus id="F275CD9B5841FF99FCF8A3E9512C85FE" box="[779,875,434,458]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
yellow labels and following label data: same data as
<typeStatus id="F275CD9B5841FF99FF45A38E535F85D9" box="[182,280,469,493]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(
<specimenCount id="3BC8B8B05841FF99FEDAA38E53C485D9" box="[297,387,469,493]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<specimenCount id="3BC8B8B05841FF99FE7CA38E53B885D9" box="[399,511,469,493]" pageId="17" pageNumber="18" type="female">4 females</specimenCount>
,
<collectionCode id="4BDFEBFC5841FF99FDF8A38E501485D9" box="[523,595,469,493]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">CNIN</collectionCode>
;
<specimenCount id="3BC8B8B05841FF99FDACA38E50FE85D9" box="[607,697,469,493]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<specimenCount id="3BC8B8B05841FF99FD36A38E516C85D9" box="[709,811,469,493]" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
,
<collectionCode id="4BDFEBFC5841FF99FCC5A38D513B85DA" box="[822,892,470,494]" pageId="17" pageNumber="18">IEXA</collectionCode>
;
<specimenCount id="3BC8B8B05841FF99FC7AA38E519F85D9" box="[905,984,469,493]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<collectionCode id="4BDFEBFC5841FF99FC17A38E567F85D9" box="[996,1080,469,493]" pageId="17" pageNumber="18">MXAL</collectionCode>
) (typeface); “
<collectingCountry id="55D933A95841FF99FB27A38E577785D9" box="[1236,1328,469,493]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
: Chiapas, Soluchistan,
<date id="597055F95841FF99FED9A3A253D98625" box="[298,414,505,529]" pageId="17" pageNumber="18" value="1985-12-11">11-XII-85</date>
. F. Arias, L. Cervantes” (typeface) (
<specimenCount id="3BC8B8B05841FF99FCB2A3A251D98625" box="[833,926,505,529]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<collectionCode id="4BDFEBFC5841FF99FC58A3A251B28625" box="[939,1013,505,529]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">CNIN</collectionCode>
); “
<collectingCountry id="55D933A95841FF99FBE6A3A256368625" box="[1045,1137,505,529]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
: Chiapas, Ixhuatán, Km 3 carr. Pozo Rosarito,
<date id="597055F95841FF99FE88A046504C8601" box="[379,523,541,565]" pageId="17" pageNumber="18" value="1985-12-12">12-XII-1985</date>
, F. Arias. R. Barba, L. Cervantes” (typeface) (
<specimenCount id="3BC8B8B05841FF99FBE6A04656258601" box="[1045,1122,541,565]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<collectionCode id="4BDFEBFC5841FF99FB9EA04656F18601" box="[1133,1206,541,565]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">CNIN</collectionCode>
); “
<collectingCountry id="55D933A95841FF99FB27A04657778601" box="[1236,1328,541,565]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
: Chiapas, Pueblo Nuevo Soluchistan. Km. 5 Pueblo Nuevo-Jitotol. 1624 msnm
<date id="597055F95841FF99FC34A01A5609866D" box="[967,1102,577,601]" pageId="17" pageNumber="18" value="2007-06-26">26-VI-2007</date>
.
<geoCoordinate id="48FA15FE5841FF99FBAEA01A56AB866D" box="[1117,1260,577,601]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="17.1425">17°08´33´´N</geoCoordinate>
<geoCoordinate id="48FA15FE5841FF99FB04A01A57DF866D" box="[1271,1432,577,601]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="-92.88778">92°53´16´´ W</geoCoordinate>
. Bosque Pino. L. Cervantes, G. Ortega, C. Mayorga” (typeface) (
<specimenCount id="3BC8B8B05841FF99FC8EA03E519D8649" box="[893,986,613,637]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<specimenCount id="3BC8B8B05841FF99FC1BA03E561C8649" box="[1000,1115,613,637]" pageId="17" pageNumber="18" type="female">3 females</specimenCount>
;
<collectionCode id="4BDFEBFC5841FF99FB9BA03E56F58649" box="[1128,1202,613,637]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">CNIN</collectionCode>
); “
<collectingCountry id="55D933A95841FF99FB21A03E576B8649" box="[1234,1324,613,637]" name="Mexico" pageId="17" pageNumber="18">México</collectingCountry>
, Chiapas, Larraizal, km. 22 Bochil-Larraizal, 1,857 msnm.
<geoCoordinate id="48FA15FE5841FF99FD48A0D251138695" box="[699,852,649,673]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="16.917778">16°55´04´´ N</geoCoordinate>
,
<geoCoordinate id="48FA15FE5841FF99FCACA0D251BB8695" box="[863,1020,649,673]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="-92.76">92° 45´36´´W</geoCoordinate>
.
<date id="597055F95841FF99FBF5A0D256CA8695" box="[1030,1165,649,673]" pageId="17" pageNumber="18" value="2007-06-26">26-VI-2007</date>
, C. Mayorga, G. Ortega, L. Cervantes” (typeface) (
<specimenCount id="3BC8B8B05841FF99FE34A0F6506A86F1" box="[455,557,685,709]" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
,
<collectionCode id="4BDFEBFC5841FF99FDCFA0F650C186F1" box="[572,646,685,709]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="18">CNIN</collectionCode>
); “Chiapas, Rancho Cuducun”, no other data (handwritten) (
<specimenCount id="3BC8B8B05841FF99FABBA0F657D086F1" box="[1352,1431,685,709]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<collectionCode id="4BDFEBFC5841FF99FF64A08A52A886DD" box="[151,239,721,745]" pageId="17" pageNumber="18">MXAL</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="65D420B25841FF9AFF3AA0AE515A8320" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="2D7173395841FF99FF3AA0AE511A8301" blockId="17.[151,1437,253,2022]" pageId="17" pageNumber="18">
<emphasis id="1FBAAF2B5841FF99FF3AA0AE50498739" bold="true" box="[201,526,757,782]" pageId="17" pageNumber="18">
Description.
<typeStatus id="F275CD9B5841FF99FE93A0AE538A8739" box="[352,461,757,781]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
male
</emphasis>
. Length
<quantity id="EA36DEDC5841FF99FD82A0AE508D8739" box="[625,714,757,781]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="17" pageNumber="18" unit="mm" value="9.8">9.8 mm</quantity>
, width
<quantity id="EA36DEDC5841FF99FCE9A0AE51348739" box="[794,883,757,781]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="17" pageNumber="18" unit="mm" value="4.4">4.4 mm</quantity>
. Color reddish yellow on clypeus, anterior half of frons, scape, pedicel, funicle, lateral margins of pronotum, prosternum, center of metasternum, abdomen, and legs; shiny black on antennal club, posterior half of frons, pronotal disc,
<taxonomicName id="EACE08BA5841FF99FC5DA16756598760" box="[942,1054,828,852]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
, elytra, and mesosternum.
<emphasis id="1FBAAF2B5841FF99FAABA16757DA8760" box="[1368,1437,828,852]" italics="true" pageId="17" pageNumber="18">Head.</emphasis>
Clypeus subtrapezoidal, anterior border curved, briefly raised, and not directed forward; anterior angles widely rounded; clypeal surface rugose with abundant, large punctures. Frontoclypeal suture straight, clearly defined. Frons slightly convex, glabrous, densely punctate; puctures rugose at center, more disperse on lateral and posterior margins. Punctures around eyes scattered. Antennal club slightly longer than the length of all preceding antennomeres combined; lamellar shape semiovate, nearly straight (
<figureCitation id="B5F56FBC5841FF99FC5EA1B656438031" box="[941,1028,1005,1029]" captionStart="FIGURES 36 – 41" captionStartId="18.[151,264,1619,1642]" captionTargetBox="[189,1372,224,1565]" captionTargetId="figure@18.[151,1436,193,1598]" captionTargetPageId="18" captionText="FIGURES 36 – 41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210549/files/figure.png" pageId="17" pageNumber="18">Fig. 37</figureCitation>
). Eyes small, ovate, ocular margin dorsally noticeable and setose. Eye canthus thick with glabrous, rounded apex and flattened surface with shallow minute punctures. Maxillary basistipe weakly convex, sparsely punctate; labrum wide and sinuate; labium sinuate, slightly longer than wide; prementum glabrous with irregular, deep pits.
<emphasis id="1FBAAF2B5841FF99FC0CA60256298044" box="[1023,1134,1113,1136]" italics="true" pageId="17" pageNumber="18">Pronotum</emphasis>
wider than long, weakly convex; surface with uniformly distributed, deep, medium size, round punctures; lateral borders angled with bulky sides and large, deep lateral pits; posterior angles not raised. Mesoepimera briefly exposed over humeral calli. Space between mesocoxae wide; mesometasternal process stout, but not very long, briefly projecting anteriorly past mesocoxa with wide base and sharply pointed apex directed forward; metasternum covered with numerous wide punctures, sides moderately setiferous; mesial region of metasternum glabrous with deep longitudinal sulcus.
<emphasis id="1FBAAF2B5841FF99FF64A776529A8171" box="[151,221,1325,1349]" italics="true" pageId="17" pageNumber="18">Elytra</emphasis>
flattened and ovate, striae clearly impressed with nearly continuous rows of large punctures; elytral costae well defined; elytral suture raised on posterior third; brief preapical slope; lateral margins nearly straight with stout, short setae on epipleural area and other slender setae on margin. Epipleural fold narrow, oblique in lateral view, gradually narrowed after metacoxa.
<taxonomicName id="EACE08BA5841FF99FDB7A7CC50F5819B" box="[580,690,1431,1455]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="1FBAAF2B5841FF99FDB7A7CC50F5819B" box="[580,690,1431,1455]" italics="true" pageId="17" pageNumber="18">Scutellum</emphasis>
</taxonomicName>
: nearly longer as wide; lateral margins rounded, apex weakly rounded; surface with uniform, wide, deep punctures.
<emphasis id="1FBAAF2B5841FF99FD02A7E0511C81E7" box="[753,859,1467,1491]" italics="true" pageId="17" pageNumber="18">Abdomen</emphasis>
: sternites with scattered setae on continuous row of large punctures.
<emphasis id="1FBAAF2B5841FF99FEBCA78453C381C2" box="[335,388,1503,1526]" italics="true" pageId="17" pageNumber="18">Legs</emphasis>
: protarsomeres 1–4 short and wide, as wide as protarsomere 5 and as long as it. Protibia stout, progressively widened apically; lateral borders divergent towards apex; external border with proximal denticle short and rounded; distal denticle long, slightly curved (
<figureCitation id="B5F56FBC5841FF99FCF6A47E511E8209" box="[773,857,1573,1597]" captionStart="FIGURES 36 – 41" captionStartId="18.[151,264,1619,1642]" captionTargetBox="[189,1372,224,1565]" captionTargetId="figure@18.[151,1436,193,1598]" captionTargetPageId="18" captionText="FIGURES 36 – 41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210549/files/figure.png" pageId="17" pageNumber="18">Fig. 36</figureCitation>
). Mesofemora with a well-defined longitudinal keel and disperse, setiferous punctures. Metafemora without keel. Metatibia with 2 deep transverse keels and spine-like, short setae; apex of metatibia clearly swollen. Metatarsomeres 1–2 longer than tarsomeres 3–5.
<emphasis id="1FBAAF2B5841FF99FB1BA437571582B0" box="[1256,1362,1644,1668]" italics="true" pageId="17" pageNumber="18">Genitalia</emphasis>
(
<figureCitation id="B5F56FBC5841FF99FA90A43752A58293" captionStart="FIGURES 36 – 41" captionStartId="18.[151,264,1619,1642]" captionTargetBox="[189,1372,224,1565]" captionTargetId="figure@18.[151,1436,193,1598]" captionTargetPageId="18" captionText="FIGURES 36 – 41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210549/files/figure.png" pageId="17" pageNumber="18">Figs. 38–41</figureCitation>
): parameres complex, shorter than the length of tectum, with upper part irregular, narrowed towards apex and weakly upturned; lower part of parameres briefly curved backwards. Basal half of tectum wider than distal half, with outstanding posterolateral calli. Basal piece with long and round, pointed apodema.
<emphasis id="1FBAAF2B5841FF99FB3CA48D57DB82DA" box="[1231,1436,1750,1774]" italics="true" pageId="17" pageNumber="18">Spiculum gastrale</emphasis>
(
<figureCitation id="B5F56FBC5841FF99FF6CA4A252B38325" box="[159,244,1785,1809]" captionStart="FIGURES 36 – 41" captionStartId="18.[151,264,1619,1642]" captionTargetBox="[189,1372,224,1565]" captionTargetId="figure@18.[151,1436,193,1598]" captionTargetPageId="18" captionText="FIGURES 36 – 41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210549/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
) narrow; basal part slightly curved, with uniform width, shorter than lateral sclerites; associated plates small, subovate, with numerous long setae near exterior border.
</paragraph>
<paragraph id="2D7173395841FF99FF3AA51B539F83D2" blockId="17.[151,1437,253,2022]" pageId="17" pageNumber="18">
<emphasis id="1FBAAF2B5841FF99FF3AA51B53C4836D" bold="true" box="[201,387,1856,1881]" pageId="17" pageNumber="18">Allotype female</emphasis>
. Length
<quantity id="EA36DEDC5841FF99FE14A51B500B836C" box="[487,588,1856,1880]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="17" pageNumber="18" unit="mm" value="10.4">10.4 mm</quantity>
, width
<quantity id="EA36DEDC5841FF99FD6EA51B50B2836C" box="[669,757,1856,1880]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="17" pageNumber="18" unit="mm" value="4.4">4.4 mm</quantity>
. Color yellowish red on most of body surface and legs; shiny black on two central spots of pronotal disc, part of epipleural border, and along elytral suture. Antennal club as long as funicle. Epipleural fold more swollen and enlarged medially. Metasternum with shallow longitudinal sulcus. Protibiae and protarsi narrower than in male. Genital plates elongate, subovate, with angled apex and numerous setae on distal half (
<figureCitation id="B5F56FBC5841FF99FE84A595538D83D2" box="[375,458,1998,2022]" captionStart="FIGURES 36 – 41" captionStartId="18.[151,264,1619,1642]" captionTargetBox="[189,1372,224,1565]" captionTargetId="figure@18.[151,1436,193,1598]" captionTargetPageId="18" captionText="FIGURES 36 – 41. Diagnostic structures in P. ixtacomitanus: 36) right proleg, 37) right antenna, 38) male genitalia dorsal view, 39) male genitalia lateral view, 40) spiculum gastrale, 41) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210549/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
).
</paragraph>
<caption id="79B123B15842FF9AFF64A408563082BC" httpUri="https://zenodo.org/record/210549/files/figure.png" pageId="18" pageNumber="19" targetBox="[189,1372,224,1565]" targetPageId="18">
<paragraph id="2D7173395842FF9AFF64A408563082BC" blockId="18.[151,1436,1619,1672]" pageId="18" pageNumber="19">
<emphasis id="1FBAAF2B5842FF9AFF64A408530A825C" bold="true" box="[151,333,1619,1642]" pageId="18" pageNumber="19">FIGURES 36–41</emphasis>
. Diagnostic structures in
<emphasis id="1FBAAF2B5842FF9AFDBDA40F50B9825D" box="[590,766,1620,1641]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="EACE08BA5842FF9AFDBDA40F50BF825D" box="[590,760,1620,1641]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ixtacomitanus">P. ixtacomitanus</taxonomicName>
:
</emphasis>
<emphasis id="1FBAAF2B5842FF9AFCF0A4085162825E" bold="true" box="[771,805,1619,1642]" pageId="18" pageNumber="19">36)</emphasis>
right proleg,
<emphasis id="1FBAAF2B5842FF9AFC5DA4085188825E" bold="true" box="[942,975,1619,1642]" pageId="18" pageNumber="19">37)</emphasis>
right antenna,
<emphasis id="1FBAAF2B5842FF9AFB95A40856CF825E" bold="true" box="[1126,1160,1619,1642]" pageId="18" pageNumber="19">38)</emphasis>
male genitalia dorsal view,
<emphasis id="1FBAAF2B5842FF9AFF64A42A52FF82BC" bold="true" box="[151,184,1649,1672]" pageId="18" pageNumber="19">39)</emphasis>
male genitalia lateral view,
<emphasis id="1FBAAF2B5842FF9AFE25A42A53B082BC" bold="true" box="[470,503,1649,1672]" pageId="18" pageNumber="19">40)</emphasis>
spiculum gastrale,
<emphasis id="1FBAAF2B5842FF9AFD4FA42A509982BC" bold="true" box="[700,734,1649,1672]" pageId="18" pageNumber="19">41)</emphasis>
female genital plates. Scale lines 1 mm.
</paragraph>
</caption>
<paragraph id="2D7173395842FF9AFF3AA482515A8320" blockId="18.[151,1437,1752,2029]" pageId="18" pageNumber="19">
<emphasis id="1FBAAF2B5842FF9AFF3AA482530682C5" bold="true" box="[201,321,1753,1777]" pageId="18" pageNumber="19">
<typeStatus id="F275CD9B5842FF9AFF3AA482530682C5" box="[201,321,1753,1777]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<emphasis id="1FBAAF2B5842FF9AFEA3A481531982C5" box="[336,350,1754,1777]" italics="true" pageId="18" pageNumber="19">n</emphasis>
= 20). Length: 9.0–11.0 mm, width:
<quantity id="EA36DEDC5842FF9AFD08A48351C382C4" box="[763,900,1752,1776]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="18" pageNumber="19" unit="mm" value="4.4" valueMax="4.5" valueMin="4.3">4.3–4.5 mm</quantity>
. Color in females vary from orange red to ochre yellow, but two black spots on the pronotum are constant.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25842FF9AFF3AA57B534483D9" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="2D7173395842FF9AFF3AA57B534483D9" blockId="18.[151,1437,1752,2029]" pageId="18" pageNumber="19">
<emphasis id="1FBAAF2B5842FF9AFF3AA57B5304830D" bold="true" box="[201,323,1824,1849]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Clypeus subrectangular. Pronotum with uniformly distributed deep, medium round punctures (approximate diameter
<quantity id="EA36DEDC5842FF9AFE6DA51F50538368" box="[414,532,1860,1884]" metricMagnitude="-5" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="19" unit="mm" value="0.033">0.033 mm</quantity>
). Elytral surface with striae indicated by uniform rows of deep, large punctures; elytral lateral margins without bend. Protarsomeres 1–4 shortened, but slightly wider than protarsomere 5. Protibia stout, progressively widened distally; external border with proximal denticle short and rounded; distal denticle long, oblique. Parameres short and complex, with upper part narrowed towards apex and weakly upturned distally (Table 1).
</paragraph>
</subSubSection>
<caption id="79B123B15843FF9BFF64A52E513483DC" httpUri="https://zenodo.org/record/210550/files/figure.png" pageId="19" pageNumber="20" targetBox="[214,1377,207,1832]" targetPageId="19">
<paragraph id="2D7173395843FF9BFF64A52E513483DC" blockId="19.[151,1436,1909,2024]" pageId="19" pageNumber="20">
<emphasis id="1FBAAF2B5843FF9BFF64A52E531783BF" bold="true" box="[151,336,1909,1932]" pageId="19" pageNumber="20">FIGURES 42–47</emphasis>
. Dorsal view of
<taxonomicName id="EACE08BA5843FF9BFDF6A52D50CD83B8" box="[517,650,1910,1932]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="1FBAAF2B5843FF9BFDF6A52D50CD83B8" box="[517,650,1910,1932]" italics="true" pageId="19" pageNumber="20">Pachystethus</emphasis>
</taxonomicName>
species:
<emphasis id="1FBAAF2B5843FF9BFD1DA52D510383B8" bold="true" box="[750,836,1909,1932]" pageId="19" pageNumber="20">42 a, b)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFCBFA52C51ED83B8" box="[844,938,1910,1932]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="1FBAAF2B5843FF9BFCBFA52C51ED83B8" box="[844,938,1910,1932]" italics="true" pageId="19" pageNumber="20">P. viduus</emphasis>
</taxonomicName>
males;
<emphasis id="1FBAAF2B5843FF9BFC0CA52D567283B8" bold="true" box="[1023,1077,1909,1932]" pageId="19" pageNumber="20">42 c)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFBCCA52C56DA83B8" box="[1087,1181,1910,1932]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="1FBAAF2B5843FF9BFBCCA52C56DA83B8" box="[1087,1181,1910,1932]" italics="true" pageId="19" pageNumber="20">P. viduus</emphasis>
</taxonomicName>
female;
<emphasis id="1FBAAF2B5843FF9BFB0FA52D577283B8" bold="true" box="[1276,1333,1909,1932]" pageId="19" pageNumber="20">42 d)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFACDA52C57DB83B8" box="[1342,1436,1910,1932]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="1FBAAF2B5843FF9BFACDA52C57DB83B8" box="[1342,1436,1910,1932]" italics="true" pageId="19" pageNumber="20">P. viduus</emphasis>
</taxonomicName>
female, lectotype;
<emphasis id="1FBAAF2B5843FF9BFEA0A5CF53CF839F" bold="true" box="[339,392,1940,1963]" pageId="19" pageNumber="20">43 a)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFE7DA5CE5058839E" box="[398,543,1941,1962]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nectoctenus">
<emphasis id="1FBAAF2B5843FF9BFE7DA5CE5058839E" box="[398,543,1941,1962]" italics="true" pageId="19" pageNumber="20">P. nectoctenus</emphasis>
</taxonomicName>
male;
<emphasis id="1FBAAF2B5843FF9BFD97A5CF50DC839F" bold="true" box="[612,667,1940,1963]" pageId="19" pageNumber="20">43 b)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFD52A5CE5176839E" box="[673,817,1941,1962]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nectoctenus">
<emphasis id="1FBAAF2B5843FF9BFD52A5CE5176839E" box="[673,817,1941,1962]" italics="true" pageId="19" pageNumber="20">P. nectoctenus</emphasis>
</taxonomicName>
female;
<emphasis id="1FBAAF2B5843FF9BFC78A5CF5187839F" bold="true" box="[907,960,1940,1963]" pageId="19" pageNumber="20">44 a)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFC34A5CE5633839E" box="[967,1140,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Pachyststhus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crassesculptus">
<emphasis id="1FBAAF2B5843FF9BFC34A5CE5633839E" box="[967,1140,1940,1962]" italics="true" pageId="19" pageNumber="20">P. crassesculptus</emphasis>
</taxonomicName>
male;
<emphasis id="1FBAAF2B5843FF9BFB4AA5CF56AF839F" bold="true" box="[1209,1256,1940,1963]" pageId="19" pageNumber="20">44b)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFB1CA5CE57DC839E" box="[1263,1435,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Pachyststhus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crassesculptus">
<emphasis id="1FBAAF2B5843FF9BFB1CA5CE57DC839E" box="[1263,1435,1940,1962]" italics="true" pageId="19" pageNumber="20">P. crassesculptus</emphasis>
</taxonomicName>
female;
<emphasis id="1FBAAF2B5843FF9BFF1AA5E8535283FC" bold="true" box="[233,277,1971,1992]" pageId="19" pageNumber="20">45 a</emphasis>
,
<emphasis id="1FBAAF2B5843FF9BFED1A5E9537E83FD" bold="true" box="[290,313,1970,1993]" pageId="19" pageNumber="20">b)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFECCA5EF53DC83FD" box="[319,411,1972,1993]" class="Insecta" family="Scarabaeidae" genus="Spilota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nutans">
<emphasis id="1FBAAF2B5843FF9BFECCA5EF53DC83FD" box="[319,411,1972,1993]" italics="true" pageId="19" pageNumber="20">P. nutans</emphasis>
</taxonomicName>
males;
<emphasis id="1FBAAF2B5843FF9BFE18A5E8505883FD" bold="true" box="[491,543,1970,1993]" pageId="19" pageNumber="20">45 c)</emphasis>
<emphasis id="1FBAAF2B5843FF9BFDD6A5EF503883FD" box="[549,639,1972,1993]" italics="true" pageId="19" pageNumber="20">P. nu t an s</emphasis>
female;
<emphasis id="1FBAAF2B5843FF9BFD28A5E8516C83FD" bold="true" box="[731,811,1970,1993]" pageId="19" pageNumber="20">46 a, b)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFCC2A5EF519C83FD" box="[817,987,1971,1993]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="matzapatlecus">
<emphasis id="1FBAAF2B5843FF9BFCC2A5EF519C83FD" box="[817,987,1971,1993]" italics="true" pageId="19" pageNumber="20">P. matzapatlecus</emphasis>
</taxonomicName>
males;
<emphasis id="1FBAAF2B5843FF9BFBD9A5E8561A83FD" bold="true" box="[1066,1117,1970,1993]" pageId="19" pageNumber="20">46 c)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFB97A5EF574983FD" box="[1124,1294,1971,1993]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="matzapatlecus">
<emphasis id="1FBAAF2B5843FF9BFB97A5EF574983FD" box="[1124,1294,1971,1993]" italics="true" pageId="19" pageNumber="20">P. matzapatlecus</emphasis>
</taxonomicName>
female;
<emphasis id="1FBAAF2B5843FF9BFA94A5E857DB83FD" bold="true" box="[1383,1436,1970,1993]" pageId="19" pageNumber="20">47 a)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFF64A588537983DC" box="[151,318,2003,2024]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ixtacomitanus">
<emphasis id="1FBAAF2B5843FF9BFF64A588537983DC" box="[151,318,2003,2024]" italics="true" pageId="19" pageNumber="20">P. ixtacomitanus</emphasis>
</taxonomicName>
male;
<emphasis id="1FBAAF2B5843FF9BFE70A58953FE83DC" bold="true" box="[387,441,2001,2024]" pageId="19" pageNumber="20">47 b)</emphasis>
<taxonomicName id="EACE08BA5843FF9BFE4CA588502183DC" box="[447,614,2003,2024]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ixtacomitanus">
<emphasis id="1FBAAF2B5843FF9BFE4CA588502183DC" box="[447,614,2003,2024]" italics="true" pageId="19" pageNumber="20">P. ixtacomitanus</emphasis>
</taxonomicName>
female. Scale lines 1 mm.
</paragraph>
</caption>
<caption id="79B123B15844FF9CFF64A6B05282810B" httpUri="https://zenodo.org/record/210551/files/figure.png" pageId="20" pageNumber="21" targetBox="[200,1411,240,1184]" targetPageId="20">
<paragraph id="2D7173395844FF9CFF64A6B05282810B" blockId="20.[151,1436,1259,1343]" pageId="20" pageNumber="21">
<emphasis id="1FBAAF2B5844FF9CFF64A6B053178135" bold="true" box="[151,336,1259,1282]" pageId="20" pageNumber="21">FIGURES 48–53</emphasis>
. Lateral view of
<taxonomicName id="EACE08BA5844FF9CFDF8A6B750D68136" box="[523,657,1260,1282]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="1FBAAF2B5844FF9CFDF8A6B750D68136" box="[523,657,1260,1282]" italics="true" pageId="20" pageNumber="21">Pachystethus</emphasis>
</taxonomicName>
species:
<emphasis id="1FBAAF2B5844FF9CFD07A6B051528136" bold="true" box="[756,789,1259,1282]" pageId="20" pageNumber="21">48)</emphasis>
<taxonomicName id="EACE08BA5844FF9CFCECA6B651398136" box="[799,894,1260,1282]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="1FBAAF2B5844FF9CFCECA6B651398136" box="[799,894,1260,1282]" italics="true" pageId="20" pageNumber="21">P. viduus</emphasis>
</taxonomicName>
<emphasis id="1FBAAF2B5844FF9CFC74A6B051DA8136" bold="true" box="[903,925,1259,1282]" pageId="20" pageNumber="21">a)</emphasis>
male,
<emphasis id="1FBAAF2B5844FF9CFC14A6B051B98136" bold="true" box="[999,1022,1259,1282]" pageId="20" pageNumber="21">b)</emphasis>
female;
<emphasis id="1FBAAF2B5844FF9CFBAEA6B056398136" bold="true" box="[1117,1150,1259,1282]" pageId="20" pageNumber="21">49)</emphasis>
<taxonomicName id="EACE08BA5844FF9CFB7BA6B6575C8136" box="[1160,1307,1261,1282]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nectoctenus">
<emphasis id="1FBAAF2B5844FF9CFB7BA6B6575C8136" box="[1160,1307,1261,1282]" italics="true" pageId="20" pageNumber="21">P. nectoctenus</emphasis>
</taxonomicName>
<emphasis id="1FBAAF2B5844FF9CFAD6A6B0577C8136" bold="true" box="[1317,1339,1259,1282]" pageId="20" pageNumber="21">a)</emphasis>
male,
<emphasis id="1FBAAF2B5844FF9CFA76A6B057DB8136" bold="true" box="[1413,1436,1259,1282]" pageId="20" pageNumber="21">b)</emphasis>
female;
<emphasis id="1FBAAF2B5844FF9CFF18A751534B8115" bold="true" box="[235,268,1290,1313]" pageId="20" pageNumber="21">50)</emphasis>
<taxonomicName id="EACE08BA5844FF9CFEE6A75053838114" box="[277,452,1290,1312]" class="Insecta" family="Scarabaeidae" genus="Pachyststhus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="crassesculptus">
<emphasis id="1FBAAF2B5844FF9CFEE6A75053838114" box="[277,452,1290,1312]" italics="true" pageId="20" pageNumber="21">P. crassesculptus</emphasis>
</taxonomicName>
male;
<emphasis id="1FBAAF2B5844FF9CFDFFA75150698115" bold="true" box="[524,558,1290,1313]" pageId="20" pageNumber="21">51)</emphasis>
<emphasis id="1FBAAF2B5844FF9CFDC4A75050D58114" box="[567,658,1291,1312]" italics="true" pageId="20" pageNumber="21">P. n u t a n s</emphasis>
male;
<emphasis id="1FBAAF2B5844FF9CFD2DA75150B88115" bold="true" box="[734,767,1290,1313]" pageId="20" pageNumber="21">52)</emphasis>
<taxonomicName id="EACE08BA5844FF9CFCFBA75051F38114" box="[776,948,1290,1312]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="matzapatlecus">
<emphasis id="1FBAAF2B5844FF9CFCFBA75051F38114" box="[776,948,1290,1312]" italics="true" pageId="20" pageNumber="21">P. matzapatlecus</emphasis>
</taxonomicName>
male;
<emphasis id="1FBAAF2B5844FF9CFC0FA75156598115" bold="true" box="[1020,1054,1290,1313]" pageId="20" pageNumber="21">53)</emphasis>
<taxonomicName id="EACE08BA5844FF9CFBD4A75056978114" box="[1063,1232,1291,1312]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ixtacomitanus">
<emphasis id="1FBAAF2B5844FF9CFBD4A75056978114" box="[1063,1232,1291,1312]" italics="true" pageId="20" pageNumber="21">P. ixtacomitanus</emphasis>
</taxonomicName>
male. Scale lines 1 mm.
</paragraph>
</caption>
<subSubSection id="65D420B25844FF9CFF3AA737514C8193" pageId="20" pageNumber="21" type="etymology">
<paragraph id="2D7173395844FF9CFF3AA737514C8193" blockId="20.[151,1436,1387,1699]" pageId="20" pageNumber="21">
<emphasis id="1FBAAF2B5844FF9CFF3AA737530881B0" bold="true" box="[201,335,1388,1412]" pageId="20" pageNumber="21">Etymology.</emphasis>
From the Nahuatl indigenous village “
<emphasis id="1FBAAF2B5844FF9CFCE7A73751E681B0" box="[788,929,1388,1412]" italics="true" pageId="20" pageNumber="21">Ixtacomitlan</emphasis>
” in the northern slopes of state of Chiapas, where the first specimens of this species were collected.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25844FF9CFF3AA7E851E58203" pageId="20" pageNumber="21" type="distribution">
<paragraph id="2D7173395844FF9CFF3AA7E851E58203" blockId="20.[151,1436,1387,1699]" pageId="20" pageNumber="21">
<emphasis id="1FBAAF2B5844FF9CFF3AA7E8509081F8" bold="true" box="[201,727,1459,1484]" pageId="20" pageNumber="21">Distribution, phenology, and natural history</emphasis>
. This species is distributed in the humid slopes of Sierra Norte de Chiapas in
<collectingCountry id="55D933A95844FF9CFECBA78C53C981DB" box="[312,398,1495,1519]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
(
<figureCitation id="B5F56FBC5844FF9CFE6EA78C53A881DB" box="[413,495,1495,1519]" captionStart="FIGURE 54" captionStartId="21.[151,250,1234,1257]" captionTargetBox="[157,1436,382,1193]" captionTargetId="figure@21.[157,1442,373,1193]" captionTargetPageId="21" captionText="FIGURE 54. Distribution of species of Pachystethus." httpUri="https://zenodo.org/record/210552/files/figure.png" pageId="20" pageNumber="21">Fig. 54</figureCitation>
) between
<quantity id="EA36DEDC5844FF9CFD90A78C50BF81C4" box="[611,760,1495,1520]" metricMagnitude="3" metricUnit="m" metricValue="1.005" metricValueMax="1.86" metricValueMin="0.15" pageId="20" pageNumber="21" unit="m" value="1005.0" valueMax="1860.0" valueMin="150.0">150–1,860 m</quantity>
in elevation, where cloud forests and pine-oak forests overlaps with tropical rain forest and coffee plantations. Adults are active during June through December. Plant hosts for adults are unknown, and the immature stages remain undescribed.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25844FF9CFF3AA41F57608297" pageId="20" pageNumber="21" type="discussion">
<paragraph id="2D7173395844FF9CFF3AA41F57608297" blockId="20.[151,1436,1387,1699]" pageId="20" pageNumber="21">
<emphasis id="1FBAAF2B5844FF9CFF3AA41F53718268" bold="true" box="[201,310,1604,1628]" pageId="20" pageNumber="21">Remarks</emphasis>
.
<taxonomicName id="EACE08BA5844FF9CFEBFA41F50CA8268" box="[332,653,1604,1628]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ixtacomitanus">
<emphasis id="1FBAAF2B5844FF9CFEBFA41F50CA8268" box="[332,653,1604,1628]" italics="true" pageId="20" pageNumber="21">Pachystethus ixtacomitanus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="EACE08BA5844FF9CFCB1A41E56408268" box="[834,1031,1604,1628]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="matzapatlecus">
<emphasis id="1FBAAF2B5844FF9CFCB1A41E56408268" box="[834,1031,1604,1628]" italics="true" pageId="20" pageNumber="21">P. matzapatlecus</emphasis>
</taxonomicName>
and
<taxonomicName id="EACE08BA5844FF9CFBBCA41E56B08268" box="[1103,1271,1605,1628]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nectoctenus">
<emphasis id="1FBAAF2B5844FF9CFBBCA41E56B08268" box="[1103,1271,1605,1628]" italics="true" pageId="20" pageNumber="21">P. nectoctenus</emphasis>
</taxonomicName>
, but may be distinguished from the former because the elytra with the posterior calli prominent, and from the latter because the elytra are clearly punctate instead of nearly smooth. The shape of parameres is also diagnostic (Table 1).
</paragraph>
</subSubSection>
</treatment>
</document>
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