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<document ID-DOI="http://dx.doi.org/10.3897/dez.65.26693" ID-GBIF-Dataset="e4872547-7d79-46ac-bcbc-07d1f39705c5" ID-GBIF-Taxon="182223627" ID-Pensoft-Pub="1860-1324-2-161" ID-Pensoft-UUID="22E30A28F3F95FB0848C3A58737B1CE9" ID-Zenodo-Dep="1343164" ID-ZooBank="196D0BD954A1481181D4B714B6B2BEEE" ModsDocID="1860-1324-65-2-161" checkinTime="1621372437338" checkinUser="pensoft" docAuthor="Massa, Bruno, Heller, Klaus-Gerhard, Warchalowska-Śliwa, Elzbieta &amp; Moulin, Nicolas" docDate="2018" docId="A01DC10D0EAD0ABEDCA67503943B8041" docLanguage="en" docName="DeutEntomolZeit 65(2): 161-175" docOrigin="Deutsche Entomologische Zeitschrift 65 (2)" docPubDate="2018-08-02" docSource="http://dx.doi.org/10.3897/dez.65.26693" docTitle="Morgenia spathulifera Griffini 1908" docType="treatment" docVersion="3" id="22E30A28F3F95FB0848C3A58737B1CE9" lastPageNumber="163" masterDocId="22E30A28F3F95FB0848C3A58737B1CE9" masterDocTitle="The tropical African genus Morgenia (Orthoptera, Tettigoniidae, Phaneropterinae) with emphasis on the spur at the mid tibia" masterLastPageNumber="175" masterPageNumber="161" pageId="2" pageNumber="163" updateTime="1632015767259" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The tropical African genus Morgenia (Orthoptera, Tettigoniidae, Phaneropterinae) with emphasis on the spur at the mid tibia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Massa, Bruno</mods:namePart>
<mods:affiliation>Department of Agricultural, Food and Forest Sciences, University of Palermo, Viale Scienze Bd. 5 A, 90128 Palermo, Italy</mods:affiliation>
<mods:nameIdentifier type="email">bruno.massa@unipa.it</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heller, Klaus-Gerhard</mods:namePart>
<mods:affiliation>Grillenstieg 18, 39120 Magdeburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Warchalowska-Śliwa, Elzbieta</mods:namePart>
<mods:affiliation>Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31515 Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moulin, Nicolas</mods:namePart>
<mods:affiliation>Route de l'Ecole, 82, Hameau de Saveaumare, 76680 Monterolier, France</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-08-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>65</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:extent unit="page">
<mods:start>161</mods:start>
<mods:end>175</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.65.26693</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.65.26693</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2-161</mods:identifier>
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<treatment ID-GBIF-Taxon="182223627" LSID="urn:lsid:plazi:treatment:A01DC10D0EAD0ABEDCA67503943B8041" httpUri="http://treatment.plazi.org/id/A01DC10D0EAD0ABEDCA67503943B8041" lastPageNumber="163" pageId="2" pageNumber="163">
<subSubSection pageId="2" pageNumber="163" type="nomenclature">
<paragraph pageId="2" pageNumber="163">
<taxonomicName authority="Griffini, 1908" authorityName="Griffini" authorityYear="1908" class="Insecta" family="Tettigoniidae" genus="Morgenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morgenia spathulifera" order="Orthoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="spathulifera">
<pageBreakToken pageId="2" pageNumber="163" start="start">Morgenia</pageBreakToken>
spathulifera Griffini, 1908
</taxonomicName>
<figureCitation captionStart="Figures 58" captionStartId="F2" captionText="Figures 5 - 8. Habitus of the male of Morgenia spathulifera (5), M. angustipinnata sp. n. (6), M. melica (7) and M. modulata (8) (left side)." figureDoi="10.3897/dez.65.26693.figures5-8" httpUri="https://binary.pensoft.net/fig/220909" pageId="2" pageNumber="163">Figs 5</figureCitation>
<figureCitation captionStart="Figures 914" captionStartId="F3" captionText="Figures 9 - 14. Stridulatory area of left tegmen and mirror of right tegmen of the male of Morgenia hamuligera (9), M. rubricornis (10), M. spathulifera (11), M. lehmannorum sp. n. (12); habitus of M. hamuligera after Brunner von Wattenwyl (1878) (13) and of M. rubricornis after Sjoestedt (1913) (14). Arrow shows the particular net of small cells in the costal area of tegmina of M. spathulifera, present also in M. lehmannorum sp. n." figureDoi="10.3897/dez.65.26693.figures9-14" httpUri="https://binary.pensoft.net/fig/220910" pageId="2" pageNumber="163">, 11</figureCitation>
<figureCitation captionStart="Figures 2326" captionStartId="F6" captionText="Figures 23 - 26. Stridulatory file of the left tegmen in the male of Morgenia spathulifera (23), M. angustipinnata sp. n. (24), M. melica (25) and M. modulata (26)." figureDoi="10.3897/dez.65.26693.figures23-26" httpUri="https://binary.pensoft.net/fig/220913" pageId="2" pageNumber="163">, 23</figureCitation>
<figureCitation captionStart="Figures 2734" captionStartId="F7" captionText="Figures 27 - 34. Mid tibia showing the inner spur in the male of Morgenia hamuligera (27), M. rubricornis (28), M. plurimaculata sp. n. (29), M. lehmannorum sp. n. (30), M. angustipinnata sp. n. (31), M. spathulifera (32), M. melica (33) and M. modulata (34)." figureDoi="10.3897/dez.65.26693.figures27-34" httpUri="https://binary.pensoft.net/fig/220914" pageId="2" pageNumber="163">, 32</figureCitation>
<figureCitation captionStart="Figures 3944" captionStartId="F9" captionText="Figures 39 - 44. Cerci and subgenital plate of male of Morgenia lehmannorum sp. n. (39 lateral, 40 ventral), M. plurimaculata sp. n. (41 lateral, 42 ventral) and M. spathulifera (43 lateral, 44 ventral)." figureDoi="10.3897/dez.65.26693.figures39-44" httpUri="https://binary.pensoft.net/fig/220916" pageId="2" pageNumber="163">, 43</figureCitation>
<figureCitation captionStart="Figures 3944" captionStartId="F9" captionText="Figures 39 - 44. Cerci and subgenital plate of male of Morgenia lehmannorum sp. n. (39 lateral, 40 ventral), M. plurimaculata sp. n. (41 lateral, 42 ventral) and M. spathulifera (43 lateral, 44 ventral)." figureDoi="10.3897/dez.65.26693.figures39-44" httpUri="https://binary.pensoft.net/fig/220916" pageId="2" pageNumber="163">, 44</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="163" type="reference_group">
<paragraph pageId="2" pageNumber="163">
<taxonomicName authorityName="Griffini" authorityYear="1908" class="Insecta" family="Tettigoniidae" genus="Morgenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morgenia spathulifera" order="Orthoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="spathulifera">Morgenia spathulifera</taxonomicName>
Griffini, 1908. Mem. Soc. entom. Belgique, Bruxelles 15: 209.
</paragraph>
<paragraph pageId="2" pageNumber="163">
<taxonomicName authorityName="Griffini" authorityYear="1908" class="Insecta" family="Tettigoniidae" genus="Morgenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morgenia spathulifera" order="Orthoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="spathulifera">Morgenia spathulifera</taxonomicName>
Type locality: Bussanga (Democratic Republic of Congo) (RBINS, Bruxelles).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="163" type="materials_examined">
<paragraph pageId="2" pageNumber="163">Material examined.</paragraph>
<paragraph pageId="2" pageNumber="163">
<materialsCitation collectingDate="1905-11-14" collectingDateMax="1939-01-09" collectingDateMin="1905-11-14" collectorName="Ngowa, J. Mertens" country="Democratic Republic of the Congo" location="Bussanga" specimenCount="2" specimenCount-male="2" typeStatus="syntype">
<emphasis bold="true" pageId="2" pageNumber="163">
<collectingCountry name="Democratic Republic of the Congo">Democratic Republic of the Congo</collectingCountry>
</emphasis>
,
<location LSID="urn:lsid:plazi:treatment:A01DC10D0EAD0ABEDCA67503943B8041:10CBEC11E60E798EE14EA7883494624B" country="Democratic Republic of the Congo" name="Bussanga">Bussanga</location>
<collectingDate value="1905-11-14">14.XI.1905</collectingDate>
(
<specimenCount type="male">1♂</specimenCount>
<typeStatus>syntype</typeStatus>
);
<collectorName>Ngowa</collectorName>
<collectingDate value="1939-01-09">9.I.1939</collectingDate>
(
<specimenCount type="male">1♂</specimenCount>
),
<collectorName>J. Mertens</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1983-01-31" collectingDateMax="1983-02-08" collectingDateMin="1983-01-31" collectorName="Mukonje Farm, R, J. van Stalle" country="Cameroon" location="Mt Koupe" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="syntype">
<emphasis bold="true" pageId="2" pageNumber="163">
<collectingCountry name="Cameroon">Cameroon</collectingCountry>
</emphasis>
,
<collectorName>Mukonje Farm, R</collectorName>
. Rohde (
<specimenCount type="female">1♀</specimenCount>
<typeStatus>syntype</typeStatus>
);
<location LSID="urn:lsid:plazi:treatment:A01DC10D0EAD0ABEDCA67503943B8041:7ECF057B919A065252AD27E8D15D74BA" country="Cameroon" name="Mt Koupe">
Mt
<normalizedToken originalValue="Koupé">Koupe</normalizedToken>
</location>
<collectingDate value="1983-01-31" valueMax="1983-02-08" valueMin="1983-01-31">31.I-8.II.1983</collectingDate>
(
<specimenCount type="male">1♂</specimenCount>
),
<collectorName>J. van Stalle</collectorName>
(RBINS)
</materialsCitation>
;
<materialsCitation collectingDate="2017-03-13" collectingDateMax="2017-04-04" collectingDateMin="2017-03-13" collectorName="B. Massa, P. Moretto" country="Ivory Coast" county="Res. Station" location="Tai Nat. Park" municipality="Tai Nat. Park" specimenCount="3" specimenCount-male="3" typeStatus="syntype">
<emphasis bold="true" pageId="2" pageNumber="163">
<collectingCountry name="Ivory Coast">Ivory Coast</collectingCountry>
</emphasis>
,
<location LSID="urn:lsid:plazi:treatment:A01DC10D0EAD0ABEDCA67503943B8041:F8AD827A5DFCABEB3EDCC212117330C5" country="Ivory Coast" county="Res. Station" municipality="Tai Nat. Park" name="Tai Nat. Park">
<normalizedToken originalValue="Taï">Tai</normalizedToken>
Nat. Park
</location>
,
<location LSID="urn:lsid:plazi:treatment:A01DC10D0EAD0ABEDCA67503943B8041:ACBC153802DE295C9286EE6F6AA52B48" country="Ivory Coast" county="Res. Station" municipality="Tai Nat. Park" name="Res. Station">Res. Station</location>
<collectingDate value="2017-03-13">13.III.2017</collectingDate>
(
<specimenCount type="male">1♂</specimenCount>
) (light),
<collectorName>B. Massa</collectorName>
;
<collectingMunicipality>
<normalizedToken originalValue="Taï">Tai</normalizedToken>
Nat. Park
</collectingMunicipality>
,
<collectingCounty>Res. Station</collectingCounty>
<collectingDate value="2017-03-22" valueMax="2017-04-04" valueMin="2017-03-22">22.III-4.IV.2017</collectingDate>
(
<specimenCount type="male">2♂</specimenCount>
) (light),
<collectorName>P. Moretto</collectorName>
(BMPC)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="163" type="distribution">
<paragraph pageId="2" pageNumber="163">Distribution.</paragraph>
<paragraph pageId="2" pageNumber="163">
<taxonomicName class="Insecta" genus="M." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="M. spathulifera" order="Saltatoria" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="spathulifera">
<emphasis italics="true" pageId="2" pageNumber="163">M. spathulifera</emphasis>
</taxonomicName>
is presently known from Cameroon, Democratic Republic of the Congo and Ivory Coast (
<bibRefCitation author="Griffini, A" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="10" pageNumber="171" publicationUrl="https://biodiversitylibrary.org/page/11121989" refId="B6" refString="Griffini, A, 1908. Phasgonuridae africane del R. Museo di Storia Naturale in Bruxelles. 6. Phaneropteridae pars 2a (reliquae species omnes). Memoires de la Societe entomologique Belgique 15: 201-226. https://biodiversitylibrary.org/page/11121989" title="Phasgonuridae africane del R. Museo di Storia Naturale in Bruxelles. 6. Phaneropteridae pars 2 a (reliquae species omnes). Memoires de la Societe entomologique Belgique 15: 201 - 226." url="https://biodiversitylibrary.org/page/11121989" year="1908">Griffini 1908</bibRefCitation>
, Massa 2017). We presume that it covers central and western tropical forests of Africa.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="163" type="remarks">
<paragraph pageId="2" pageNumber="163">Remarks.</paragraph>
<paragraph pageId="2" pageNumber="163">
<taxonomicName class="Insecta" genus="M." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="M. spathulifera" order="Saltatoria" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="spathulifera">
<emphasis italics="true" pageId="2" pageNumber="163">M. spathulifera</emphasis>
</taxonomicName>
may be easily separated from other known species by the ratio length/width of the tegmina (6.4-6.8). It is also characterised by the presence of small grey dots on the tegmina and costal area of the tegmina with a net of small cells (Figs
<figureCitation captionStart="Figures 58" captionStartId="F2" captionText="Figures 5 - 8. Habitus of the male of Morgenia spathulifera (5), M. angustipinnata sp. n. (6), M. melica (7) and M. modulata (8) (left side)." figureDoi="10.3897/dez.65.26693.figures5-8" httpUri="https://binary.pensoft.net/fig/220909" pageId="2" pageNumber="163">5</figureCitation>
,
<figureCitation captionStart="Figures 914" captionStartId="F3" captionText="Figures 9 - 14. Stridulatory area of left tegmen and mirror of right tegmen of the male of Morgenia hamuligera (9), M. rubricornis (10), M. spathulifera (11), M. lehmannorum sp. n. (12); habitus of M. hamuligera after Brunner von Wattenwyl (1878) (13) and of M. rubricornis after Sjoestedt (1913) (14). Arrow shows the particular net of small cells in the costal area of tegmina of M. spathulifera, present also in M. lehmannorum sp. n." figureDoi="10.3897/dez.65.26693.figures9-14" httpUri="https://binary.pensoft.net/fig/220910" pageId="2" pageNumber="163">11</figureCitation>
). The stridulatory area of the left tegmen is protruding below its posterior margin and it is composed of the stridulatory file and has a raised parallel posterior bulge (Fig.
<figureCitation captionStart="Figures 914" captionStartId="F3" captionText="Figures 9 - 14. Stridulatory area of left tegmen and mirror of right tegmen of the male of Morgenia hamuligera (9), M. rubricornis (10), M. spathulifera (11), M. lehmannorum sp. n. (12); habitus of M. hamuligera after Brunner von Wattenwyl (1878) (13) and of M. rubricornis after Sjoestedt (1913) (14). Arrow shows the particular net of small cells in the costal area of tegmina of M. spathulifera, present also in M. lehmannorum sp. n." figureDoi="10.3897/dez.65.26693.figures9-14" httpUri="https://binary.pensoft.net/fig/220910" pageId="2" pageNumber="163">11</figureCitation>
). The mirror of the right tegmen is smaller than in
<taxonomicName class="Insecta" genus="M." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="M. hamuligera" order="Saltatoria" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="hamuligera">
<emphasis italics="true" pageId="2" pageNumber="163">M. hamuligera</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 914" captionStartId="F3" captionText="Figures 9 - 14. Stridulatory area of left tegmen and mirror of right tegmen of the male of Morgenia hamuligera (9), M. rubricornis (10), M. spathulifera (11), M. lehmannorum sp. n. (12); habitus of M. hamuligera after Brunner von Wattenwyl (1878) (13) and of M. rubricornis after Sjoestedt (1913) (14). Arrow shows the particular net of small cells in the costal area of tegmina of M. spathulifera, present also in M. lehmannorum sp. n." figureDoi="10.3897/dez.65.26693.figures9-14" httpUri="https://binary.pensoft.net/fig/220910" pageId="2" pageNumber="163">11</figureCitation>
). The stridulatory file is 1.2 mm long, arched and composed of ca. 60 densely spaced teeth (Fig.
<figureCitation captionStart="Figures 2326" captionStartId="F6" captionText="Figures 23 - 26. Stridulatory file of the left tegmen in the male of Morgenia spathulifera (23), M. angustipinnata sp. n. (24), M. melica (25) and M. modulata (26)." figureDoi="10.3897/dez.65.26693.figures23-26" httpUri="https://binary.pensoft.net/fig/220913" pageId="2" pageNumber="163">23</figureCitation>
). The inner ventral spur of mid tibiae is more apical, short, up- and incurved and does not exceed the base of the first tarsal article; its length is about 2.5-3.0% the length of tibia. On the outer ventral margin of the mid tibiae, three close short spines are present (Fig.
<figureCitation captionStart="Figures 2734" captionStartId="F7" captionText="Figures 27 - 34. Mid tibia showing the inner spur in the male of Morgenia hamuligera (27), M. rubricornis (28), M. plurimaculata sp. n. (29), M. lehmannorum sp. n. (30), M. angustipinnata sp. n. (31), M. spathulifera (32), M. melica (33) and M. modulata (34)." figureDoi="10.3897/dez.65.26693.figures27-34" httpUri="https://binary.pensoft.net/fig/220914" pageId="2" pageNumber="163">32</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="1343190" doi="10.3897/dez.65.26693.figures5-8" httpUri="https://binary.pensoft.net/fig/220909" pageId="2" pageNumber="163" start="Figures 58" startId="F2">
<paragraph pageId="2" pageNumber="163">
<emphasis bold="true" pageId="2" pageNumber="163">Figures 5-8.</emphasis>
Habitus of the male of
<taxonomicName authorityName="Griffini" authorityYear="1908" class="Insecta" family="Tettigoniidae" genus="Morgenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morgenia spathulifera" order="Orthoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="spathulifera">
<emphasis italics="true" pageId="2" pageNumber="163">Morgenia spathulifera</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="2" pageNumber="163">5</emphasis>
),
<taxonomicName class="Insecta" genus="M." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="M. angustipinnata" order="Saltatoria" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="angustipinnata">
<emphasis italics="true" pageId="2" pageNumber="163">M. angustipinnata</emphasis>
</taxonomicName>
sp. n. (
<emphasis bold="true" pageId="2" pageNumber="163">6</emphasis>
),
<taxonomicName class="Insecta" genus="M." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="M. melica" order="Saltatoria" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="melica">
<emphasis italics="true" pageId="2" pageNumber="163">M. melica</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="2" pageNumber="163">7</emphasis>
) and
<taxonomicName class="Insecta" genus="M." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="M. modulata" order="Saltatoria" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="modulata">
<emphasis italics="true" pageId="2" pageNumber="163">M. modulata</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="2" pageNumber="163">8</emphasis>
) (left side).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>