454 lines
41 KiB
XML
454 lines
41 KiB
XML
<document ID-DOI="10.11646/zootaxa.5270.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7850665" ID-ZooBank="8544401E-7B8D-4552-AD15-BFECAF29710A" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.tables_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1681979080773" checkinUser="plazi" docAuthor="Cock, Matthew J. W., Laguerre, Michel, Buddie, Alan G., Cafa, Giovanni, Alston-Smith, Scott, Morrall, John & Gosula, Venkata Siva" docDate="2023" docId="9B35878EBC5FE268FF65FBA5FE30FE70" docLanguage="en" docName="zootaxa.5270.2.4.pdf" docOrigin="Zootaxa 5270 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5270.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calodesma pseudocollaris Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula 2023, new species" docType="treatment" docVersion="1" lastPageNumber="256" masterDocId="670CFFF6BC48E271FFF2FF9BFFE4FF85" masterDocTitle="Using DNA barcodes to test the association of sexes and morphs in Calodesma spp (Lepidoptera, Erebidae, Arctiinae, Arctiini, Pericopina) of Trinidad, West Indies with an overview of the genus, taxonomic changes and a new species" masterLastPageNumber="261" masterPageNumber="231" pageNumber="254" updateTime="1682450417206" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Using DNA barcodes to test the association of sexes and morphs in Calodesma spp (Lepidoptera, Erebidae, Arctiinae, Arctiini, Pericopina) of Trinidad, West Indies with an overview of the genus, taxonomic changes and a new species</mods:title>
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</mods:titleInfo>
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||
<mods:name type="personal">
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||
<mods:role>
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||
<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Cock, Matthew J. W.</mods:namePart>
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<mods:affiliation>CABI, Bakeham Lane, Egham, Surrey TW 20 9 TY, UK.</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Laguerre, Michel</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-6524-7069</mods:nameIdentifier>
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<mods:affiliation>31, rue de la Haute-Lande, 33850 Léognan, France. mlaguerre @ wanadoo. fr; https: // orcid. org / 0000 - 0002 - 6524 - 7069</mods:affiliation>
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<mods:nameIdentifier type="email">mlaguerre@wanadoo.fr</mods:nameIdentifier>
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</mods:name>
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||
<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Buddie, Alan G.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-7677-9300</mods:nameIdentifier>
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<mods:affiliation>CABI, Bakeham Lane, Egham, Surrey TW 20 9 TY, UK. & a. buddie @ cabi. org; https: // orcid. org / 0000 - 0002 - 7677 - 9300</mods:affiliation>
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<mods:nameIdentifier type="email">a.buddie@cabi.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cafa, Giovanni</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-0819-2126</mods:nameIdentifier>
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<mods:affiliation>CABI, Bakeham Lane, Egham, Surrey TW 20 9 TY, UK. & Present address: Eurofins County Pathology Ltd., 8 Huxley Road, Guildford, Surrey GU 2 7 RE, UK. giovanni. cafa @ biomnis. co. uk; https: // orcid. org / 0000 - 0003 - 0819 - 2126</mods:affiliation>
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<mods:nameIdentifier type="email">giovanni.cafa@biomnis.co.uk</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Alston-Smith, Scott</mods:namePart>
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<mods:affiliation>13 Hopeton Crown Trace, Black Rock, Tobago, Trinidad & Tobago. s _ alstonsmith @ yahoo. com</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Morrall, John</mods:namePart>
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<mods:affiliation>Upper Schoolfields, Blithbury Road, Hamstall Ridware, WS 15 3 RR, UK. ridware @ outlook. com</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gosula, Venkata Siva</mods:namePart>
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<mods:affiliation>Bldg 56 FFE, French Village Apts., EWMSC, Mt. Hope, Trinidad & Tobago. vgosula 2021 @ gmail. com</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>Zootaxa</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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||
<mods:date>2023</mods:date>
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||
<mods:detail type="pubDate">
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||
<mods:number>2023-04-20</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
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||
<mods:number>5270</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>2</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>231</mods:start>
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||
<mods:end>261</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5270.2.4</mods:url>
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||
</mods:location>
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||
<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.11646/zootaxa.5270.2.4</mods:identifier>
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||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">7850665</mods:identifier>
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||
<mods:identifier type="ZooBank">8544401E-7B8D-4552-AD15-BFECAF29710A</mods:identifier>
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||
</mods:mods>
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||
<treatment LSID="urn:lsid:plazi:treatment:9B35878EBC5FE268FF65FBA5FE30FE70" httpUri="http://treatment.plazi.org/id/9B35878EBC5FE268FF65FBA5FE30FE70" lastPageId="25" lastPageNumber="256" pageId="23" pageNumber="254">
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||
<subSubSection pageId="23" pageNumber="254" type="nomenclature">
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||
<paragraph blockId="23.[151,967,1086,1221]" box="[151,706,1086,1113]" pageId="23" pageNumber="254">
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||
<heading bold="true" box="[151,706,1086,1113]" fontSize="11" level="1" pageId="23" pageNumber="254" reason="1">
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||
<emphasis bold="true" box="[151,706,1086,1113]" pageId="23" pageNumber="254">
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<taxonomicName authority="Cock" authorityName="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula" authorityYear="2023" box="[151,541,1086,1113]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="pseudocollaris" status="new species">
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<emphasis bold="true" box="[151,466,1086,1113]" italics="true" pageId="23" pageNumber="254">Calodesma pseudocollaris</emphasis>
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||
Cock
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||
</taxonomicName>
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,
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<taxonomicNameLabel box="[554,700,1086,1113]" pageId="23" pageNumber="254" rank="species">new species</taxonomicNameLabel>
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||
,
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph blockId="23.[151,967,1086,1221]" box="[151,400,1123,1149]" pageId="23" pageNumber="254">BIN: BOLD:AEI2170</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="23" pageNumber="254" type="description">
|
||
<paragraph blockId="23.[151,967,1086,1221]" box="[151,967,1158,1185]" pageId="23" pageNumber="254">
|
||
<uri box="[151,967,1158,1185]" pageId="23" pageNumber="254">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid box="[424,967,1158,1185]" pageId="23" pageNumber="254">464505E0-2DFE-4146-9C03-0D74A6A5E8AE</uuid>
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||
</uri>
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||
</paragraph>
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||
<paragraph blockId="23.[151,967,1086,1221]" box="[151,330,1195,1221]" pageId="23" pageNumber="254">
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||
(
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<figureCitation box="[159,239,1195,1221]" captionStart="FIGURE 9" captionStartId="10.[151,250,676,700]" captionTargetBox="[207,1379,200,641]" captionTargetId="figure-18@10.[189,1398,181,653]" captionTargetPageId="10" captionText="FIGURE 9. Type material Calodesma pseudocollaris new species Trinidad, W.I., Cat’s Hill, at eupatorium flowers, 24.ix.2017, J. Morrall; dorsal view above, ventral view below. A, holotype, ♁ DNA 137. B, paratype ♁, DNA 135, dissection 1001. C, allotype ♀, DNA 143. Life size." figureDoi="http://doi.org/10.5281/zenodo.7850685" httpUri="https://zenodo.org/record/7850685/files/figure.png" pageId="23" pageNumber="254">Figs. 9</figureCitation>
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,
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<figureCitation box="[251,281,1195,1221]" captionStart="FIGURE 10" captionStartId="11.[151,250,1102,1126]" captionTargetBox="[194,1394,179,1071]" captionTargetId="figure-18@11.[194,1394,179,1076]" captionTargetPageId="11" captionText="FIGURE 10. Male genitalia of Calodesma spp. from Trinidad, W.I. Top row, C. pseudocollaris new species, paratype, Trinidad, W.I., Cat’s Hill, eupatorium flowers, 24.ix.2017, John Morrall [MJWC, DNA 135, dissection 1001]. Middle row, C. militta BOLD:AAK1660 collaris morph, Parrylands, eupatorium flowers, x.2017, S. Alston-Smith [MJWC, DNA 162, dissection 1003]. Bottom row, C. militta BOLD:AAK1660 dioptis morph, data as middle row [MJWC, DNA 161, dissection 1004]. Each row from left to right: dorsal view; ventral view; lateral view from left; aedeagus lateral view from left; aedeagus ventral or ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.7850687" httpUri="https://zenodo.org/record/7850687/files/figure.png" pageId="23" pageNumber="254">10</figureCitation>
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,
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<figureCitation box="[293,322,1195,1221]" captionStart="FIGURE 16" captionStartId="24.[151,250,1878,1902]" captionTargetBox="[151,1436,1098,1852]" captionTargetId="figure-495@24.[151,1436,1098,1852]" captionTargetPageId="24" captionText="FIGURE 16 Views of the head and anterior thorax of Calodesma pseudocollaris new species. A–C paratype male. D–F allotype female." figureDoi="http://doi.org/10.5281/zenodo.7850707" httpUri="https://zenodo.org/record/7850707/files/figure.png" pageId="23" pageNumber="254">16</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="23" pageNumber="254" type="materials_examined">
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||
<paragraph blockId="23.[151,1437,1266,2013]" pageId="23" pageNumber="254">
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||
<emphasis bold="true" box="[151,324,1266,1293]" pageId="23" pageNumber="254">Type material.</emphasis>
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||
<materialsCitation collectingDate="2017-09-24" collectorName="John Morrall & M. J. W. Cock" country="Trinidad and Tobago" location="Cat's Hill" pageId="23" pageNumber="254" specimenCount="1" typeStatus="holotype">
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||
<typeStatus box="[330,434,1267,1293]" pageId="23" pageNumber="254">Holotype</typeStatus>
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||
♁:
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<collectingCountry box="[475,573,1267,1293]" name="Trinidad and Tobago" pageId="23" pageNumber="254">Trinidad</collectingCountry>
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||
, W.I.,
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||
<location LSID="urn:lsid:plazi:treatment:9B35878EBC5FE268FF65FBA5FE30FE70:16436043BC5FE266FD71FB68FD15FA88" box="[643,753,1267,1293]" country="Trinidad and Tobago" name="Cat's Hill" pageId="23" pageNumber="254">Cat’s Hill</location>
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||
, eupatorium flowers,
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||
<date box="[994,1117,1267,1293]" pageId="23" pageNumber="254" value="2017-09-24">
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||
<collectingDate box="[994,1117,1267,1293]" pageId="23" pageNumber="254" value="2017-09-24">24.ix.2017</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[1127,1272,1267,1293]" pageId="23" pageNumber="254">John Morrall</collectorName>
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||
/ DNA sample 135,
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<collectorName box="[208,356,1303,1329]" pageId="23" pageNumber="254">M.J.W. Cock</collectorName>
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||
/
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<collectorName box="[381,529,1303,1329]" pageId="23" pageNumber="254">M.J.W. Cock</collectorName>
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dissection 1001
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||
</materialsCitation>
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||
/
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||
<materialsCitation collectionCode="NHMUK, MJWC" country="Trinidad and Tobago" pageId="23" pageNumber="254" specimenCount="1" typeStatus="holotype">
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||
<typeStatus box="[738,842,1303,1329]" pageId="23" pageNumber="254">Holotype</typeStatus>
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||
<taxonomicName authority="Cock" authorityName="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula" authorityYear="2023" box="[850,1216,1303,1329]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="pseudocollaris">
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||
<emphasis box="[850,1149,1303,1329]" italics="true" pageId="23" pageNumber="254">Calodesma pseudocollaris</emphasis>
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Cock
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</taxonomicName>
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(to be deposited in
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<collectionCode box="[151,260,1339,1365]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="23" pageNumber="254" type="Museum">NHMUK</collectionCode>
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||
, ex
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||
<collectionCode box="[304,390,1339,1365]" pageId="23" pageNumber="254">MJWC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph blockId="23.[151,1437,1266,2013]" pageId="23" pageNumber="254">
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||
<materialsCitation collectingDate="2017-09-24" collectorName="John Morrall & M. J. W. Cock" country="Trinidad and Tobago" location="Cat's Hill" pageId="23" pageNumber="254" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
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||
<emphasis bold="true" box="[199,327,1374,1404]" pageId="23" pageNumber="254">
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||
<typeStatus box="[199,300,1374,1400]" pageId="23" pageNumber="254">Allotype</typeStatus>
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||
<specimenCount box="[306,327,1376,1404]" pageId="23" pageNumber="254" type="female">♀</specimenCount>
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||
</emphasis>
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||
:
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||
<collectingCountry box="[340,438,1375,1401]" name="Trinidad and Tobago" pageId="23" pageNumber="254">Trinidad</collectingCountry>
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||
, W.I.,
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||
<location LSID="urn:lsid:plazi:treatment:9B35878EBC5FE268FF65FBA5FE30FE70:16436043BC5FE266FE09FAC4FD8CFAFC" box="[507,616,1375,1401]" country="Trinidad and Tobago" name="Cat's Hill" pageId="23" pageNumber="254">Cat’s Hill</location>
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||
, eupatorium flowers,
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||
<date box="[856,979,1375,1401]" pageId="23" pageNumber="254" value="2017-09-24">
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||
<collectingDate box="[856,979,1375,1401]" pageId="23" pageNumber="254" value="2017-09-24">24.ix.2017</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[988,1133,1375,1401]" pageId="23" pageNumber="254">John Morrall</collectorName>
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||
/ DNA sample 143,
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||
<collectorName pageId="23" pageNumber="254">M.J.W. Cock</collectorName>
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||
</materialsCitation>
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||
/
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||
<materialsCitation box="[231,1308,1411,1437]" collectionCode="NHMUK, MJWC" country="Trinidad and Tobago" pageId="23" pageNumber="254" specimenCount="1" typeStatus="allotype">
|
||
<typeStatus box="[231,329,1411,1437]" pageId="23" pageNumber="254">Allotype</typeStatus>
|
||
<taxonomicName authority="Cock" authorityName="Cock" box="[336,839,1411,1437]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="subSpecies" species="maculifrons" subSpecies="pseudocollaris">
|
||
<emphasis box="[336,773,1411,1437]" italics="true" pageId="23" pageNumber="254">Calodesma maculifrons pseudocollaris</emphasis>
|
||
Cock
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||
</taxonomicName>
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||
(to be deposited in
|
||
<collectionCode box="[1061,1170,1411,1437]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="23" pageNumber="254" type="Museum">NHMUK</collectionCode>
|
||
, ex
|
||
<collectionCode box="[1214,1300,1411,1437]" pageId="23" pageNumber="254">MJWC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,1266,2013]" pageId="23" pageNumber="254">
|
||
<materialsCitation collectingDate="2017-09-24" collectorName="John Morrall & M. J. W. Cock" country="Trinidad and Tobago" location="Cat's Hill" pageId="23" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
<emphasis bold="true" box="[199,340,1447,1476]" pageId="23" pageNumber="254">
|
||
<typeStatus box="[199,308,1447,1473]" pageId="23" pageNumber="254">Paratype</typeStatus>
|
||
♁
|
||
</emphasis>
|
||
:
|
||
<collectingCountry box="[359,457,1447,1473]" name="Trinidad and Tobago" pageId="23" pageNumber="254">Trinidad</collectingCountry>
|
||
, W.I.,
|
||
<location LSID="urn:lsid:plazi:treatment:9B35878EBC5FE268FF65FBA5FE30FE70:16436043BC5FE266FDEBFA3CFD68FA44" box="[537,652,1447,1473]" country="Trinidad and Tobago" name="Cat's Hill" pageId="23" pageNumber="254">Cat’s Hill</location>
|
||
, eupatorium flowers,
|
||
<date box="[909,1031,1447,1473]" pageId="23" pageNumber="254" value="2017-09-24">
|
||
<collectingDate box="[909,1031,1447,1473]" pageId="23" pageNumber="254" value="2017-09-24">24.ix.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1047,1197,1447,1473]" pageId="23" pageNumber="254">John Morrall</collectorName>
|
||
/ DNA sample 137,
|
||
<collectorName box="[151,298,1483,1509]" pageId="23" pageNumber="254">M.J.W. Cock</collectorName>
|
||
</materialsCitation>
|
||
/
|
||
<materialsCitation box="[320,1397,1483,1509]" collectionCode="NHMUK, MJWC" country="Trinidad and Tobago" pageId="23" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus box="[320,418,1483,1509]" pageId="23" pageNumber="254">Paratype</typeStatus>
|
||
<taxonomicName authority="Cock" authorityName="Cock" box="[425,928,1483,1509]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="subSpecies" species="maculifrons" subSpecies="pseudocollaris">
|
||
<emphasis box="[425,862,1483,1509]" italics="true" pageId="23" pageNumber="254">Calodesma maculifrons pseudocollaris</emphasis>
|
||
Cock
|
||
</taxonomicName>
|
||
(to be deposited in
|
||
<collectionCode box="[1150,1259,1483,1509]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="23" pageNumber="254" type="Museum">NHMUK</collectionCode>
|
||
, ex
|
||
<collectionCode box="[1303,1389,1483,1509]" pageId="23" pageNumber="254">MJWC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="23" pageNumber="254" type="diagnosis">
|
||
<paragraph blockId="23.[151,1437,1266,2013]" pageId="23" pageNumber="254">
|
||
<emphasis bold="true" box="[199,322,1518,1544]" pageId="23" pageNumber="254">Diagnosis.</emphasis>
|
||
<taxonomicName authority="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula, 2023" authorityName="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula" authorityYear="2023" box="[329,628,1519,1545]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="pseudocollaris" status="new species">
|
||
<emphasis box="[329,628,1519,1545]" italics="true" pageId="23" pageNumber="254">Calodesma pseudocollaris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[636,774,1518,1545]" pageId="23" pageNumber="254">
|
||
<taxonomicNameLabel box="[636,774,1518,1545]" pageId="23" pageNumber="254" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
has males of a
|
||
<taxonomicName authorityName="Drury" authorityYear="1782" box="[953,1039,1519,1545]" class="Insecta" family="Geometridae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis box="[953,1039,1519,1545]" italics="true" pageId="23" pageNumber="254">collaris</emphasis>
|
||
</taxonomicName>
|
||
morph and the only known female is a white morph resembling the
|
||
<typeStatus box="[532,580,1555,1581]" pageId="23" pageNumber="254">type</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Kaye. I" authorityYear="1923" box="[622,750,1555,1581]" class="Magnoliopsida" family="Melastomataceae" genus="Centronia" kingdom="Plantae" order="Myrtales" pageId="23" pageNumber="254" phylum="Tracheophyta" rank="species" species="plorator">
|
||
<emphasis box="[622,750,1555,1581]" italics="true" pageId="23" pageNumber="254">C. plorator</emphasis>
|
||
</taxonomicName>
|
||
. It cannot be distinguished from these morphs of
|
||
<taxonomicName authorityName="BIN" baseAuthorityName="Stoll" baseAuthorityYear="1781" box="[1328,1436,1555,1581]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="militta">
|
||
<emphasis box="[1328,1436,1555,1581]" italics="true" pageId="23" pageNumber="254">C. militta</emphasis>
|
||
</taxonomicName>
|
||
on external morphology or reliably on male genitalia, but can be separated by its DNA barcode. Having said that, the aedeagus of
|
||
<taxonomicName authority="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula, 2023" authorityName="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula" authorityYear="2023" box="[334,532,1627,1653]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="pseudocollaris" status="new species">
|
||
<emphasis box="[334,532,1627,1653]" italics="true" pageId="23" pageNumber="254">C. pseudocollaris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[539,677,1626,1653]" pageId="23" pageNumber="254">
|
||
<taxonomicNameLabel box="[539,677,1626,1653]" pageId="23" pageNumber="254" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
was noted to be less elongate distally, with a shorter distal opening compared to
|
||
<taxonomicName authorityName="BIN" baseAuthorityName="Stoll" baseAuthorityYear="1781" box="[298,403,1663,1689]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="militta">
|
||
<emphasis box="[298,317,1663,1689]" italics="true" pageId="23" pageNumber="254">C</emphasis>
|
||
.
|
||
<emphasis box="[330,403,1663,1689]" italics="true" pageId="23" pageNumber="254">militta</emphasis>
|
||
</taxonomicName>
|
||
, but individual variation would need to be evaluated on a larger sample before this could be relied upon as a diagnostic feature. Further, the
|
||
<specimenCount box="[674,786,1699,1725]" pageId="23" pageNumber="254" type="male">two males</specimenCount>
|
||
are slightly smaller than typical for
|
||
<collectingCountry box="[1183,1278,1699,1725]" name="Trinidad and Tobago" pageId="23" pageNumber="254">Trinidad</collectingCountry>
|
||
material of
|
||
<taxonomicName authorityName="BIN" baseAuthorityName="Stoll" baseAuthorityYear="1781" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="militta">
|
||
<emphasis italics="true" pageId="23" pageNumber="254">C. militta</emphasis>
|
||
</taxonomicName>
|
||
, i.e. forewing length
|
||
<quantity box="[466,587,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="23" pageNumber="254" unit="mm" value="18.0" valueMax="19.0" valueMin="17.0">17–19 mm</quantity>
|
||
(male),
|
||
<quantity box="[681,760,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="254" unit="mm" value="20.0">20 mm</quantity>
|
||
(female), compared to male
|
||
<taxonomicName authorityName="Drury" authorityYear="1782" box="[1084,1170,1735,1761]" class="Insecta" family="Geometridae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis box="[1084,1170,1735,1761]" italics="true" pageId="23" pageNumber="254">collaris</emphasis>
|
||
</taxonomicName>
|
||
morph
|
||
<quantity box="[1258,1358,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="23" pageNumber="254" unit="mm" value="19.2">19.2 mm</quantity>
|
||
(range 18–20, n=6) and white female morphs
|
||
<quantity box="[587,687,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="23" pageNumber="254" unit="mm" value="21.3">21.3 mm</quantity>
|
||
(range
|
||
<quantity box="[773,896,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="254" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">20–22 mm</quantity>
|
||
, n=18) of
|
||
<taxonomicName authorityName="BIN" baseAuthorityName="Stoll" baseAuthorityYear="1781" box="[1013,1119,1771,1797]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="militta">
|
||
<emphasis box="[1013,1119,1771,1797]" italics="true" pageId="23" pageNumber="254">C. militta</emphasis>
|
||
</taxonomicName>
|
||
, but again we have not seen enough material of
|
||
<taxonomicName authority="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula, 2023" authorityName="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula" authorityYear="2023" box="[363,558,1807,1833]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="pseudocollaris" status="new species">
|
||
<emphasis box="[363,382,1807,1833]" italics="true" pageId="23" pageNumber="254">C</emphasis>
|
||
.
|
||
<emphasis box="[393,558,1807,1833]" italics="true" pageId="23" pageNumber="254">pseudocollaris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[563,698,1806,1833]" pageId="23" pageNumber="254">
|
||
<taxonomicNameLabel box="[563,698,1806,1833]" pageId="23" pageNumber="254" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
to know whether this could be a useful character. Males resembling
|
||
<taxonomicName authorityName="C. Felder & R. Felder" authorityYear="1874" box="[151,261,1843,1869]" class="Insecta" genus="Chamesthema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="dioptis">
|
||
<emphasis box="[151,261,1843,1869]" italics="true" pageId="23" pageNumber="254">C. dioptis</emphasis>
|
||
</taxonomicName>
|
||
and females with other variants of the white markings or females with orange or red markings as found in
|
||
<taxonomicName authorityName="BIN" baseAuthorityName="Stoll" baseAuthorityYear="1781" box="[182,290,1879,1905]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="254" phylum="Arthropoda" rank="species" species="militta">
|
||
<emphasis box="[182,290,1879,1905]" italics="true" pageId="23" pageNumber="254">C. militta</emphasis>
|
||
</taxonomicName>
|
||
are not known for this new species, but the small sample size does not preclude these possibilities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="25" lastPageNumber="256" pageId="23" pageNumber="254" type="description">
|
||
<paragraph blockId="23.[151,1437,1266,2013]" lastBlockId="24.[151,1437,151,1077]" lastPageId="24" lastPageNumber="255" pageId="23" pageNumber="254">
|
||
<emphasis bold="true" box="[199,337,1914,1940]" pageId="23" pageNumber="254">Description</emphasis>
|
||
.
|
||
<emphasis bold="true" box="[355,416,1915,1941]" pageId="23" pageNumber="254">Male</emphasis>
|
||
as shown in
|
||
<figureCitation box="[576,656,1915,1941]" captionStart="FIGURE 9" captionStartId="10.[151,250,676,700]" captionTargetBox="[207,1379,200,641]" captionTargetId="figure-18@10.[189,1398,181,653]" captionTargetPageId="10" captionText="FIGURE 9. Type material Calodesma pseudocollaris new species Trinidad, W.I., Cat’s Hill, at eupatorium flowers, 24.ix.2017, J. Morrall; dorsal view above, ventral view below. A, holotype, ♁ DNA 137. B, paratype ♁, DNA 135, dissection 1001. C, allotype ♀, DNA 143. Life size." figureDoi="http://doi.org/10.5281/zenodo.7850685" httpUri="https://zenodo.org/record/7850685/files/figure.png" pageId="23" pageNumber="254">Figs. 9</figureCitation>
|
||
A-B (full view), 16 A–C (head and anterior thorax), and 10 (male genitalia).
|
||
<emphasis bold="true" box="[273,337,1951,1977]" pageId="23" pageNumber="254">Head</emphasis>
|
||
. Lateral and ventral head orange; dorsum dark, with two white dots above the scape; frons dark with some orange setae and a white border; antennae black, with white scaling on pedicel, bipectinate almost to apex; eyes black; labial palpi segment 1 pale orange; labial palpi segments 2 and 3 black with inner lower margin white.
|
||
<emphasis bold="true" box="[151,240,187,213]" pageId="24" pageNumber="255">Thorax</emphasis>
|
||
. Collar orange. Dorsally black apart from inner margins of tegulae are white-orange and there is a very short orange dorsal line between the tegulae. Ventrally prothorax and mesothorax orange shading into blackish posteriorly on mesothorax; metathorax black. Forelegs back, with two white-orange lines on the femur, two white lines on the posterior tibia and white scales on the outer margin of the remainder of the leg. Midlegs black, with one (
|
||
<typeStatus box="[1327,1428,295,321]" pageId="24" pageNumber="255">holotype</typeStatus>
|
||
) or two (
|
||
<typeStatus box="[243,343,331,357]" pageId="24" pageNumber="255">paratype</typeStatus>
|
||
) white lines the length of posterior tibia, and the remainder of the leg dark (
|
||
<typeStatus box="[1228,1332,331,357]" pageId="24" pageNumber="255">holotype</typeStatus>
|
||
), or with white scales on outer margin (
|
||
<typeStatus box="[482,585,367,393]" pageId="24" pageNumber="255">paratype</typeStatus>
|
||
). Hindlegs missing on
|
||
<typeStatus box="[835,937,367,393]" pageId="24" pageNumber="255">holotype</typeStatus>
|
||
; as midlegs on
|
||
<typeStatus box="[1105,1205,367,393]" pageId="24" pageNumber="255">paratype</typeStatus>
|
||
.Abdomen black, the posterior margin of the sternites white, and orange scales around the edge of the genitalia.
|
||
<emphasis bold="true" box="[1162,1238,402,428]" pageId="24" pageNumber="255">Wings</emphasis>
|
||
. Forewing length
|
||
<quantity box="[151,232,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="255" unit="mm" value="17.0">17 mm</quantity>
|
||
(
|
||
<typeStatus box="[248,352,439,465]" pageId="24" pageNumber="255">holotype</typeStatus>
|
||
),
|
||
<quantity box="[373,454,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="24" pageNumber="255" unit="mm" value="19.0">19 mm</quantity>
|
||
(
|
||
<typeStatus box="[470,573,439,465]" pageId="24" pageNumber="255">paratype</typeStatus>
|
||
). Dorsal forewing black, with the veins of the dorsal half paler; large white postdiscal spot; fringe black, weakly white at apex. Ventral forewing black; white postdiscal spot as dorsally; fringe white at apex, weakly so in spaces 1B and 2 (CU
|
||
<subScript attach="left" box="[715,723,529,543]" fontSize="6" pageId="24" pageNumber="255">2</subScript>
|
||
-2A and Cu
|
||
<subScript attach="left" box="[856,864,529,543]" fontSize="6" pageId="24" pageNumber="255">1</subScript>
|
||
-Cu
|
||
<subScript attach="right" box="[906,914,529,543]" fontSize="6" pageId="24" pageNumber="255">2</subScript>
|
||
). Dorsal hindwing black, with a narrow, faint white spot at apex; fringe white. Ventral hindwing black, with veins paler in the basal part of the wing; apical spot more clearly marked than in dorsal view; fringe white.
|
||
<emphasis bold="true" box="[778,888,582,608]" pageId="24" pageNumber="255">Genitalia</emphasis>
|
||
. Small and weakly sclerotised. Tegumen domed and bulbous. Uncus simple, acute, strongly downcurved distally; a broad, bilobed, very weakly sclerotised dorsal project projects posteriorly behind tegumen. Valvae almost membranous, simple. Saccus well developed. Aedeagus short, with a strong bulbous ventral lobe at base; the apex obliquely truncate; no indication of structures associated with the vesica.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,1077]" pageId="24" pageNumber="255">
|
||
<emphasis bold="true" box="[199,286,763,789]" pageId="24" pageNumber="255">Female</emphasis>
|
||
as shown in
|
||
<figureCitation box="[433,529,763,789]" captionStart="FIGURE 9" captionStartId="10.[151,250,676,700]" captionTargetBox="[207,1379,200,641]" captionTargetId="figure-18@10.[189,1398,181,653]" captionTargetPageId="10" captionText="FIGURE 9. Type material Calodesma pseudocollaris new species Trinidad, W.I., Cat’s Hill, at eupatorium flowers, 24.ix.2017, J. Morrall; dorsal view above, ventral view below. A, holotype, ♁ DNA 137. B, paratype ♁, DNA 135, dissection 1001. C, allotype ♀, DNA 143. Life size." figureDoi="http://doi.org/10.5281/zenodo.7850685" httpUri="https://zenodo.org/record/7850685/files/figure.png" pageId="24" pageNumber="255">Figs 9 C</figureCitation>
|
||
(full view) and 16 D–F (head and anterior thorax). Generally similar to the male apart from the weakly serrate antennae, no orange scales around genital opening, and the larger, broader, less pointed forewings, and more rounded hindwing. Forewing length
|
||
<quantity box="[795,874,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="255" unit="mm" value="20.0">20 mm</quantity>
|
||
(
|
||
<typeStatus box="[888,984,835,861]" pageId="24" pageNumber="255">allotype</typeStatus>
|
||
). The white markings of the wings differ from the male. Dorsal forewing with the white discal spot narrower, extending to the costa and in a narrow arc to the dorsum just basal to tornus; the white markings on the costa and in space 3 (M
|
||
<subScript attach="left" box="[1013,1021,925,939]" fontSize="6" pageId="24" pageNumber="255">3</subScript>
|
||
-Cu
|
||
<subScript attach="left" box="[1063,1071,925,939]" fontSize="6" pageId="24" pageNumber="255">1</subScript>
|
||
) are suffused with black. Ventral forewing as dorsal; fringe black. Dorsal hindwing with no white apical spot; a weak white marginal-submarginal band, strongly suffused with black and with the veins black; the band starts narrow on the margin in space 1B (Cu
|
||
<subScript attach="left" box="[1419,1427,997,1011]" fontSize="6" pageId="24" pageNumber="255">2</subScript>
|
||
- 2A) and widens and separates from the margin until space 5 (M
|
||
<subScript attach="right" box="[891,899,1033,1047]" fontSize="6" pageId="24" pageNumber="255">1</subScript>
|
||
-M
|
||
<subScript attach="left" box="[933,941,1033,1047]" fontSize="6" pageId="24" pageNumber="255">2</subScript>
|
||
). Ventral hindwing as dorsal but marginal-submarginal band less suffused with black; fringe black, slightly paler at apex. Genitalia not examined.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7850707" ID-Zenodo-Dep="7850707" httpUri="https://zenodo.org/record/7850707/files/figure.png" pageId="24" pageNumber="255" startId="24.[151,250,1878,1902]" targetBox="[151,1436,1098,1852]" targetPageId="24" targetType="figure">
|
||
<paragraph blockId="24.[151,1437,1878,1939]" pageId="24" pageNumber="255">
|
||
<emphasis bold="true" box="[151,280,1878,1903]" pageId="24" pageNumber="255">FIGURE 16</emphasis>
|
||
Views of the head and anterior thorax of
|
||
<taxonomicName authority="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula, 2023" authorityName="Cock & Laguerre & Buddie & Cafa & Alston-Smith & Morrall & Gosula" authorityYear="2023" box="[690,958,1879,1903]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="255" phylum="Arthropoda" rank="species" species="pseudocollaris" status="new species">
|
||
<emphasis box="[690,958,1879,1903]" italics="true" pageId="24" pageNumber="255">Calodesma pseudocollaris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[964,1093,1878,1903]" pageId="24" pageNumber="255">
|
||
<taxonomicNameLabel box="[964,1088,1878,1903]" pageId="24" pageNumber="255" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</emphasis>
|
||
A–C paratype male. D–F allotype female.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="25.[151,1436,150,501]" pageId="25" pageNumber="256">
|
||
<emphasis bold="true" box="[199,379,150,177]" pageId="25" pageNumber="256">DNA barcodes.</emphasis>
|
||
Three identical barcodes were obtained, which form BIN BOLD:AEI2170, most similar to
|
||
<emphasis box="[1411,1430,151,177]" italics="true" pageId="25" pageNumber="256">C</emphasis>
|
||
. sp.
|
||
<taxonomicName authority="BOLD" authorityName="BOLD" box="[190,396,187,213]" class="Insecta" family="Geometridae" genus="Phalaena" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="collaris">
|
||
cf.
|
||
<emphasis box="[226,312,187,213]" italics="true" pageId="25" pageNumber="256">collaris</emphasis>
|
||
BOLD
|
||
</taxonomicName>
|
||
:ABZ2392 and
|
||
<taxonomicName authority="BOLD" authorityName="BOLD" box="[569,819,187,213]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="maculifrons">
|
||
<emphasis box="[569,734,187,213]" italics="true" pageId="25" pageNumber="256">C. maculifrons</emphasis>
|
||
BOLD
|
||
</taxonomicName>
|
||
:AAA1430 (
|
||
<figureCitation box="[957,1041,187,213]" captionStart="FIGURE 12" captionStartId="17.[151,250,1612,1636]" captionTargetBox="[172,1415,189,1578]" captionTargetId="figure-18@17.[162,1425,179,1586]" captionTargetPageId="17" captionText="FIGURE 12. Similarity relationships of Calodesma spp. based on this work, barcodes from the ML collection, and selected public sequences in BOLD. Many duplicate DNA barcodes available for C. maculifrons from Costa Rica and some for C. militta from Trinidad are omitted so as to facilitate presentation. The taxonomy used is that developed in this work, and where relevant the names used here for the various morphs are included in parentheses after the scientific name. Bootstrap values are shown only if>70% and distances only if>0.002.The evolutionary history was inferred using the Neighbor-Joining method (Saitou and Nei 1987). The optimal tree with the sum of branch length = 0.15482979 is shown. The percentage of replicate trees in which the associated taxa clustered together in the bootstrap test (1000 replicates) are shown next to the branches (Felsenstein 1985). The tree is drawn to scale, with branch lengths (next to the branches) in the same units as those of the evolutionary distances used to infer the phylogenetic tree. The evolutionary distances were computed using the Kimura 2-parameter method (Tamura et al. 2004) and are in the units of the number of base substitutions per site. The analysis involved 73 nucleotide sequences. Codon positions included were 1st+2nd+3rd+Noncoding. All ambiguous positions were removed for each sequence pair. There was a total of 689 positions in the final dataset. Evolutionary analyses were conducted in MEGAX (Kumar et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.7850695" httpUri="https://zenodo.org/record/7850695/files/figure.png" pageId="25" pageNumber="256">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="25.[151,1436,150,501]" pageId="25" pageNumber="256">
|
||
<emphasis bold="true" box="[199,327,222,248]" pageId="25" pageNumber="256">Variability</emphasis>
|
||
. Only
|
||
<specimenCount box="[407,522,223,249]" pageId="25" pageNumber="256" type="male">two males</specimenCount>
|
||
of the
|
||
<taxonomicName box="[605,691,223,249]" class="Insecta" family="Geometridae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis box="[605,691,223,249]" italics="true" pageId="25" pageNumber="256">collaris</emphasis>
|
||
</taxonomicName>
|
||
morph and one white female morph have been sequenced. Given these small numbers and the relative frequency of the different morphs in
|
||
<taxonomicName authorityName="BIN" baseAuthorityName="Stoll" baseAuthorityYear="1781" box="[1010,1118,259,285]" class="Insecta" family="Arctiidae" genus="Calodesma" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="militta">
|
||
<emphasis box="[1010,1118,259,285]" italics="true" pageId="25" pageNumber="256">C. militta</emphasis>
|
||
</taxonomicName>
|
||
, our sample is too small to comment on the probability of this subspecies occurring in other morphs and colours.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="25" pageNumber="256" type="distribution">
|
||
<paragraph blockId="25.[151,1436,150,501]" pageId="25" pageNumber="256">
|
||
<emphasis bold="true" box="[199,353,330,356]" pageId="25" pageNumber="256">Distribution.</emphasis>
|
||
<collectingCountry box="[358,591,331,357]" name="Trinidad and Tobago" pageId="25" pageNumber="256">Trinidad and Tobago</collectingCountry>
|
||
,
|
||
<collectingCountry box="[601,701,331,357]" name="Trinidad and Tobago" pageId="25" pageNumber="256">Trinidad</collectingCountry>
|
||
: Cat’s Hill (
|
||
<typeStatus box="[831,881,331,357]" pageId="25" pageNumber="256">type</typeStatus>
|
||
series). This is an area of disturbed lowland forest with scattered on-going oil extraction using pumpjacks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="25" pageNumber="256" type="etymology">
|
||
<paragraph blockId="25.[151,1436,150,501]" pageId="25" pageNumber="256">
|
||
<emphasis bold="true" box="[199,334,403,429]" pageId="25" pageNumber="256">Etymology.</emphasis>
|
||
The name refers to the resemblance of the males to
|
||
<taxonomicName box="[911,1029,403,429]" class="Insecta" family="Geometridae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis box="[911,1029,403,429]" italics="true" pageId="25" pageNumber="256">C. collaris</emphasis>
|
||
</taxonomicName>
|
||
, but that they are a different species, i.e. false
|
||
<taxonomicName box="[254,340,439,465]" class="Insecta" family="Geometridae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis box="[254,340,439,465]" italics="true" pageId="25" pageNumber="256">collaris</emphasis>
|
||
</taxonomicName>
|
||
. The original name,
|
||
<taxonomicName box="[574,660,439,465]" class="Insecta" family="Geometridae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis box="[574,660,439,465]" italics="true" pageId="25" pageNumber="256">collaris</emphasis>
|
||
</taxonomicName>
|
||
, is adjectival in the genitive case, unchanging for gender agreement; it refers to the orange collar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |