313 lines
39 KiB
XML
313 lines
39 KiB
XML
<document id="94FE64590D5E42B5A6EA4C20ECE01124" ID-DOI="10.11646/zootaxa.3734.2.1" ID-GBIF-Dataset="06b7c7bc-cbcc-4314-9718-c2349641c8bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="247499" ID-ZooBank="9A217106-EDDF-4129-BA53-55B4F48BF82E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460695387635" checkinUser="plazi" docAuthor="Maza-Benignos, Mauricio De La & Lozano-Vilano, Ma. De Lourdes" docDate="2013" docId="9746878AFFB8FFAE76CAFB8AFDFB5B67" docLanguage="en" docName="zt03734p129.pdf" docOrigin="Zootaxa 3734 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Herichthys pantostictus Taylor & Miller 1983" docType="treatment" docVersion="9" lastPageNumber="109" masterDocId="6B7FFFF2FFBDFFA6765DFFC8FFC9583A" masterDocTitle="Description of three new species of the genus Herichthys (Perciformes: Cichlidae) from eastern Mexico, with redescription of H. labridens, H. steindachneri, and H. pantostictus" masterLastPageNumber="129" masterPageNumber="101" pageNumber="106" updateTime="1698353782969" updateUser="plazi">
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<mods:titleInfo id="9B0519B4B20911633534AF46F733D0C1">
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<mods:title id="088E67994C2B4BBFBC1FB2D5F4E97F9D">Description of three new species of the genus Herichthys (Perciformes: Cichlidae) from eastern Mexico, with redescription of H. labridens, H. steindachneri, and H. pantostictus</mods:title>
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<mods:name id="8360B6E50E7CC369361F2CF37A22EEA2" type="personal">
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<mods:namePart id="AD4B5154C9E76710B9BB73D6B81BD471">Maza-Benignos, Mauricio De La</mods:namePart>
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<mods:name id="8C8C866D7DB51DEC930506008AA09E15" type="personal">
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<mods:roleTerm id="234CBA196CCB16AC7BF10D50513535E2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C70D8936EBD4DB6D233A4DAA04641DBC">Lozano-Vilano, Ma. De Lourdes</mods:namePart>
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<mods:typeOfResource id="482927A48E8BD88924259937642099D1">text</mods:typeOfResource>
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<mods:title id="AAF43E51E09189322ABDD1D92521003C">Zootaxa</mods:title>
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<mods:part id="1A17AE9DD0D959E64949E45C7E10A914">
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<mods:date id="4DE8ADC2640ACAE0A832301B3A5725A1">2013</mods:date>
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<mods:number id="C379683295A71533037F5B33636009C8">3734</mods:number>
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<mods:detail id="C5E455C371E4964D6C92C519892FDF69" type="issue">
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<mods:number id="7B0450A1DD5B1C46F76346AABC9ABFDA">2</mods:number>
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<mods:classification id="FBA842ED4376F1C3D7D7F1B7D65E1674">journal article</mods:classification>
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<mods:identifier id="96BEA3E12B1E750B58F722A9ED60B50B" type="DOI">10.11646/zootaxa.3734.2.1</mods:identifier>
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<mods:identifier id="6F9F3E93B971E40B83738CE6CD887830" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BCBA9AD105575D10B7B7EF8FF5842A8E" type="Zenodo-Dep">247499</mods:identifier>
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<treatment id="9746878AFFB8FFAE76CAFB8AFDFB5B67" ID-DOI="http://doi.org/10.5281/zenodo.5619848" ID-GBIF-Taxon="119581834" ID-Zenodo-Dep="5619848" LSID="urn:lsid:plazi:treatment:9746878AFFB8FFAE76CAFB8AFDFB5B67" httpUri="http://treatment.plazi.org/id/9746878AFFB8FFAE76CAFB8AFDFB5B67" lastPageId="8" lastPageNumber="109" pageId="5" pageNumber="106">
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<subSubSection id="57F56517FFB8FFA376CAFB8AFDEF5C44" pageId="5" pageNumber="106" type="nomenclature">
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<paragraph id="1F50369CFFB8FFA376CAFB8AFD115C67" blockId="5.[151,728,1090,1150]" box="[151,728,1090,1117]" pageId="5" pageNumber="106">
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<heading id="441881F0FFB8FFA376CAFB8AFD115C67" bold="true" box="[151,728,1090,1117]" fontSize="11" level="1" pageId="5" pageNumber="106" reason="1">
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<taxonomicName id="D8EF4D1FFFB8FFA376CAFB8AFD115C67" ID-CoL="3KYWX" authority="Taylor & Miller 1983" authorityName="Taylor & Miller" authorityYear="1983" box="[151,728,1090,1117]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="pantostictus">
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<emphasis id="2D9BEA8EFFB8FFA376CAFB8AFD115C67" bold="true" box="[151,728,1090,1117]" pageId="5" pageNumber="106">
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<emphasis id="2D9BEA8EFFB8FFA376CAFB8AFE7C5C66" bold="true" box="[151,437,1090,1116]" italics="true" pageId="5" pageNumber="106">Herichthys pantostictus</emphasis>
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(Taylor & Miller 1983)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="1F50369CFFB8FFA376CAFBADFDEF5C44" blockId="5.[151,728,1090,1150]" box="[151,550,1125,1150]" pageId="5" pageNumber="106">
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<figureCitation id="87D42A19FFB8FFA376CAFBADFECB5C44" box="[151,258,1125,1150]" captionStart="FIGURE 4" captionStartId="6.[151,250,1843,1865]" captionTargetBox="[151,1436,1137,1821]" captionTargetId="figure@6.[151,1436,1137,1821]" captionTargetPageId="6" captionText="FIGURE 4. Herichthys pantostictus (Taylor & Miller 1983) collected from Rio Naranjos in the Tamiahua lagoon system (~ 95 mm TL)." httpUri="https://zenodo.org/record/247503/files/figure.png" pageId="5" pageNumber="106">Figures 4</figureCitation>
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,
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<figureCitation id="87D42A19FFB8FFA37753FBADFED75C44" box="[270,286,1125,1150]" captionStart="FIGURE 5" captionStartId="7.[151,250,819,841]" captionTargetBox="[217,1369,193,796]" captionTargetId="figure@7.[217,1369,193,797]" captionTargetPageId="7" captionText="FIGURE 5. UANL 14685 Herichthys pantostictus collected from Laguna la Puerta (Tampico), Tamaulipas (62.4 mm SL). Mauricio De la Maza-Benignos, Figures 6 – 7." httpUri="https://zenodo.org/record/247504/files/figure.png" pageId="5" pageNumber="106">5</figureCitation>
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, and 15; Tables 4 and 5
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</paragraph>
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</subSubSection>
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<subSubSection id="57F56517FFB8FFA376CAFB65FD825E4C" pageId="5" pageNumber="106" type="materials_examined">
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<paragraph id="1F50369CFFB8FFA376CAFB65FD4E5CD0" blockId="5.[151,1437,1197,2014]" pageId="5" pageNumber="106">
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<emphasis id="2D9BEA8EFFB8FFA376CAFB65FEC25CFC" bold="true" box="[151,267,1197,1222]" pageId="5" pageNumber="106">
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<typeStatus id="C054883EFFB8FFA376CAFB65FECF5CFC" box="[151,262,1197,1222]" pageId="5" pageNumber="106" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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UMMZ 207699 (1:
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<quantity id="D8179B79FFB8FFA377B8FB65FD9A5CFF" box="[485,595,1197,1222]" metricMagnitude="-2" metricUnit="m" metricValue="7.95" pageId="5" pageNumber="106" unit="mm" value="79.5">79.5 mm</quantity>
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SL, nongravid female) Laguna de Chairel near Tampico,
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<collectingCountry id="67F8760CFFB8FFA372BAFB65FAF65CFC" box="[1255,1343,1197,1222]" name="Mexico" pageId="5" pageNumber="106">México</collectingCountry>
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, R.M. y J.H. Darnell y E. Liner,
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<date id="6B51105CFFB8FFA377FEFB19FD4A5CD0" box="[419,643,1233,1258]" pageId="5" pageNumber="106" value="1952-12-29">December 29, 1952</date>
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.
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</paragraph>
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<paragraph id="1F50369CFFB8FFA3769AFB3DFD825E4C" blockId="5.[151,1437,1197,2014]" pageId="5" pageNumber="106">
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<emphasis id="2D9BEA8EFFB8FFA3769AFB3DFE795D34" bold="true" box="[199,432,1269,1294]" pageId="5" pageNumber="106">Material examined.</emphasis>
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Eighty-one specimens,
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<quantity id="D8179B79FFB8FFA374E0FB3DFC905D37" box="[701,857,1269,1294]" metricMagnitude="-2" metricUnit="m" metricValue="8.045" metricValueMax="12.69" metricValueMin="3.4" pageId="5" pageNumber="106" unit="mm" value="80.45" valueMax="126.9" valueMin="34.0">34–126.9 mm</quantity>
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SL.
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<emphasis id="2D9BEA8EFFB8FFA375D2FB3DFBEA5D34" bold="true" box="[911,1059,1269,1294]" pageId="5" pageNumber="106">Tamaulipas:</emphasis>
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UANL 17428 (13:
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<quantity id="D8179B79FFB8FFA372AFFB3DFA555D37" box="[1266,1436,1269,1294]" metricMagnitude="-2" metricUnit="m" metricValue="7.275" metricValueMax="8.98" metricValueMin="5.57" pageId="5" pageNumber="106" unit="mm" value="72.75" valueMax="89.8" valueMin="55.7">55.7–89.8 mm</quantity>
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SL), Ojito de Jaumave, Lat. 23.933, Long. −100.05, 778 masl,
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<collectingCountry id="67F8760CFFB8FFA3753AFAD1FC075D08" box="[871,974,1305,1330]" name="Mauritius" pageId="5" pageNumber="106">Mauricio</collectingCountry>
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de la Maza Benignos (MMB),
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<date id="6B51105CFFB8FFA3736AFAD1FECA5D6C" pageId="5" pageNumber="106" value="2006-02-25">February 25, 2006</date>
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; UANL 17431 (6:
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<quantity id="D8179B79FFB8FFA37783FAF5FD475D6F" box="[478,654,1341,1366]" metricMagnitude="-2" metricUnit="m" metricValue="5.665" metricValueMax="7.39" metricValueMin="3.94" pageId="5" pageNumber="106" unit="mm" value="56.65" valueMax="73.9" valueMin="39.4">39.4–73.9 mm</quantity>
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SL), Guayalejo, Lat. 23.665, Long. −100.223, 568 masl, MMB,
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<date id="6B51105CFFB8FFA376CAFAA9FEA05D40" box="[151,361,1377,1402]" pageId="5" pageNumber="106" value="2006-02-22">February 22, 2006</date>
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; UANL 17473 (12:
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<quantity id="D8179B79FFB8FFA3741DFAA9FD235D43" box="[576,746,1377,1402]" metricMagnitude="-2" metricUnit="m" metricValue="5.145" metricValueMax="6.36" metricValueMin="3.9299999999999997" pageId="5" pageNumber="106" unit="mm" value="51.45" valueMax="63.6" valueMin="39.3">39.3–63.6 mm</quantity>
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SL), Río Mante, Lat. 22.73071667, Long. −98.994, 80 masl, MMB, May 0 9, 2006; UANL 17477, (10:
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<quantity id="D8179B79FFB8FFA37439FA4DFCC65DA7" box="[612,783,1413,1438]" metricMagnitude="-2" metricUnit="m" metricValue="6.83" metricValueMax="9.12" metricValueMin="4.54" pageId="5" pageNumber="106" unit="mm" value="68.3" valueMax="91.2" valueMin="45.4">45.4–91.2 mm</quantity>
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SL), Canal Mante, Lat. 22.717, Long. −98.9902, 82 masl, MMB May 0 9, 2006; UANL 17480 (6:
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<quantity id="D8179B79FFB8FFA37411FA61FD265DFB" box="[588,751,1449,1474]" metricMagnitude="-2" metricUnit="m" metricValue="8.33" metricValueMax="10.4" metricValueMin="6.26" pageId="5" pageNumber="106" unit="mm" value="83.3" valueMax="104.0" valueMin="62.6">62.6–104 mm</quantity>
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SL), El Encino, Lat. 23.13 Long. −99.617, 119 masl, MMB,
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<date id="6B51105CFFB8FFA376CAFA06FEE65DDC" box="[151,303,1485,1510]" pageId="5" pageNumber="106" value="2006-05-13">May13, 2006</date>
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; UANL 14685 (6:
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<quantity id="D8179B79FFB8FFA377A8FA05FD415DDF" box="[501,648,1485,1510]" metricMagnitude="-2" metricUnit="m" metricValue="4.82" metricValueMax="6.24" metricValueMin="3.4" pageId="5" pageNumber="106" unit="mm" value="48.2" valueMax="62.4" valueMin="34.0">34–62.4 mm</quantity>
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SL), Laguna de la Puerta,
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<quantity id="D8179B79FFB8FFA375EFFA06FC305DDC" box="[946,1017,1485,1510]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="106" unit="km" value="12.0">12 km</quantity>
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northwest (NW) of Tampico on Hwy 100, SCB,
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<date id="6B51105CFFB8FFA37747FA39FDCA5E30" box="[282,515,1521,1546]" pageId="5" pageNumber="106" value="1964-12-29">December 29, 1964</date>
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;
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<emphasis id="2D9BEA8EFFB8FFA3744FFA39FD405E30" bold="true" box="[530,649,1521,1546]" pageId="5" pageNumber="106">Veracruz:</emphasis>
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UANL 17443 (5:
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<quantity id="D8179B79FFB8FFA3750BFA39FBCD5E33" box="[854,1028,1521,1546]" metricMagnitude="-2" metricUnit="m" metricValue="6.595" metricValueMax="7.67" metricValueMin="5.52" pageId="5" pageNumber="106" unit="mm" value="65.95" valueMax="76.7" valueMin="55.2">55.2–76.7 mm</quantity>
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SL), Naranjos, Lat. 22.228, Long. −98.605, 48 masl, MMB,
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<date id="6B51105CFFB8FFA37799F9DDFDB75E14" box="[452,638,1557,1582]" pageId="5" pageNumber="106" value="2006-03-17">March 17, 2006</date>
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; UANL 17511 (1:
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<quantity id="D8179B79FFB8FFA3751AF9DDFC0C5E17" box="[839,965,1557,1582]" metricMagnitude="-1" metricUnit="m" metricValue="1.269" pageId="5" pageNumber="106" unit="mm" value="126.9">126.9 mm</quantity>
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SL), Platón Sánchez, Lat. 21.295, Long. −98.364, 55 masl, MMB,
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<date id="6B51105CFFB8FFA3778CF9F2FDB45E68" box="[465,637,1593,1618]" pageId="5" pageNumber="106" value="2006-06-30">June 30, 2006</date>
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; UANL 17513 (22:
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<quantity id="D8179B79FFB8FFA37539F9F1FC345E6B" box="[868,1021,1593,1618]" metricMagnitude="-2" metricUnit="m" metricValue="6.62" metricValueMax="7.04" metricValueMin="6.2" pageId="5" pageNumber="106" unit="mm" value="66.2" valueMax="70.4" valueMin="62.0">62–70.4 mm</quantity>
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SL), Naranjos, Lat. 22.228, Long. −98.605, 48 masl, MMB,
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<date id="6B51105CFFB8FFA377E7F995FD815E4C" box="[442,584,1629,1654]" pageId="5" pageNumber="106" value="2006-07-01">July 1, 2006</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="57F56517FFB8FFA3769AF949FE3E5F83" pageId="5" pageNumber="106" type="diagnosis">
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<paragraph id="1F50369CFFB8FFA3769AF949FE3E5F83" blockId="5.[151,1437,1197,2014]" pageId="5" pageNumber="106">
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<emphasis id="2D9BEA8EFFB8FFA3769AF949FE885EA0" bold="true" box="[199,321,1665,1690]" pageId="5" pageNumber="106">Diagnosis.</emphasis>
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Distinguished using the following combination of characters: shallow cheeks (mean 28%, SD 3%), large eye diameter (26%, SD 2% HL). In live fish the ground color is olive brown, but in preservation the body is brown with lighter color in the flanks. Head and dorsal areas dark; ventral areas light brown or whitish. Lower pharyngeal plate moderately stout and broad; indentation
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<typeStatus id="C054883EFFB8FFA37547F926FC835F3C" box="[794,842,1774,1798]" pageId="5" pageNumber="106">type</typeStatus>
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polymorphic with 2 rows of 6–7 combined enlargedconic or medium-sized molariform teeth, increasing in size posteriorly, flanking the midline. Indentation
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<typeStatus id="C054883EFFB8FFA37331F8DAFA555F10" box="[1388,1436,1810,1834]" pageId="5" pageNumber="106">type</typeStatus>
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polymorphic; <20 conic and enlarged teeth along the posterior margin. Coastal and lagoon populations (i.e., Chairel and Rio Tigre) as well as many river populations have the entire body covered with dark dots placed at the base of the scales, darker and less marked on edges, except for the abdomen and venter of head. Dots show irregular rows in the body area.
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</paragraph>
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</subSubSection>
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<subSubSection id="57F56517FFB8FFAE769AF80DFD1859F5" lastPageId="8" lastPageNumber="109" pageId="5" pageNumber="106" type="description">
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<paragraph id="1F50369CFFB8FFA0769AF80DFE8E5BB3" blockId="5.[151,1437,1197,2014]" lastBlockId="6.[151,1436,151,1085]" lastPageId="6" lastPageNumber="107" pageId="5" pageNumber="106">
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<emphasis id="2D9BEA8EFFB8FFA3769AF80DFE905FE4" bold="true" box="[199,345,1989,2014]" pageId="5" pageNumber="106">Description.</emphasis>
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Derived from sexually mature specimens>
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<quantity id="D8179B79FFB8FFA3753CF80EFC7F5FE7" box="[865,950,1990,2014]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="106" unit="mm" value="39.0">39 mm</quantity>
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SL. Morphometric and meristic data are summarized in Tables 4 and 5. Species is polymorphic throughout its range; body elongated, depth, 36%–44% SL (mean 40%, SD 2%); head long, 36%–39% (mean 37%, SD 1%) SL; caudal peduncle, shallow, 14%–17% (mean 15%, SD 1%) SL. Predorsal contour steep and non-acute, curved out before the eye, forehead inconspicuous, and nuchal hump absent; dorsal and ventral contours straight to moderately convex with a moderate slope; mouth terminal and pointed; frenum weakly developed or absent. Dorsal fin XVI–XVII (mode XVI, freq 45%), 9–11 (mode 11, freq 35%); fin set forward of the opercle (intersects between the 1st and 3rd rays); depressed fin short, tip rarely expands beyond the anterior 3rd of the caudal fin. Anal fin V–VI (modeVI, freq 70%), 8–10 (mode 8, freq 70%); scales in longitudinal series 28–31 (mode 30, freq 50%). Anterior teeth regularly set, well-spaced, conic, unicuspid, strongly recurved, and pointed, with erect implantation; upper jaw with a pair of well-developed recurved fangs that are reddish in color. Lower jaw with 2nd, less-developed, pair of recurved fangs; 15–22 teeth in outer series of the premaxilla, 6–8 regularly set of acutely pointed, caniniform frontal teeth with moderately stout necks; lateral and posterior teeth small, non-caniniform, and irregularly set. Lower jaw with 2–3 rows of teeth; upper jaw with 3 rows of teeth; posterior rows of diminutive, irregularly set, and hardly visible teeth. Lower pharyngeal plate moderately stout and broad; indentation
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<typeStatus id="C054883EFFBBFFA0750DFDBDFC495AB7" box="[848,896,629,653]" pageId="6" pageNumber="107">type</typeStatus>
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polymorphic with moderately robust, lightly pigmented molars in some populations (i.e., Guayalejo) and enlarged, conic-flattened to moderately pointed and non-enlarged pigmented teeth in others (i.e., Tamiahua); 2 rows of 6–7 medium-sized enlarged conic teeth or molars increasing posteriorly in size; molarization flanking midline. Conic teeth along posterior margin conic and numbering 18–20. Stomach robust-walled, moderately saccular (16.42% of SL), rugged with longitudinal folds, and adhered at its anterodorsal section to a well-elongated, elastic, smooth caecum (27% of SL). Gut-coiling pattern with a double medial loop; secondary loop projecting away and passing under the gut. Peritoneum uniformly dark.
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</paragraph>
|
||
<paragraph id="1F50369CFFBBFFA0769AFC5CFA5F5C07" blockId="6.[151,1436,151,1085]" pageId="6" pageNumber="107">
|
||
<emphasis id="2D9BEA8EFFBBFFA0769AFC5CFDC45B97" bold="true" box="[199,525,916,941]" pageId="6" pageNumber="107">Coloration in preservative.</emphasis>
|
||
Body brown, lighter on flanks. Head and dorsal areas dark; ventral areas light brown or whitish. Coastal forms and some lotic populations and individuals with bodies entirely covered with dots, except for the belly and underside of the head. Caudal, soft dorsal, and anal fins marked with streaks. Patterns along the flanks vary between populations from a faint discontinuous horizontal stripe to 3–5 irregularly formed blotches and a dark spot at the base of the caudal fin. Markings may be faint or completely lacking in older adults.
|
||
</paragraph>
|
||
<caption id="4B906614FFBBFFA076CAF8FBFF3A5F52" httpUri="https://zenodo.org/record/247503/files/figure.png" pageId="6" pageNumber="107" targetBox="[151,1436,1137,1821]" targetPageId="6">
|
||
<paragraph id="1F50369CFFBBFFA076CAF8FBFF3A5F52" blockId="6.[151,1435,1843,1896]" pageId="6" pageNumber="107">
|
||
<emphasis id="2D9BEA8EFFBBFFA076CAF8FBFEDA5F72" bold="true" box="[151,275,1843,1865]" pageId="6" pageNumber="107">FIGURE 4.</emphasis>
|
||
<taxonomicName id="D8EF4D1FFFBBFFA07744F8FBFD355F73" authority="Taylor & Miller 1983" authorityName="Taylor & Miller" authorityYear="1983" box="[281,764,1843,1865]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFBBFFA07744F8FBFDC35F73" box="[281,522,1843,1865]" italics="true" pageId="6" pageNumber="107">Herichthys pantostictus</emphasis>
|
||
(Taylor & Miller 1983)
|
||
</taxonomicName>
|
||
collected from Rio Naranjos in the Tamiahua lagoon system (~95 mm TL).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="4B906614FFBAFFA176CAFCFBFDAB5B5D" httpUri="https://zenodo.org/record/247504/files/figure.png" pageId="7" pageNumber="108" targetBox="[217,1369,193,796]" targetPageId="7">
|
||
<paragraph id="1F50369CFFBAFFA176CAFCFBFDAB5B5D" blockId="7.[151,1436,819,871]" pageId="7" pageNumber="108">
|
||
<emphasis id="2D9BEA8EFFBAFFA176CAFCFBFEDC5B72" bold="true" box="[151,277,819,841]" pageId="7" pageNumber="108">FIGURE 5.</emphasis>
|
||
UANL 14685
|
||
<taxonomicName id="D8EF4D1FFFBAFFA177EBFCFBFD605B73" box="[438,681,819,841]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="108" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFBAFFA177EBFCFBFD605B73" box="[438,681,819,841]" italics="true" pageId="7" pageNumber="108">Herichthys pantostictus</emphasis>
|
||
</taxonomicName>
|
||
collected from Laguna la Puerta (Tampico), Tamaulipas (62.4 mm SL). Mauricio De la Maza-Benignos, Figures 6–7.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1F50369CFFBAFFA1769AFC5CFBA25CBF" blockId="7.[151,1437,916,2023]" pageId="7" pageNumber="108">
|
||
<emphasis id="2D9BEA8EFFBAFFA1769AFC5CFE9A5B97" bold="true" box="[199,339,916,941]" pageId="7" pageNumber="108">Live colors.</emphasis>
|
||
Body olive brown; pectoral, caudal, and soft sections of dorsal and anal fins tan to light orange. Inter-radial membranes of dorsal and anal soft fins and caudal fin with dots and streaks, all centered on basal areas, particularly in spiny sections. Coastal and lagoon populations (i.e., Chairel and Rio Tigre), as well as many river populations, of
|
||
<taxonomicName id="D8EF4D1FFFBAFFA17711FBC9FE315C22" box="[332,504,1025,1048]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="108" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFBAFFA17711FBC9FE315C22" box="[332,504,1025,1048]" italics="true" pageId="7" pageNumber="108">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
with entire body covered with dark dots at bases of scales; dots bigger and less marked at scale edges, except for abdomen and ventral region of the head. Dots are in irregular rows along body; other river populations (i.e., most Tamiahua, few Guayalejo, and none of the
|
||
<taxonomicName id="D8EF4D1FFFBAFFA175AAFB81FB805C5A" box="[1015,1097,1097,1120]" class="Actinopterygii" family="Poeciliidae" genus="Gambusia" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="119" phylum="Chordata" rank="species" species="panuco">Panuco</taxonomicName>
|
||
lotic populations) with few to no dots on flanks. All populations with a small, red–purple mark on axil of pectoral fin.
|
||
</paragraph>
|
||
<paragraph id="1F50369CFFBAFFA1769AFB58FE7F5E0D" blockId="7.[151,1437,916,2023]" pageId="7" pageNumber="108">
|
||
<emphasis id="2D9BEA8EFFBAFFA1769AFB58FE205C93" bold="true" box="[199,489,1168,1193]" pageId="7" pageNumber="108">Breeding pigmentation.</emphasis>
|
||
<collectingCountry id="67F8760CFFBAFFA177A4FB59FDD15C93" box="[505,536,1169,1193]" name="American Samoa" pageId="7" pageNumber="108">As</collectingCountry>
|
||
noted by Salazar-Gonzáles (2007), breeding pigmentation varies between populations. Populations of Tamaulipas, including those from Guayalejo, Mante, Jaumave, and El Encino, develop darkening over anteroventral and posterior halves (regions) of the body. This forms a continuum that is interrupted by a 70°-angled vertical, pale area up the side and across the flank. The pale area begins anterior to the midsection of the abdomen, extends posteriorly to become horizontal with the hypural fan, and ends frontal to the anal fin base at 1/3rd the distance between the anal and pectoral fin origins. Populations of San Luis Potosí and Veracruz, including Tampaón and Platón Sánchez, develop darkening over the anteroventral and posterior halves (regions) of the body, separated by a pale area with a frontal point of origin halfway between anal and pectoral fin origins and up the side at 90° until it merges with the dorsal pale section; the posterior point of origin of the pale area begins at the midpoint of the spiny anal fin base and extends up the side at 100° to merge with the dorsal pale section. Darkening on the posterior half does not reach the dorsal contour anterior to the interception of the dorsal fin base with the caudal peduncle.
|
||
</paragraph>
|
||
<paragraph id="1F50369CFFBAFFA1769AF98BFE565ED1" blockId="7.[151,1437,916,2023]" pageId="7" pageNumber="108">
|
||
<emphasis id="2D9BEA8EFFBAFFA1769AF98BFDC25E66" bold="true" box="[199,523,1603,1628]" pageId="7" pageNumber="108">Geographical distribution.</emphasis>
|
||
Río Tigre in Tamaulipas (San Andrés lagoon system), the Rio Naranjos in Veracruz (the Tamiahua lagoon system), coastal lagoons of Tampico, including Chairel, the Tamesí-Guayalejo subbasin from Jaumave Spring; The western limit of the species is the Rio
|
||
<taxonomicName id="D8EF4D1FFFBAFFA17581F943FBE75E99" box="[988,1070,1675,1699]" class="Actinopterygii" family="Poeciliidae" genus="Gambusia" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="119" phylum="Chordata" rank="species" species="panuco">Pánuco</taxonomicName>
|
||
basin, excluding the Rio Verde valley, Media Luna, “Rio El Salto,” “Rio Gallinas,” and the headwaters of the Rio Moctezuma in the states of Hidalgo and Queretaro.
|
||
</paragraph>
|
||
<paragraph id="1F50369CFFBAFFA1769AF93EFB0B5FA5" blockId="7.[151,1437,916,2023]" pageId="7" pageNumber="108">
|
||
<emphasis id="2D9BEA8EFFBAFFA1769AF93EFE155F35" bold="true" box="[199,476,1782,1807]" pageId="7" pageNumber="108">Habitat and associates.</emphasis>
|
||
<taxonomicName id="D8EF4D1FFFBAFFA177B9F93EFD275F35" box="[484,750,1782,1807]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="108" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFBAFFA177B9F93EFD275F35" box="[484,750,1782,1807]" italics="true" pageId="7" pageNumber="108">Herichthys pantostictus</emphasis>
|
||
</taxonomicName>
|
||
is sympatric with
|
||
<taxonomicName id="D8EF4D1FFFBAFFA1759EF930FB835F35" box="[963,1098,1784,1807]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="108" phylum="Chordata" rank="species" species="carpintis">
|
||
<emphasis id="2D9BEA8EFFBAFFA1759EF930FB835F35" box="[963,1098,1784,1807]" italics="true" pageId="7" pageNumber="108">H. carpintis</emphasis>
|
||
</taxonomicName>
|
||
throughout most of its range, except in some high elevation sites such as Jaumave spring, Tamaulipas, where it shares its habitat with
|
||
<taxonomicName id="D8EF4D1FFFBAFFA176CAF8F6FEE15F6D" box="[151,296,1854,1879]" class="Actinopterygii" family="Poeciliidae" genus="Xiphophorus" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="108" phylum="Chordata" rank="genus">
|
||
<emphasis id="2D9BEA8EFFBAFFA176CAF8F6FEE15F6D" box="[151,296,1854,1879]" italics="true" pageId="7" pageNumber="108">Xiphophorus</emphasis>
|
||
</taxonomicName>
|
||
sp. cf.
|
||
<emphasis id="2D9BEA8EFFBAFFA17727F888FE1F5F6D" box="[378,470,1856,1879]" italics="true" pageId="7" pageNumber="108">variatus</emphasis>
|
||
and
|
||
<emphasis id="2D9BEA8EFFBAFFA17450F8F7FD315F6D" box="[525,760,1855,1879]" italics="true" pageId="7" pageNumber="108">
|
||
<taxonomicName id="D8EF4D1FFFBAFFA17450F8F7FD3C5F6D" box="[525,757,1855,1879]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="108" phylum="Chordata" rank="species" species="mexicanus">Astyanax mexicanus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
among other species. It prospers in habitats that range from turbid coastal lagoon systems with a muddy substrate to clear running waters and springs with rocky or sandy substrate. Stomach contents include detritus (10%), filamentous algae (80%), and snails (10%).
|
||
</paragraph>
|
||
<paragraph id="1F50369CFFBAFFA1769AF863FCF75FFE" blockId="7.[151,1437,916,2023]" box="[199,830,1963,1988]" pageId="7" pageNumber="108">
|
||
<emphasis id="2D9BEA8EFFBAFFA1769AF863FE6E5FFE" bold="true" box="[199,423,1963,1988]" pageId="7" pageNumber="108">Vernacular names.</emphasis>
|
||
Mojarra de Chairel, Chairel cichlid.
|
||
</paragraph>
|
||
<paragraph id="1F50369CFFBAFFAE769AF806FEA3595F" blockId="7.[151,1437,916,2023]" lastBlockId="8.[151,1436,151,861]" lastPageId="8" lastPageNumber="109" pageId="7" pageNumber="108">
|
||
<emphasis id="2D9BEA8EFFBAFFA1769AF806FE0D5FDD" bold="true" box="[199,452,1998,2023]" pageId="7" pageNumber="108">Geographic variants.</emphasis>
|
||
<taxonomicName id="D8EF4D1FFFBAFFA17791F806FD1E5FDD" box="[460,727,1998,2023]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="108" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFBAFFA17791F806FD1E5FDD" box="[460,727,1998,2023]" italics="true" pageId="7" pageNumber="108">Herichthys pantostictus</emphasis>
|
||
</taxonomicName>
|
||
is composed of a number of parapatric populations inhabiting the lower
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE775AFF50FE905895" box="[263,345,152,175]" class="Actinopterygii" family="Poeciliidae" genus="Gambusia" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="panuco">Panuco</taxonomicName>
|
||
and the Rio Tamesí basins as well as tributaries to the adjacent Tamiahua and San Andrés lagoon systems. The species displays geographic variation in morphometric, meristic, and breeding pigmentation as observed by Salazar-Gonzáles (2007).
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE7417FF29FD3D58C2" box="[586,756,225,248]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE7417FF29FD3D58C2" box="[586,756,225,248]" italics="true" pageId="8" pageNumber="109">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
is parapatric with
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE759BFF29FB9A58C2" box="[966,1107,223,248]" class="Actinopterygii" family="Cichlidae" genus="Heros" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="labridens">
|
||
<emphasis id="2D9BEA8EFFB5FFAE759BFF29FB9A58C2" box="[966,1107,223,248]" italics="true" pageId="8" pageNumber="109">H. labridens</emphasis>
|
||
</taxonomicName>
|
||
and possibly
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE72B3FF29FABA58C2" box="[1262,1395,223,248]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="molango" status="sp. nov.">
|
||
<emphasis id="2D9BEA8EFFB5FFAE72B3FF29FABA58C2" box="[1262,1395,223,248]" italics="true" pageId="8" pageNumber="109">H. molango</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2D9BEA8EFFB5FFAE7327FF17FF015927" bold="true" pageId="8" pageNumber="109">
|
||
<taxonomicNameLabel id="36A857F5FFB5FFAE7327FF17FF015927" pageId="8" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Geographic variation in
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE77B5FECDFD5A5926" box="[488,659,261,284]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE77B5FECDFD5A5926" box="[488,659,261,284]" italics="true" pageId="8" pageNumber="109">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
is the most conspicuous between coastal lagoon forms and riverine forms of the Rio Tamesí-Guayalejo and Rio
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE74D5FEE0FD135905" box="[648,730,296,319]" class="Actinopterygii" family="Poeciliidae" genus="Gambusia" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="panuco">Panuco</taxonomicName>
|
||
and their tributaries. River forms are generally more elongated than lagoon forms.
|
||
</paragraph>
|
||
<paragraph id="1F50369CFFB5FFAE769AFEA7FD1859F5" blockId="8.[151,1436,151,861]" pageId="8" pageNumber="109">
|
||
<emphasis id="2D9BEA8EFFB5FFAE769AFEA7FE7459B2" bold="true" box="[199,445,367,392]" pageId="8" pageNumber="109">Conservation status.</emphasis>
|
||
Not listed in NORMA Oficial
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE757AFEA7FC5C59B2" box="[807,917,367,392]" class="Actinopterygii" family="Poeciliidae" genus="Poecilia" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="129" phylum="Chordata" rank="species" species="mexicana">Mexicana</taxonomicName>
|
||
NOM-059-ECOL-2010 or on the IUCN Red List of Threatened Species 2009.
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE744DFE5CFCD35996" box="[528,794,404,429]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE744DFE5CFCD35996" box="[528,794,404,429]" italics="true" pageId="8" pageNumber="109">Herichthys pantostictus</emphasis>
|
||
</taxonomicName>
|
||
appears to be stable throughout most of its range, except in the coastal lagoon areas where it appears scarce.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="57F56517FFB5FFAE769AFE14FDFB5B67" pageId="8" pageNumber="109" type="discussion">
|
||
<paragraph id="1F50369CFFB5FFAE769AFE14FDFB5B67" blockId="8.[151,1436,151,861]" pageId="8" pageNumber="109">
|
||
<emphasis id="2D9BEA8EFFB5FFAE769AFE14FEF259CF" bold="true" box="[199,315,476,501]" pageId="8" pageNumber="109">Remarks.</emphasis>
|
||
This species was described by Taylor & Miller (1983) as
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE725FFE14FAD559CE" box="[1026,1308,476,501]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE725FFE14FAD559CE" box="[1026,1308,476,501]" italics="true" pageId="8" pageNumber="109">Cichlasoma pantostictus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<typeStatus id="C054883EFFB5FFAE7336FE15FA5259CF" box="[1387,1435,477,501]" pageId="8" pageNumber="109">type</typeStatus>
|
||
locality of Laguna de Chairel (Tampico), and a population in the Rio Sabinas (the Tamesí-Guayalejo river system in Tamaulipas). Specimens can be described as having the entire body covered by small dark dots, except for the belly and ventral surface of the head. While lentic populations of
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE75DFFD81FBE45A5A" box="[898,1069,585,608]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE75DFFD81FBE45A5A" box="[898,1069,585,608]" italics="true" pageId="8" pageNumber="109">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
show the “entirely dotted” trait, which is a key to distinguishing between
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE7428FDA5FCE85ABE" box="[629,801,621,644]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE7428FDA5FCE85ABE" box="[629,801,621,644]" italics="true" pageId="8" pageNumber="109">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE7501FDA5FC235ABF" box="[860,1002,620,645]" class="Actinopterygii" family="Cichlidae" genus="Heros" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="labridens">
|
||
<emphasis id="2D9BEA8EFFB5FFAE7501FDA5FC235ABF" box="[860,1002,620,645]" italics="true" pageId="8" pageNumber="109">H. labridens</emphasis>
|
||
</taxonomicName>
|
||
according to Taylor & Miller (1983), many riverine populations and individuals are not entirely dotted. It is possible that the geographic distribution of the trait drove Taylor & Miller (1983) to erroneously conclude that an “entirely dotted pattern” is key to separating
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE76CAFD11FE8B5ACA" box="[151,322,729,752]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE76CAFD11FE8B5ACA" box="[151,322,729,752]" italics="true" pageId="8" pageNumber="109">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE77D7FD11FDD15ACA" box="[394,536,727,752]" class="Actinopterygii" family="Cichlidae" genus="Heros" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="labridens">
|
||
<emphasis id="2D9BEA8EFFB5FFAE77D7FD11FDD15ACA" box="[394,536,727,752]" italics="true" pageId="8" pageNumber="109">H. labridens</emphasis>
|
||
</taxonomicName>
|
||
and that
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE74D8FD11FCF95ACA" box="[645,816,729,752]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE74D8FD11FCF95ACA" box="[645,816,729,752]" italics="true" pageId="8" pageNumber="109">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
were sympatric or parapatric with
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE729EFD11FA985ACA" box="[1219,1361,727,752]" class="Actinopterygii" family="Cichlidae" genus="Heros" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="labridens">
|
||
<emphasis id="2D9BEA8EFFB5FFAE729EFD11FA985ACA" box="[1219,1361,727,752]" italics="true" pageId="8" pageNumber="109">H. labridens</emphasis>
|
||
</taxonomicName>
|
||
of the Guayalejo, Sabinas at Gómez Farías, Ojo de Jaumave, Río Tigre, and Río Mante. Hulsey
|
||
<emphasis id="2D9BEA8EFFB5FFAE7221FD35FB7D5B2F" box="[1148,1204,764,789]" italics="true" pageId="8" pageNumber="109">et al.</emphasis>
|
||
(2004) recovered
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE73DCFD35FED75B02" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="2D9BEA8EFFB5FFAE73DCFD35FED75B02" italics="true" pageId="8" pageNumber="109">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
of Rio Tamesí, Rio Guayalejo, and Rio Tigre as sister taxa to a clade shared with
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE72E1FCE9FAD55B02" box="[1212,1308,801,824]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="pame" status="sp. nov.">
|
||
<emphasis id="2D9BEA8EFFB5FFAE72E1FCE9FAD55B02" box="[1212,1308,801,824]" italics="true" pageId="8" pageNumber="109">H. pame</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2D9BEA8EFFB5FFAE737EFCD7FAB45B02" bold="true" box="[1315,1405,799,824]" pageId="8" pageNumber="109">
|
||
<taxonomicNameLabel id="36A857F5FFB5FFAE737EFCD7FAB45B02" box="[1315,1405,799,824]" pageId="8" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
of Rio Tamasopo and
|
||
<emphasis id="2D9BEA8EFFB5FFAE7732FC8DFDFB5B67" box="[367,562,836,861]" italics="true" pageId="8" pageNumber="109">
|
||
<taxonomicName id="D8EF4D1FFFB5FFAE7732FC8DFDE75B67" box="[367,558,836,861]" class="Actinopterygii" family="Cichlidae" genus="Herichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="109" phylum="Chordata" rank="species" species="steindachneri">H. steindachneri</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |