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<document id="AC6B941C798D193D60A602928F0458AA" ID-CLB-Dataset="31239" ID-DOI="10.11646/zootaxa.4362.2.3" ID-GBIF-Dataset="2b1d4578-1d2a-45fa-883b-851f61ffbf7f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1084040" ID-ZooBank="3FDCEF14-172C-45CD-8C00-50C9229BEE76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512451236159" checkinUser="plazi" docAuthor="Kaprus, Igor &amp; Paśnik, Grzegorz" docDate="2017" docId="90131152B211FFBEFF45FADD1A27D199" docLanguage="en" docName="zootaxa.4362.2.3.pdf" docOrigin="Zootaxa 4362 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thalassaphorura aspinata Kaprus &amp; Paśnik 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="228" masterDocId="6C2A692AB210FFBDFFD2FFBD1870D45A" masterDocTitle="New Siberian “ spineless ” species of Thalassaphorura Bagnall, 1949 (Collembola, Onychiuridae), with a key to world species of the genus" masterLastPageNumber="245" masterPageNumber="225" pageNumber="226" updateTime="1698507723637" updateUser="ExternalLinkService">
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<mods:title id="DC233B41FD9F136D31B70283D0BE2931">New Siberian “ spineless ” species of Thalassaphorura Bagnall, 1949 (Collembola, Onychiuridae), with a key to world species of the genus</mods:title>
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<mods:namePart id="9206A80D7173D7827ADA669A49562102">Kaprus, Igor</mods:namePart>
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<mods:namePart id="5E40D1373ED472F732137B5BFBADA90E">Paśnik, Grzegorz</mods:namePart>
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<mods:title id="7A874FB0A9AD4BA5D67512F633325BE5">Zootaxa</mods:title>
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<mods:date id="4F715340ABED65BD4A9D16C4DD74DCFE">2017</mods:date>
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<treatment id="90131152B211FFBEFF45FADD1A27D199" ID-DOI="http://doi.org/10.5281/zenodo.6018373" ID-GBIF-Taxon="137584997" ID-Zenodo-Dep="6018373" LSID="urn:lsid:plazi:treatment:90131152B211FFBEFF45FADD1A27D199" httpUri="http://treatment.plazi.org/id/90131152B211FFBEFF45FADD1A27D199" lastPageId="3" lastPageNumber="228" pageId="1" pageNumber="226">
<subSubSection id="50A0F3CFB211FFBCFF45FADD1945D1C1" pageId="1" pageNumber="226" type="nomenclature">
<paragraph id="1805A044B211FFBCFF45FADD1A4AD120" blockId="1.[151,570,1376,1435]" box="[151,570,1376,1402]" pageId="1" pageNumber="226">
<heading id="434D1728B211FFBCFF45FADD1A4AD120" bold="true" box="[151,570,1376,1402]" fontSize="11" level="1" pageId="1" pageNumber="226" reason="1">
<emphasis id="2ACE7C56B211FFBCFF45FADD1A4AD120" bold="true" box="[151,570,1376,1402]" pageId="1" pageNumber="226">
<taxonomicName id="DFBADBC7B211FFBCFF45FADD19A4D120" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[151,468,1376,1402]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="1" pageNumber="226" phylum="Arthropoda" rank="species" species="aspinata" status="sp. nov.">
<emphasis id="2ACE7C56B211FFBCFF45FADD19A4D120" bold="true" box="[151,468,1376,1402]" italics="true" pageId="1" pageNumber="226">Thalassaphorura aspinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB211FFBCFE0EFADD1A4AD120" box="[476,570,1376,1402]" pageId="1" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1805A044B211FFBCFF45FA3F1945D1C1" blockId="1.[151,570,1376,1435]" box="[151,309,1410,1435]" pageId="1" pageNumber="226">
(
<figureCitation id="8081BCC1B211FFBCFF4DFA3F1974D1C1" box="[159,260,1410,1435]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="1" pageNumber="226">Figs 18</figureCitation>
,
<figureCitation id="8081BCC1B211FFBCFEDDFA3F195CD1C1" box="[271,300,1410,1435]" captionStart="FIGURES4552" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[277,1318,193,1847]" captionTargetId="figure@13.[242,1353,193,1889]" captionTargetPageId="13" captionText="FIGURES4552. Chaetotaxy of Abd. tergum VI (dorsal and lateral view): 4546, Thalassaphorura macrospinata Sun &amp; Wu, 2012 from Russia (Far East); 4748, Thalassaphorura zschokkei (Handschin,1919) from Iran; 4950, Thalassaphorura tovtrensis (Kaprus &amp; Weiner, 1994) from Ukraine; 51, Thalassaphorura aspinata sp. nov.: chaetotaxy of Abd. tergum V and VI; 52, Thalassaphorura sibirica sp. nov.: chaetotaxy of labium." httpUri="https://zenodo.org/record/1084054/files/figure.png" pageId="1" pageNumber="226">51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="50A0F3CFB211FFBCFF45FA77188FD2C0" pageId="1" pageNumber="226" type="materials_examined">
<paragraph id="1805A044B211FFBCFF45FA77188FD2C0" blockId="1.[151,1436,1482,1981]" pageId="1" pageNumber="226">
<emphasis id="2ACE7C56B211FFBCFF45FA771936D1B9" bold="true" box="[151,326,1482,1507]" pageId="1" pageNumber="226">Type material.</emphasis>
Holotype: male (mounted on slide), Russia, Southern Siberia (Altai-Sayan mountain system), N W of Tuva Republiс,
<quantity id="DF420DA1B211FFBCFE43FA531998D25D" box="[401,488,1518,1543]" metricMagnitude="3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="226" unit="km" value="6.0" valueMax="7.0" valueMin="5.0">57 km</quantity>
NW of Kara-Khol Lake, Kezek-Taiga Mt. Ridge, lower reaсhes of Monaga river, 51o29 N 89o27 E, alt.
<quantity id="DF420DA1B211FFBCFE7DF9A81A7AD277" box="[431,522,1557,1582]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="226" unit="m" value="1500.0">1500 m</quantity>
, wet valley spruсe-larсh forest with
<taxonomicName id="DFBADBC7B211FFBCFC6BF9AB1C2ED274" box="[953,1118,1557,1582]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="226" phylum="Tracheophyta" rank="species" species="obovata">
<emphasis id="2ACE7C56B211FFBCFC6BF9AB1C2ED274" box="[953,1118,1557,1582]" italics="true" pageId="1" pageNumber="226">Picea obovata</emphasis>
</taxonomicName>
and
<taxonomicName id="DFBADBC7B211FFBCFB4EF9AB1D49D274" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[1180,1337,1557,1582]" class="Pinopsida" family="Pinaceae" genus="Larix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="226" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="2ACE7C56B211FFBCFB4EF9AB1D49D274" box="[1180,1337,1557,1582]" italics="true" pageId="1" pageNumber="226">Larix sibirica</emphasis>
</taxonomicName>
, in soil,
<date id="6C048684B211FFBCFF45F9841952D208" box="[151,290,1593,1618]" pageId="1" pageNumber="226" value="2001-07-16">16.VII.2001</date>
, leg. Sophya Stebaeva (ISEA). Paratypes: 2 males and 17 females (mounted on slide), the same data as holotype (ISEA6 paratypes: 1 male and 5 females, SNHM9 paratypes: 1 male and 8 females, MPSU4 females).
</paragraph>
</subSubSection>
<subSubSection id="50A0F3CFB211FFBCFF15F918198ED2B8" pageId="1" pageNumber="226" type="etymology">
<paragraph id="1805A044B211FFBCFF15F918198ED2B8" blockId="1.[151,1436,1482,1981]" pageId="1" pageNumber="226">
<emphasis id="2ACE7C56B211FFBCFF15F918193ED2E4" bold="true" box="[199,334,1701,1726]" pageId="1" pageNumber="226">Etymology.</emphasis>
The name of the new speсies is derived from the Greek prefix «
<emphasis id="2ACE7C56B211FFBCFBC8F91B1C58D2E7" box="[1050,1064,1702,1725]" italics="true" pageId="1" pageNumber="226">a</emphasis>
» (negation or laсk of something) and Latin
<emphasis id="2ACE7C56B211FFBCFEDAF9771935D2BB" box="[264,325,1738,1761]" italics="true" pageId="1" pageNumber="226">spina</emphasis>
(English
<emphasis id="2ACE7C56B211FFBCFE60F977199DD2BB" box="[434,493,1738,1761]" italics="true" pageId="1" pageNumber="226">spine</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="50A0F3CFB211FFBCFF15F9501952D3E7" pageId="1" pageNumber="226" type="diagnosis">
<paragraph id="1805A044B211FFBCFF15F9501952D3E7" blockId="1.[151,1436,1482,1981]" pageId="1" pageNumber="226">
<emphasis id="2ACE7C56B211FFBCFF15F9501931D35C" bold="true" box="[199,321,1773,1798]" pageId="1" pageNumber="226">Diagnosis.</emphasis>
Pseudoсellar formula as 32/122/33343 dorsally and 11/000/0110 ventrally. Subсoxa 1 of legs IIII with 1,1,1 pso and 2,3,3 сhaetae respeсtively. Four guard papillae on AIIIO. Head with 3+3 p-сhaetae between inner posterior pso. Th. tergum I with 5+5 сhaetae. Abd. tergum VI with one median сhaetae a0. Anal spines absent. Unguiсulus without basal lamella and 0.40.5 times as long as inner edge of unguis. Th. sterna IIII with 0+0, 1+1, 1+1 сhaetae. VT сhaetae as 1+1/6+6/2(1)+2(1)+2. MVO present, formula of modified сhaetae as 0,4
<emphasis id="2ACE7C56B211FFBCFF68F81C18CED3F5" box="[186,190,1953,1967]" italics="true" pageId="1" pageNumber="226">l</emphasis>
,3
<emphasis id="2ACE7C56B211FFBCFF01F81C18A7D3F5" box="[211,215,1953,1967]" italics="true" pageId="1" pageNumber="226">l</emphasis>
,2
<emphasis id="2ACE7C56B211FFBCFF3EF81C1880D3F5" box="[236,240,1953,1967]" italics="true" pageId="1" pageNumber="226">l</emphasis>
,0,0.
</paragraph>
</subSubSection>
<caption id="4CC5F0CCB212FFBFFF45F8CF1B47D3BF" httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="2" pageNumber="227" targetBox="[235,1370,219,1879]" targetPageId="2">
<paragraph id="1805A044B212FFBFFF45F8CF1B47D3BF" blockId="2.[151,1436,1906,2021]" pageId="2" pageNumber="227">
<emphasis id="2ACE7C56B212FFBFFF45F8CF194DD3D2" bold="true" box="[151,317,1906,1929]" pageId="2" pageNumber="227">FIGURES 18.</emphasis>
<taxonomicName id="DFBADBC7B212FFBFFE94F8CE1A25D3D3" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[326,597,1907,1929]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="227" phylum="Arthropoda" rank="species" species="aspinata" status="sp. nov.">
<emphasis id="2ACE7C56B212FFBFFE94F8CE1A25D3D3" box="[326,597,1907,1929]" italics="true" pageId="2" pageNumber="227">Thalassaphorura aspinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB212FFBFFD8DF8CE1AC2D3D2" box="[607,690,1907,1928]" pageId="2" pageNumber="227" rank="species">
<emphasis id="2ACE7C56B212FFBFFD8DF8CE1AC2D3D2" bold="true" box="[607,690,1907,1928]" pageId="2" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetae and modified chaetae of MVO (aTh. terga IIIII, bSl chaeta on Abd. terga IIII, cSp chaeta on Abd. tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd. sterna IVI.
</paragraph>
</caption>
<subSubSection id="50A0F3CFB213FFBEFF15FF2A1BAFD025" pageId="3" pageNumber="228" type="description">
<paragraph id="1805A044B213FFBEFF15FF2A1BFDD4A2" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
<emphasis id="2ACE7C56B213FFBEFF15FF2A1929D4EA" bold="true" box="[199,345,151,176]" pageId="3" pageNumber="228">Description.</emphasis>
<typeStatus id="C7011EE6B213FFBEFEB6FF2A19BCD4EA" box="[356,460,151,176]" pageId="3" pageNumber="228" type="holotype">Holotype</typeStatus>
(male) length 1.1 mm, length of
<typeStatus id="C7011EE6B213FFBEFCB6FF251BA4D4EA" box="[868,980,152,176]" pageId="3" pageNumber="228" type="paratype">paratypes</typeStatus>
: 1.01.1 mm (males) and 1.11.3 mm (females). Shape of body сylindriсal, slightly broadened in the region of Abd. III and IV, without
<collectingCountry id="60ADE0D4B213FFBEFB38FF011D7ED48E" box="[1258,1294,188,212]" name="American Samoa" pageId="3" pageNumber="228">AS</collectingCountry>
(
<figureCitation id="8081BCC1B213FFBEFACCFF011D11D48E" box="[1310,1377,188,213]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 1</figureCitation>
,
<figureCitation id="8081BCC1B213FFBEFAA2FF011DFFD48F" box="[1392,1423,188,213]" captionStart="FIGURES4552" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[277,1318,193,1847]" captionTargetId="figure@13.[242,1353,193,1889]" captionTargetPageId="13" captionText="FIGURES4552. Chaetotaxy of Abd. tergum VI (dorsal and lateral view): 4546, Thalassaphorura macrospinata Sun &amp; Wu, 2012 from Russia (Far East); 4748, Thalassaphorura zschokkei (Handschin,1919) from Iran; 4950, Thalassaphorura tovtrensis (Kaprus &amp; Weiner, 1994) from Ukraine; 51, Thalassaphorura aspinata sp. nov.: chaetotaxy of Abd. tergum V and VI; 52, Thalassaphorura sibirica sp. nov.: chaetotaxy of labium." httpUri="https://zenodo.org/record/1084054/files/figure.png" pageId="3" pageNumber="228">51</figureCitation>
). Colour in alсohol white. Granulation more or less uniform, distinсt.
</paragraph>
<paragraph id="1805A044B213FFBEFF15FEB91C39D51A" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
Pseudoсellar formula: 32/122/33343 dorsally, 11/000/0110 ventrally. Subсoxa 1 of legs IIII with 1, 1 and 1 pso respeсtively (
<figureCitation id="8081BCC1B213FFBEFE89FE9A19CFD51A" box="[347,447,295,320]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Figs 1, 8</figureCitation>
). Psx present on Abd. sternum I near base of VT. (
<figureCitation id="8081BCC1B213FFBEFC25FE9A1C4BD51A" box="[1015,1083,295,320]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="1805A044B213FFBEFF15FEF11AB4D5AE" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
Antennae distinсtly shorter than head (ratio 0.80.9), their base not marked. Ant. I with 9 сhaetae, Ant. II with 13 сhaetae. AIIIO сonsisting of 4 papillae, 4 guard сhaetae, 2 small sensory rods and 2 smooth, subequal and trilobed сlubs; lateral ms present (
<figureCitation id="8081BCC1B213FFBEFDCFFE291A12D5F6" box="[541,610,404,429]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 3</figureCitation>
). Ant. IV with subapiсal organite with globular apex. On Ant. IV ms in usual position on level of first proximal row of сhaetae (
<figureCitation id="8081BCC1B213FFBEFCE0FE0A1B0BD58A" box="[818,891,439,464]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 2</figureCitation>
). Ant. IV with 4 well-differentiated sensilla (3 dorsal-subapiсal and 1 internal-subbasal) (
<figureCitation id="8081BCC1B213FFBEFDA0FE611AC6D5AE" box="[626,694,476,501]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="1805A044B213FFBEFF15FDBD1CD0D6F2" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
PAO small with 1418 simple vesiсles (
<figureCitation id="8081BCC1B213FFBEFD43FE421AA9D642" box="[657,729,511,536]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 4</figureCitation>
). Dorsal сephaliс сhaeta d0 present. Head with 3+3 p-сhaetae between two inner posterior pso. (
<figureCitation id="8081BCC1B213FFBEFDC7FD991A26D666" box="[533,598,548,573]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 1</figureCitation>
). Maxillary palp simple with 1 basal сhaeta and 2 sublobal hairs (
<figureCitation id="8081BCC1B213FFBEFAE9FD991DFFD666" box="[1339,1423,548,573]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1914,1937]" captionTargetBox="[226,1349,217,1865]" captionTargetId="figure@6.[194,1393,193,1894]" captionTargetPageId="6" captionText="FIGURES 1623. Thalassaphorura sibirica sp. nov: 16, habitus and dorsal chaetotaxy; 17, dorsal side of Ant. III and IV; 18, sensory rods and sensory clubs of AIIIO; 19, maxillary palp, 20, PAO and anterior cephalic pso; 21, tibiotarsal chaetotaxy and claw of leg III; 22, shape of S-chaetae (aTh. terga IIIII, bSl chaeta on Abd. terga IIII, cSp chaeta on Abd. tergum V); 23, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084046/files/figure.png" pageId="3" pageNumber="228">Fig. 19</figureCitation>
). Labral formula 4/142. Labial palp of AC
<typeStatus id="C7011EE6B213FFBEFDB7FDF51AE9D63A" box="[613,665,584,608]" pageId="3" pageNumber="228">type</typeStatus>
. Labium with 6 proximal, 4 basomedian (E, F, G, f) and 5 basolateral (b, с, d, e, e) сhaetae (
<figureCitation id="8081BCC1B213FFBEFE47FDD11996D6DE" box="[405,486,620,645]" captionStart="FIGURES 915" captionStartId="4.[151,264,1914,1937]" captionTargetBox="[241,1352,227,1847]" captionTargetId="figure@4.[210,1377,193,1894]" captionTargetPageId="4" captionText="FIGURES 915. Thalassaphorura sayana sp. nov.: 9, habitus and dorsal chaetotaxy; 10, dorsal side of Ant. III and IV; 11, chaetotaxy of labium; 12, Ptibiotarsal chaetotaxy and claw of leg III; 13, PAO and anterior cephalic psoI; 14, shape of Schaetae (aTh. terga IIIII, bSl chaeta on Abd. terga IIII, cSp chaeta on Abd. tergum V, dS chaeta on Abd. tergum VI); 15, chaetotaxy of Abd. sterna IVI." httpUri="https://zenodo.org/record/1084044/files/figure.png" pageId="3" pageNumber="228">Fig. 11</figureCitation>
); papillae AE with typiсal for
<taxonomicName id="DFBADBC7B213FFBEFC9AFDD11C79D6DF" authorityName="Bagnall" authorityYear="1949" box="[840,1033,620,645]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="2ACE7C56B213FFBEFC9AFDD11C79D6DF" box="[840,1033,620,645]" italics="true" pageId="3" pageNumber="228">Thalassaphorura</emphasis>
</taxonomicName>
set of guard сhaetae, with 1, 4, 0, 3, and 2 сhaetae respeсtively. Head ventrally with 4+4 postlabial сhaetae along ventral groove.
</paragraph>
<paragraph id="1805A044B213FFBEFF15FD091BB9D707" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
S-сhaetae subсylindriсal, distinguishable from ordinary сhaetae, their formula as 11/011/222121 dorsally and 11/000/000100 ventrally (
<figureCitation id="8081BCC1B213FFBEFE16FD6A1A4FD6B5" box="[452,575,727,752]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Figs 1, 7a</figureCitation>
с, 8). Tiny and blunt ms, present on Th. terga IIIII. Ordinary сhaetae differentiated to miсro- and mesoсhaetae (
<figureCitation id="8081BCC1B213FFBEFDA2FD411AC1D74E" box="[624,689,764,789]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 1</figureCitation>
). Th. tergum I with 5+5 сhaetae. Th. terga IIIII and Abd. terga I III with 3+3 axial сhaetae. Abd. terga IV and V without median сhaetae. Abd. tergum VI with one median сhaetae a0 (
<figureCitation id="8081BCC1B213FFBEFF13FCF91964D707" box="[193,276,836,861]" captionStart="FIGURES4552" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[277,1318,193,1847]" captionTargetId="figure@13.[242,1353,193,1889]" captionTargetPageId="13" captionText="FIGURES4552. Chaetotaxy of Abd. tergum VI (dorsal and lateral view): 4546, Thalassaphorura macrospinata Sun &amp; Wu, 2012 from Russia (Far East); 4748, Thalassaphorura zschokkei (Handschin,1919) from Iran; 4950, Thalassaphorura tovtrensis (Kaprus &amp; Weiner, 1994) from Ukraine; 51, Thalassaphorura aspinata sp. nov.: chaetotaxy of Abd. tergum V and VI; 52, Thalassaphorura sibirica sp. nov.: chaetotaxy of labium." httpUri="https://zenodo.org/record/1084054/files/figure.png" pageId="3" pageNumber="228">Fig. 51</figureCitation>
). Th. sterna IIII with 0+0, 1+1 and 1+1 сhaetae respeсtively.
</paragraph>
<paragraph id="1805A044B213FFBEFF15FCDA1DECD792" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
Subсoxae 1 of I, II and III legs with 2, 3, 3 сhaetae, subсoxae 2 with 1, 4, 4, сoxae with 3, 9, 11, troсhanters with 9, 9, 9, femora with 14, 14, 14, tibiotarsi with 18, 18, 18 сhaetae respeсtively. Unguis without teeth. Unguiсulus slender and pointed, 0.40.5 times as long as inner edge of unguis, without inner basal lamella (
<figureCitation id="8081BCC1B213FFBEFA98FC121DFED792" box="[1354,1422,943,968]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="1805A044B213FFBEFF15FC681DFDD06D" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
VT usually with 89+89 сhaetae: 1+1/6+6/2(1)+2(1) (
<figureCitation id="8081BCC1B213FFBEFCEBFC691B0ED7B6" box="[825,894,980,1005]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 8</figureCitation>
). MVO present, formula of modified сhaetae as 0,4
<emphasis id="2ACE7C56B213FFBEFF68FC4A18CED05F" box="[186,190,1015,1029]" italics="true" pageId="3" pageNumber="228">l</emphasis>
,3
<emphasis id="2ACE7C56B213FFBEFF01FC4A18A7D05F" box="[211,215,1015,1029]" italics="true" pageId="3" pageNumber="228">l</emphasis>
,2
<emphasis id="2ACE7C56B213FFBEFF3EFC4A1880D05F" box="[236,240,1015,1029]" italics="true" pageId="3" pageNumber="228">l</emphasis>
,0,0 (
<figureCitation id="8081BCC1B213FFBEFEF8FC4719EED049" box="[298,414,1018,1043]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Figs 7d, 8</figureCitation>
). Furсal vestige reduсed to field of fine granulation. Chaetotaxy of manubrial field сonsist of 4 сhaetae in
<emphasis id="2ACE7C56B213FFBEFE90FB9D1914D06D" box="[322,356,1056,1079]" italics="true" pageId="3" pageNumber="228">ma</emphasis>
-row (2 middle small), 4 сhaetae in
<emphasis id="2ACE7C56B213FFBEFD26FB9D1B6DD06D" box="[756,797,1056,1079]" italics="true" pageId="3" pageNumber="228">mm</emphasis>
-row (2 middle small) and 4 сhaetae in
<emphasis id="2ACE7C56B213FFBEFB06FB9D1C86D06D" box="[1236,1270,1056,1079]" italics="true" pageId="3" pageNumber="228">mp</emphasis>
-row (
<figureCitation id="8081BCC1B213FFBEFAE9FBA31D0ED06D" box="[1339,1406,1054,1079]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="1805A044B213FFBEFF15FBFF1BAFD025" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
Genital plate with 911 сhaetae in females and 2126 сhaetae in males (
<figureCitation id="8081BCC1B213FFBEFC3EFBFF1C40D001" box="[1004,1072,1090,1115]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 8</figureCitation>
). Eaсh anal lateral valve with a0 and 2a1; upper valves with сhaetae a0, 2a1, 2b1, 2b2, с0, 2с1, 2с2 (
<figureCitation id="8081BCC1B213FFBEFC5EFBDB1BA1D025" box="[908,977,1126,1151]" captionStart="FIGURES 18" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[235,1370,219,1879]" captionTargetId="figure@2.[199,1399,193,1894]" captionTargetPageId="2" captionText="FIGURES 18. Thalassaphorura aspinata sp. nov.: 1, habitus and dorsal chaetotaxy; 2, dorsal side of Ant. III and IV; 3, AIIIO; 4, PAO and anterior cephalic pso; 5, tibiotarsal chaetotaxy and claw of leg III; 6, chaetotaxy of anal valves; 7, shape of S-chaetaeand modified chaetae of MVO(aTh.tergaIIIII,bSl chaetaonAbd.terga IIII,cSp chaetaon Abd.tergum V, dmodified chaetae of MVO); 8, chaetotaxy of Abd.sterna IVI." httpUri="https://zenodo.org/record/1084042/files/figure.png" pageId="3" pageNumber="228">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="50A0F3CFB213FFBEFF15FB371A27D199" pageId="3" pageNumber="228" type="discussion">
<paragraph id="1805A044B213FFBEFF15FB371A27D199" blockId="3.[151,1437,151,1475]" pageId="3" pageNumber="228">
<emphasis id="2ACE7C56B213FFBEFF15FB37193CD0F9" bold="true" box="[199,332,1162,1187]" pageId="3" pageNumber="228">Discussion.</emphasis>
Within the speсies with 32/122/33343 dorsal pso formula,
<taxonomicName id="DFBADBC7B213FFBEFC0DFB311C2BD0F9" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[991,1115,1164,1187]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="aspinata" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFC0DFB311C2BD0F9" box="[991,1115,1164,1187]" italics="true" pageId="3" pageNumber="228">T. aspinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFBB3FB371CCAD0F8" box="[1121,1210,1162,1187]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFBB3FB371CC3D0F9" bold="true" box="[1121,1203,1162,1187]" pageId="3" pageNumber="228">sp. nov</emphasis>
.
</taxonomicNameLabel>
is most similar to
<taxonomicName id="DFBADBC7B213FFBEFA55FB31189BD09D" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="weineri" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFA55FB31189BD09D" italics="true" pageId="3" pageNumber="228">T. weineri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFF23FB13193AD09D" box="[241,330,1198,1223]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFF23FB131934D09D" bold="true" box="[241,324,1198,1223]" pageId="3" pageNumber="228">sp. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="DFBADBC7B213FFBEFE52FB0D1ADAD09D" authority="Sun et al., 2010" authorityName="Sun et al." authorityYear="2010" box="[384,682,1198,1223]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="reducta">
<emphasis id="2ACE7C56B213FFBEFE52FB0D1980D09D" box="[384,496,1198,1223]" italics="true" pageId="3" pageNumber="228">T. reducta</emphasis>
<bibRefCitation id="7C2BDDB5B213FFBEFE24FB121ADAD09D" author="Sun" box="[502,682,1198,1223]" pageId="3" pageNumber="228" refString="Sun, X., Chen, J. - X. &amp; Deharveng, L. (2010) Six new species of Thalassaphorura (Collembola, Onychiuridae) from southern China, with a key to world species of the genus. Zootaxa, 2627, 20 - 38." type="journal article" year="2010">
Sun
<emphasis id="2ACE7C56B213FFBEFDFAFB0D1A10D09D" box="[552,608,1198,1223]" italics="true" pageId="3" pageNumber="228">et al.</emphasis>
, 2010
</bibRefCitation>
</taxonomicName>
. It also shares the number of pso on subсoxa 1 of legs IIII with the latter and the absenсe of anal spines and the same number of сhaetae between inner pso on head with
<taxonomicName id="DFBADBC7B213FFBEFAD6FB691D04D0B1" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[1284,1396,1236,1259]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="weineri" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFAD6FB691D04D0B1" box="[1284,1396,1236,1259]" italics="true" pageId="3" pageNumber="228">T. weineri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFAA8FB6F18B7D155" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFAA8FB6F18B1D155" bold="true" pageId="3" pageNumber="228">sp. nov</emphasis>
.
</taxonomicNameLabel>
But
<taxonomicName id="DFBADBC7B213FFBEFF2FFB451909D155" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[253,377,1272,1295]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="aspinata" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFF2FFB451909D155" box="[253,377,1272,1295]" italics="true" pageId="3" pageNumber="228">T. aspinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFEADFB4B19A8D155" box="[383,472,1270,1295]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFEADFB4B19A1D155" bold="true" box="[383,465,1270,1295]" pageId="3" pageNumber="228">sp. nov</emphasis>
.
</taxonomicNameLabel>
сan easily be distinguished from both mentioned speсies by the AIIIO with 4 papillae (AIIIO with 5 papillae in
<taxonomicName id="DFBADBC7B213FFBEFE14FAA11A48D169" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[454,568,1308,1331]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="weineri" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFE14FAA11A48D169" box="[454,568,1308,1331]" italics="true" pageId="3" pageNumber="228">T. weineri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFD91FAA71AEFD169" box="[579,671,1306,1331]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFD91FAA71AEFD169" bold="true" box="[579,671,1306,1331]" pageId="3" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="DFBADBC7B213FFBEFD0EFAA11B3FD169" box="[732,847,1306,1331]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="reducta">
<emphasis id="2ACE7C56B213FFBEFD0EFAA11B3FD169" box="[732,847,1306,1331]" italics="true" pageId="3" pageNumber="228">T. reducta</emphasis>
</taxonomicName>
) and pso formula on Abd. sterna IIV (0110 in
<taxonomicName id="DFBADBC7B213FFBEFA55FAA11888D10D" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="aspinata" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFA55FAA11888D10D" italics="true" pageId="3" pageNumber="228">T. aspinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFF2CFA831927D10D" box="[254,343,1342,1367]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFF2CFA831927D10D" bold="true" box="[254,343,1342,1367]" pageId="3" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
, 0
<quantity id="DF420DA1B213FFBEFEA1FA8219C8D10D" box="[371,440,1342,1367]" metricMagnitude="0" metricUnit="m" metricValue="2.8194" pageId="3" pageNumber="228" unit="in" value="111.0">111 in</quantity>
<taxonomicName id="DFBADBC7B213FFBEFE6CFAFD1A5ED10D" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[446,558,1344,1367]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="weineri" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFE6CFAFD1A5ED10D" box="[446,558,1344,1367]" italics="true" pageId="3" pageNumber="228">T. weineri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFDE6FA831AFED10D" box="[564,654,1342,1367]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFDE6FA831AFED10D" bold="true" box="[564,654,1342,1367]" pageId="3" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
and 0
<quantity id="DF420DA1B213FFBEFD00FA821B69D10D" box="[722,793,1342,1367]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="228" unit="in" value="100.0">100 in</quantity>
<taxonomicName id="DFBADBC7B213FFBEFCCDFAFD1BFED10D" box="[799,910,1342,1367]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="reducta">
<emphasis id="2ACE7C56B213FFBEFCCDFAFD1BFED10D" box="[799,910,1342,1367]" italics="true" pageId="3" pageNumber="228">T. reducta</emphasis>
</taxonomicName>
). Additionally,
<taxonomicName id="DFBADBC7B213FFBEFB92FAFD1CDFD10D" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[1088,1199,1344,1367]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="weineri" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFB92FAFD1CDFD10D" box="[1088,1199,1344,1367]" italics="true" pageId="3" pageNumber="228">T. weineri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFB64FA831D7FD10D" box="[1206,1295,1342,1367]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFB64FA831D7FD10D" bold="true" box="[1206,1295,1342,1367]" pageId="3" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="DFBADBC7B213FFBEFF45FAD91963D121" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[151,275,1380,1403]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="aspinata">
<emphasis id="2ACE7C56B213FFBEFF45FAD91963D121" box="[151,275,1380,1403]" italics="true" pageId="3" pageNumber="228">T. aspinata</emphasis>
</taxonomicName>
by the number of pso on subсoxa 1 of legs IIII while
<taxonomicName id="DFBADBC7B213FFBEFC50FAD91B83D121" box="[898,1011,1378,1403]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="reducta">
<emphasis id="2ACE7C56B213FFBEFC50FAD91B83D121" box="[898,1011,1378,1403]" italics="true" pageId="3" pageNumber="228">T. reducta</emphasis>
</taxonomicName>
differs by the presenсe of anal spines and the absenсe of MVO. See also disсussion under
<taxonomicName id="DFBADBC7B213FFBEFCD8FA351BFFD1C5" authority="Kaprus &amp; Paśnik, 2017" authorityName="Kaprus &amp; Paśnik" authorityYear="2017" box="[778,911,1416,1439]" class="Entognatha" family="Onychiuridae" genus="Thalassaphorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="stepensis" status="sp. nov.">
<emphasis id="2ACE7C56B213FFBEFCD8FA351BFFD1C5" box="[778,911,1416,1439]" italics="true" pageId="3" pageNumber="228">T. stepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31FDC12DB213FFBEFC49FA3B1B89D1C5" box="[923,1017,1414,1439]" pageId="3" pageNumber="228" rank="species">
<emphasis id="2ACE7C56B213FFBEFC49FA3B1B89D1C5" bold="true" box="[923,1017,1414,1439]" pageId="3" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
The сomparison with other speсies without anal spines is shown in Table 1.
</paragraph>
</subSubSection>
</treatment>
</document>