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<document id="23EB3AB1C9F1DBEE4652ABDFA7AA99EA" ID-DOI="10.11646/zootaxa.5311.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8089905" ID-ZooBank="0E28EEF2-38B4-412F-80C1-CC958A9B7B26" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1687949212707" checkinUser="plazi" docAuthor="Poorani, J." docDate="2023" docId="8A2A223EEA70FFBF02D67400B15EFCCB" docLanguage="en" docName="zootaxa.5311.1.1.pdf" docOrigin="Zootaxa 5311 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5311.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Novius octoguttatus" docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="76135A46EA69FFA402417269B37AFF96" masterDocTitle="A review of the tribe Noviini (Coleoptera: Coccinellidae) of the Indian subcontinent" masterLastPageNumber="47" masterPageNumber="1" pageNumber="26" updateTime="1687949915969" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B690843DDB1B78DF8D6026BD75CBA250" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9070898BD4BB4FFF6006D50C563AAABB">A review of the tribe Noviini (Coleoptera: Coccinellidae) of the Indian subcontinent</mods:title>
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<mods:title id="CE67B9573EB26AC5C6EBFCE6B143B1B1">Zootaxa</mods:title>
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<mods:date id="4FD24E1EAD55D506853B9C4FD4CE11A6">2023</mods:date>
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<mods:classification id="301FBE17B7899DE9474189B71833C78B">journal article</mods:classification>
<mods:identifier id="DE9C62512756D9797AAB542D83A6C23C" type="DOI">10.11646/zootaxa.5311.1.1</mods:identifier>
<mods:identifier id="1F897B3C00465751BCA2114B9BF60569" type="ISSN">1175-5326</mods:identifier>
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<treatment id="8A2A223EEA70FFBF02D67400B15EFCCB" ID-DOI="http://doi.org/10.5281/zenodo.8089955" ID-Zenodo-Dep="8089955" LSID="urn:lsid:plazi:treatment:8A2A223EEA70FFBF02D67400B15EFCCB" httpUri="http://treatment.plazi.org/id/8A2A223EEA70FFBF02D67400B15EFCCB" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="4A99C0A3EA70FFBD02D67400B296F915" box="[151,492,1641,1668]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="023C9328EA70FFBD02D67400B296F915" blockId="25.[151,492,1641,1703]" box="[151,492,1641,1668]" pageId="25" pageNumber="26">
<heading id="59742444EA70FFBD02D67400B296F915" bold="true" box="[151,492,1641,1668]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="C583E8ABEA70FFBD02D67400B296F915" authority="(Weise)" baseAuthorityName="Weise" box="[151,492,1641,1668]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="30F74F3AEA70FFBD02D67400B296F915" bold="true" box="[151,492,1641,1668]" pageId="25" pageNumber="26">
<emphasis id="30F74F3AEA70FFBD02D67400B2FDF912" bold="true" box="[151,391,1641,1668]" italics="true" pageId="25" pageNumber="26">Novius octoguttatus</emphasis>
(Weise)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4A99C0A3EA70FFBD02D674E4B380F931" box="[151,250,1677,1703]" pageId="25" pageNumber="26" type="description">
<paragraph id="023C9328EA70FFBD02D674E4B380F931" blockId="25.[151,492,1641,1703]" box="[151,250,1677,1703]" pageId="25" pageNumber="26">
(
<figureCitation id="9AB88FADEA70FFBD02DE74E4B38BF931" box="[159,241,1677,1703]" captionStart="FIGURE 19" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[288,1298,181,1918]" captionTargetId="figure-21@26.[288,1299,181,1919]" captionTargetPageId="26" captionText="FIGURE 19. Novius octoguttatus (Weise):a.adult, dorsal view (BMNH); e. adult(from Rajasthan, India); f.abdominal postcoxal line; g–i. male genitalia: g. tegmen, lateral view; h. tegmen, inner view; i. penis; b–d. Novius quadrimaculatus (Mader): b. adult, dorsal view; c. adult, lateral view; d. adult, frontal view (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.8089969" httpUri="https://zenodo.org/record/8089969/files/figure.png" pageId="25" pageNumber="26">Fig. 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4A99C0A3EA70FFBD02D674BBB13BF89F" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="023C9328EA70FFBD02D674BBB168F97C" blockId="25.[151,577,1746,1801]" box="[151,530,1746,1770]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="2293B406EA70FFBD02D674BBB168F97C" box="[151,530,1746,1770]" pageId="25" pageNumber="26">
<treatmentCitation id="8322B539EA70FFBD02D674BBB177F97C" author="Weise, J." box="[151,525,1746,1770]" page="51" pageId="25" pageNumber="26" year="1910">
<taxonomicName id="C583E8ABEA70FFBD02D674BBB177F97C" authority="Weise, 1910: 51" authorityName="Weise" authorityPageNumber="51" authorityYear="1910" box="[151,525,1746,1770]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="octoguttata">
<emphasis id="30F74F3AEA70FFBD02D674BBB21BF97C" box="[151,353,1746,1770]" italics="true" pageId="25" pageNumber="26">Rodolia octoguttata</emphasis>
<bibRefCitation id="6612EED9EA70FFBD032674BBB177F97C" author="Weise, J." box="[359,525,1746,1770]" pageId="25" pageNumber="26" pagination="25 - 53" refId="ref16948" refString="Weise, J. (1910) Chrysomeliden und Coccinelliden. Brunn Verhandlungen des Naturforschenden Vereines, 48 (1909), 25 - 53." type="journal article" year="1910">Weise, 1910: 51</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="023C9328EA70FFBD02D67498B13BF89F" blockId="25.[151,577,1746,1801]" box="[151,577,1776,1801]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="2293B406EA70FFBD02D67498B13BF89F" box="[151,577,1776,1801]" pageId="25" pageNumber="26">
<treatmentCitation id="8322B539EA70FFBD02D67498B147F89F" ID-DOI="http://doi.org/10.5281/zenodo.3796065" ID-Zenodo-Dep="3796065" author="Pang, H. &amp; Tang, X. F. &amp; Booth, R. G. &amp; Vandenberg, N. &amp; Forrester, J. &amp; McHugh, J. &amp; Slipinski, A." box="[151,573,1776,1801]" httpUri="http://treatment.plazi.org/id/03EEFD2AFFB1FF92FF6F1A3AD6C538FB" page="20" pageId="25" pageNumber="26" year="2020">
<taxonomicName id="C583E8ABEA70FFBD02D67498B147F89F" authority=": Pang et al. 2020: 20" authorityName="Pang" authorityPageNumber="20" authorityYear="2020" box="[151,573,1776,1801]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="30F74F3AEA70FFBD02D67498B21BF89E" box="[151,353,1777,1800]" italics="true" pageId="25" pageNumber="26">Novius octoguttatus</emphasis>
:
<bibRefCitation id="6612EED9EA70FFBD032E7498B147F89F" author="Pang, H. &amp; Tang, X. F. &amp; Booth, R. G. &amp; Vandenberg, N. &amp; Forrester, J. &amp; McHugh, J. &amp; Slipinski, A." box="[367,573,1776,1801]" pageId="25" pageNumber="26" pagination="1 - 24" refId="ref15453" refString="Pang, H., Tang, X. F., Booth, R. G., Vandenberg, N., Forrester, J., McHugh, J. &amp; Slipinski, A. (2020) Revision of the Australian Coccinellidae (Coleoptera). Genus Novius Mulsant of tribe Noviini. Annales Zoologici, 70 (1), 1 - 24. https: // doi. org / 10.3161 / 00034541 ANZ 2020.70.1.001" type="journal article" year="2020">
Pang
<emphasis id="30F74F3AEA70FFBD03E97498B2AEF89E" box="[424,468,1776,1800]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2020: 20
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="56FCC3A0EA73FFBE02D675FEB04EF861" ID-DOI="http://doi.org/10.5281/zenodo.8089969" ID-Zenodo-Dep="8089969" httpUri="https://zenodo.org/record/8089969/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1943,1967]" targetBox="[288,1298,181,1918]" targetPageId="26" targetType="figure">
<paragraph id="023C9328EA73FFBE02D675FEB04EF861" blockId="26.[151,1436,1943,2039]" pageId="26" pageNumber="27">
<emphasis id="30F74F3AEA73FFBE02D675FEB264F839" bold="true" box="[151,286,1943,1967]" pageId="26" pageNumber="27">FIGURE 19.</emphasis>
<taxonomicName id="C583E8ABEA73FFBE036375F1B145F839" authority="(Weise)" baseAuthorityName="Weise" box="[290,575,1943,1967]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="30F74F3AEA73FFBE036375F1B293F839" box="[290,489,1944,1967]" italics="true" pageId="26" pageNumber="27">Novius octoguttatus</emphasis>
(Weise)
</taxonomicName>
: a. adult, dorsal view (BMNH); e. adult (from Rajasthan, India); f. abdominal postcoxal line; g–i. male genitalia: g. tegmen, lateral view; h. tegmen, inner view; i. penis; b–d.
<taxonomicName id="C583E8ABEA73FFBE01AF75D5B638F845" authority="(Mader)" baseAuthorityName="Mader" box="[1006,1346,1979,2003]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="30F74F3AEA73FFBE01AF75D5B79CF845" box="[1006,1254,1979,2003]" italics="true" pageId="26" pageNumber="27">Novius quadrimaculatus</emphasis>
(Mader)
</taxonomicName>
: b. adult, dorsal view; c. adult, lateral view; d. adult, frontal view (BMNH).
</paragraph>
</caption>
<subSubSection id="4A99C0A3EA72FFBF02D672FFB753FED7" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="023C9328EA72FFBF02D672FFB753FED7" blockId="27.[151,1437,150,861]" pageId="27" pageNumber="28">
<emphasis id="30F74F3AEA72FFBF02D672FFB268FF26" bold="true" box="[151,274,150,176]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Length: 3.50–5.00 mm; width:
<quantity id="C57B3ECDEA72FFBF003072FEB06CFF27" box="[625,790,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.6" metricValueMin="3.2" pageId="27" pageNumber="28" unit="mm" value="3.9" valueMax="4.6" valueMin="3.2">3.20–4.60 mm</quantity>
. Form (
<figureCitation id="9AB88FADEA72FFBF012B72FEB0A5FF27" box="[874,991,151,177]" captionStart="FIGURE 19" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[288,1298,181,1918]" captionTargetId="figure-21@26.[288,1299,181,1919]" captionTargetPageId="26" captionText="FIGURE 19. Novius octoguttatus (Weise):a.adult, dorsal view (BMNH); e. adult(from Rajasthan, India); f.abdominal postcoxal line; g–i. male genitalia: g. tegmen, lateral view; h. tegmen, inner view; i. penis; b–d. Novius quadrimaculatus (Mader): b. adult, dorsal view; c. adult, lateral view; d. adult, frontal view (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.8089969" httpUri="https://zenodo.org/record/8089969/files/figure.png" pageId="27" pageNumber="28">Fig. 19a, e</figureCitation>
) elongate oval, dorsum strongly convex and densely pubescent. Ground colour dark reddish brown or rusty reddish brown; pronotum with two black spots on either side of midline above posterior margin, occasionally obsolete or absent; nominate form (
<figureCitation id="9AB88FADEA72FFBF06A272B6B621FF6F" box="[1251,1371,223,249]" captionStart="FIGURE 19" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[288,1298,181,1918]" captionTargetId="figure-21@26.[288,1299,181,1919]" captionTargetPageId="26" captionText="FIGURE 19. Novius octoguttatus (Weise):a.adult, dorsal view (BMNH); e. adult(from Rajasthan, India); f.abdominal postcoxal line; g–i. male genitalia: g. tegmen, lateral view; h. tegmen, inner view; i. penis; b–d. Novius quadrimaculatus (Mader): b. adult, dorsal view; c. adult, lateral view; d. adult, frontal view (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.8089969" httpUri="https://zenodo.org/record/8089969/files/figure.png" pageId="27" pageNumber="28">Fig. 19a, e</figureCitation>
) with each elytron having four black spots arranged in a 1-1-1-1 pattern, elytral spots often much enlarged; ventral side uniformly rusty red to yellowish brown. Male genitalia (
<figureCitation id="9AB88FADEA72FFBF014F734EB0FEFED7" box="[782,900,295,321]" captionStart="FIGURE 19" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[288,1298,181,1918]" captionTargetId="figure-21@26.[288,1299,181,1919]" captionTargetPageId="26" captionText="FIGURE 19. Novius octoguttatus (Weise):a.adult, dorsal view (BMNH); e. adult(from Rajasthan, India); f.abdominal postcoxal line; g–i. male genitalia: g. tegmen, lateral view; h. tegmen, inner view; i. penis; b–d. Novius quadrimaculatus (Mader): b. adult, dorsal view; c. adult, lateral view; d. adult, frontal view (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.8089969" httpUri="https://zenodo.org/record/8089969/files/figure.png" pageId="27" pageNumber="28">
Fig.
<quantity id="C57B3ECDEA72FFBF0102734EB016FED7" box="[835,876,295,321]" metricMagnitude="-2" metricUnit="kg" metricValue="1.9" pageId="27" pageNumber="28" unit="g" value="19.0">19g</quantity>
–i
</figureCitation>
) as illustrated.
</paragraph>
</subSubSection>
<subSubSection id="4A99C0A3EA72FFBF02867323B04BFE47" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="023C9328EA72FFBF02867323B04BFE47" blockId="27.[151,1437,150,861]" pageId="27" pageNumber="28">
<emphasis id="30F74F3AEA72FFBF02867323B2C8FEF2" bold="true" box="[199,434,330,356]" pageId="27" pageNumber="28">Material examined.</emphasis>
N.
<materialsCitation id="B2EB9975EA72FFBF039F7322B03CFEF3" box="[478,838,331,357]" collectorName="W. Almora &amp; Kumaon" country="India" location="India" pageId="27" pageNumber="28" specimenCount="1">
<collectingCountry id="7A94D3B8EA72FFBF039F7322B160FEF3" box="[478,538,331,357]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
,
<collectorName id="AF76F6FEEA72FFBF00667322B1DFFEF3" box="[551,677,331,357]" pageId="27" pageNumber="28">W. Almora</collectorName>
,
<collectorName id="AF76F6FEEA72FFBF00F37322B06FFEF3" box="[690,789,331,357]" pageId="27" pageNumber="28">Kumaon</collectorName>
, UP
</materialsCitation>
,
<materialsCitation id="B2EB9975EA72FFBF01137322B009FE1F" collectionCode="BMNH" country="India" location="Champion Coll." pageId="27" pageNumber="28" specimenCount="1">
<collectingCountry id="7A94D3B8EA72FFBF01137322B0F4FEF3" box="[850,910,331,357]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
, H.G.C/G.C.
<location id="075CC5F3EA72FFBF066B7322B7A5FEF3" LSID="urn:lsid:plazi:treatment:8A2A223EEA70FFBF02D67400B15EFCCB:075CC5F3EA72FFBF066B7322B7A5FEF3" box="[1066,1247,331,357]" country="India" name="Champion Coll." pageId="27" pageNumber="28">Champion Coll.</location>
B.M. 1927-409/
<taxonomicName id="C583E8ABEA72FFBF02D67306B1BFFE1F" authority="Weise, Det. R. G. Booth 1991" authorityName="Weise, Det. R. G. Booth" authorityYear="1991" box="[151,709,367,393]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="octoguttata">Rodolia octoguttata Weise, Det. R.G. Booth 1991</taxonomicName>
, 1 ex (
<collectionCode id="64920BEDEA72FFBF01507306B011FE1F" box="[785,875,367,393]" pageId="27" pageNumber="28">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B2EB9975EA72FFBF013F7306B1D1FE3B" collectingDate="1918-07" collectionCode="BMNH" collectorName="GC Champion Coll. B. M." country="India" location="Almora Divn. Kumaon" pageId="27" pageNumber="28" specimenCount="1">
W.
<location id="075CC5F3EA72FFBF01E27306B7DCFE1F" LSID="urn:lsid:plazi:treatment:8A2A223EEA70FFBF02D67400B15EFCCB:075CC5F3EA72FFBF01E27306B7DCFE1F" box="[931,1190,367,393]" country="India" name="Almora Divn. Kumaon" pageId="27" pageNumber="28">Almora Divn. Kumaon</location>
UP,
<date id="763DB5E8EA72FFBF069C7306B634FE1F" box="[1245,1358,367,393]" pageId="27" pageNumber="28" value="1918-07">
<collectingDate id="66794C00EA72FFBF069C7306B634FE1F" box="[1245,1358,367,393]" pageId="27" pageNumber="28" value="1918-07">July 1918</collectingDate>
</date>
, HGC/
<collectorName id="AF76F6FEEA72FFBF02D673FAB2B8FE3B" box="[151,450,403,429]" pageId="27" pageNumber="28">GC Champion Coll. B.M.</collectorName>
1927-401 (
<collectionCode id="64920BEDEA72FFBF000873FAB1D9FE3B" box="[585,675,403,429]" pageId="27" pageNumber="28">BMNH</collectionCode>
)
</materialsCitation>
; W.
<materialsCitation id="B2EB9975EA72FFBF00A073FAB057FE47" collectingDate="1988-03-25" collectingDateMax="1988-03-27" collectingDateMin="1988-03-25" collectionCode="BMNH" collectorName="Farmland &amp; Kwai, M. J. D &amp; Brendell" country="Thailand" elevation="400" location="Kanchanaburi" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Kanchanaburi">
<collectingCountry id="7A94D3B8EA72FFBF00A073FAB03DFE3B" box="[737,839,403,429]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
,
<collectingRegion id="C0475DCAEA72FFBF011773FAB08CFE3B" box="[854,1014,403,429]" country="Thailand" name="Kanchanaburi" pageId="27" pageNumber="28">Kanchanaburi</collectingRegion>
,
<quantity id="C57B3ECDEA72FFBF064573FAB72AFE3B" box="[1028,1104,403,429]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="400.0">
<elevation id="89AE741BEA72FFBF064573FAB72AFE3B" box="[1028,1104,403,429]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="763DB5E8EA72FFBF061E73FBB781FE3B" box="[1119,1275,402,429]" pageId="27" pageNumber="28" value="1988-03-25" valueMax="1988-03-27" valueMin="1988-03-25">
<collectingDate id="66794C00EA72FFBF061E73FBB781FE3B" box="[1119,1275,402,429]" pageId="27" pageNumber="28" value="1988-03-25" valueMax="1988-03-27" valueMin="1988-03-25">25–27.iii.1988</collectingDate>
</date>
/
<collectorName id="AF76F6FEEA72FFBF074573FBB60CFE3B" box="[1284,1398,402,429]" pageId="27" pageNumber="28">Farmland</collectorName>
nr. river
<collectorName id="AF76F6FEEA72FFBF029273DEB21DFE47" box="[211,359,439,465]" pageId="27" pageNumber="28">Kwai, M.J.D</collectorName>
.
<collectorName id="AF76F6FEEA72FFBF033373DEB2AFFE47" box="[370,469,439,465]" pageId="27" pageNumber="28">Brendell</collectorName>
, BM 1988-183, 1 ex (
<collectionCode id="64920BEDEA72FFBF008B73DEB05EFE47" box="[714,804,439,465]" pageId="27" pageNumber="28">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4A99C0A3EA72FFBF028673B3B15EFCCB" pageId="27" pageNumber="28" type="distribution">
<paragraph id="023C9328EA72FFBF028673B3B61BFE63" blockId="27.[151,1437,150,861]" box="[199,1377,474,501]" pageId="27" pageNumber="28">
<emphasis id="30F74F3AEA72FFBF028673B3B21BFE62" bold="true" box="[199,353,474,500]" pageId="27" pageNumber="28">Distribution.</emphasis>
<collectingCountry id="7A94D3B8EA72FFBF032673B2B2DBFE63" box="[359,417,475,501]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
(
<collectingRegion id="C0475DCAEA72FFBF03F173B2B14BFE63" box="[432,561,475,501]" country="India" name="Meghalaya" pageId="27" pageNumber="28">Meghalaya</collectingRegion>
,
<collectingRegion id="C0475DCAEA72FFBF007D73B2B1B3FE63" box="[572,713,475,501]" country="India" name="Uttarakhand" pageId="27" pageNumber="28">Uttarakhand</collectingRegion>
,
<collectingRegion id="C0475DCAEA72FFBF009573B2B032FE63" box="[724,840,475,501]" country="India" name="Rajasthan" pageId="27" pageNumber="28">Rajasthan</collectingRegion>
);
<collectingCountry id="7A94D3B8EA72FFBF011A73B2B0B6FE63" box="[859,972,475,501]" name="Sri Lanka" pageId="27" pageNumber="28">Sri Lanka</collectingCountry>
;
<collectingCountry id="7A94D3B8EA72FFBF019673B2B767FE63" box="[983,1053,475,501]" name="Nepal" pageId="27" pageNumber="28">Nepal</collectingCountry>
;
<collectingCountry id="7A94D3B8EA72FFBF066873B2B7E0FE63" box="[1065,1178,475,501]" name="Myanmar" pageId="27" pageNumber="28">Myanmar</collectingCountry>
;
<collectingCountry id="7A94D3B8EA72FFBF06E573B2B671FE63" box="[1188,1291,475,501]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
;
<collectingCountry id="7A94D3B8EA72FFBF075773B2B627FE63" box="[1302,1373,475,501]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
.
</paragraph>
<paragraph id="023C9328EA72FFBF02867397B226FDF7" blockId="27.[151,1437,150,861]" pageId="27" pageNumber="28">
<emphasis id="30F74F3AEA72FFBF02867397B29DFD8E" bold="true" box="[199,487,510,536]" pageId="27" pageNumber="28">Prey/associated habitat.</emphasis>
<taxonomicName id="C583E8ABEA72FFBF03B37396B109FD8F" box="[498,627,511,537]" class="Insecta" family="Margarodidae" genus="Icerya" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="30F74F3AEA72FFBF03B37396B140FD8E" box="[498,570,511,536]" italics="true" pageId="27" pageNumber="28">Icerya</emphasis>
spp.
</taxonomicName>
<bibRefCitation id="6612EED9EA72FFBF003F7396B02EFD8F" author="Iqbal, Z. &amp; Nasir, M. F. &amp; Bodlah, I. &amp; Qureshi, R." box="[638,852,511,537]" pageId="27" pageNumber="28" pagination="1103 - 1111" refId="ref14578" refString="Iqbal, Z., Nasir, M. F., Bodlah, I. &amp; Qureshi, R. (2018) A Contribution to the Genus Rodolia Mulsant, 1850 (Coleoptera: Coccinellidae) from Pothwar Plateau of Pakistan. The Journal of Animal &amp; Plant Sciences, 28 (4), 1103 - 1111." type="journal article" year="2018">
Iqbal
<emphasis id="30F74F3AEA72FFBF00837396B18DFD8F" box="[706,759,511,537]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. (2018)
</bibRefCitation>
recorded it as a predator of
<taxonomicName id="C583E8ABEA72FFBF06EA7396B6E7FD8F" box="[1195,1437,511,537]" class="Insecta" family="Margarodidae" genus="Drosicha" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mangiferae">
<emphasis id="30F74F3AEA72FFBF06EA7396B6E7FD8F" box="[1195,1437,511,537]" italics="true" pageId="27" pageNumber="28">Drosicha mangiferae</emphasis>
</taxonomicName>
and the host plants were
<taxonomicName id="C583E8ABEA72FFBF0381704AB1D2FDAA" box="[448,680,547,573]" class="Magnoliopsida" family="Rosaceae" genus="Eriobotrya" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis id="30F74F3AEA72FFBF0381704AB1D2FDAA" box="[448,680,547,573]" italics="true" pageId="27" pageNumber="28">Eriobotrya japonica</emphasis>
</taxonomicName>
,
<taxonomicName id="C583E8ABEA72FFBF00FB704AB0F3FDAB" box="[698,905,547,573]" class="Magnoliopsida" family="Apocynaceae" genus="Alstonia" kingdom="Plantae" order="Gentianales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="scholaris">
<emphasis id="30F74F3AEA72FFBF00FB704AB0F3FDAB" box="[698,905,547,573]" italics="true" pageId="27" pageNumber="28">Alstonia scholaris</emphasis>
</taxonomicName>
,
<taxonomicName id="C583E8ABEA72FFBF01DA704AB7EAFDAB" box="[923,1168,547,573]" class="Magnoliopsida" family="Lythraceae" genus="Lagerstroemia" kingdom="Plantae" order="Myrtales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="30F74F3AEA72FFBF01DA704AB7EAFDAB" box="[923,1168,547,573]" italics="true" pageId="27" pageNumber="28">Lagerstroemia indica</emphasis>
</taxonomicName>
,
<taxonomicName id="C583E8ABEA72FFBF06E3704AB613FDAB" box="[1186,1385,547,573]" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" kingdom="Plantae" order="Sapindales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="30F74F3AEA72FFBF06E3704AB613FDAB" box="[1186,1385,547,573]" italics="true" pageId="27" pageNumber="28">Mangifera indica</emphasis>
</taxonomicName>
and
<taxonomicName id="C583E8ABEA72FFBF02D6702EB22EFDF6" box="[151,340,583,609]" class="Magnoliopsida" family="Fabaceae" genus="Dalbergia" kingdom="Plantae" order="Fabales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="sissoo">
<emphasis id="30F74F3AEA72FFBF02D6702EB22EFDF6" box="[151,340,583,609]" italics="true" pageId="27" pageNumber="28">Dalbergia sissoo</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="023C9328EA72FFBF02867003B755FD13" blockId="27.[151,1437,150,861]" box="[199,1071,618,645]" pageId="27" pageNumber="28">
<emphasis id="30F74F3AEA72FFBF02867003B2BBFD13" bold="true" box="[199,449,618,645]" pageId="27" pageNumber="28">Seasonal occurrence.</emphasis>
April–June and October in
<collectingCountry id="7A94D3B8EA72FFBF00B67002B02CFD13" box="[759,854,619,645]" name="Pakistan" pageId="27" pageNumber="28">Pakistan</collectingCountry>
(
<bibRefCitation id="6612EED9EA72FFBF01247002B759FD13" author="Iqbal, Z. &amp; Nasir, M. F. &amp; Bodlah, I. &amp; Qureshi, R." box="[869,1059,619,645]" pageId="27" pageNumber="28" pagination="1103 - 1111" refId="ref14578" refString="Iqbal, Z., Nasir, M. F., Bodlah, I. &amp; Qureshi, R. (2018) A Contribution to the Genus Rodolia Mulsant, 1850 (Coleoptera: Coccinellidae) from Pothwar Plateau of Pakistan. The Journal of Animal &amp; Plant Sciences, 28 (4), 1103 - 1111." type="journal article" year="2018">
Iqbal
<emphasis id="30F74F3AEA72FFBF01E67002B0A5FD13" box="[935,991,619,645]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
<paragraph id="023C9328EA72FFBF028670E6B15EFCCB" blockId="27.[151,1437,150,861]" pageId="27" pageNumber="28">
<emphasis id="30F74F3AEA72FFBF028670E6B26BFD3F" bold="true" box="[199,273,655,681]" pageId="27" pageNumber="28">Notes.</emphasis>
<bibRefCitation id="6612EED9EA72FFBF035970E6B2D5FD3F" author="Kapur, A. P." box="[280,431,654,681]" pageId="27" pageNumber="28" pagination="1 - 7" refId="ref14683" refString="Kapur, A. P. (1951) Further notes on the Indian species of Rodolia Mulsant (Coleoptera: Coccinellidae). Records of the Indian Museum, 48, 1 - 7. https: // doi. org / 10.26515 / rzsi / v 48 / i 2 / 1951 / 162144" type="journal article" year="1951">Kapur (1951)</bibRefCitation>
described it in detail and illustrated only truncated parts of the male and female genitalia. He mentioned that it belonged to the same group as
<taxonomicName id="C583E8ABEA72FFBF009270DAB031FD5B" authorityName="BMNH" baseAuthorityName="Mulsant" baseAuthorityYear="1850" box="[723,843,691,717]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fumidus">
<emphasis id="30F74F3AEA72FFBF009270DAB031FD5B" box="[723,843,691,717]" italics="true" pageId="27" pageNumber="28">N. fumidus</emphasis>
</taxonomicName>
,
<taxonomicName id="C583E8ABEA72FFBF011970DAB0BCFD5A" box="[856,966,691,716]" class="Insecta" family="Coccinellidae" genus="Rodolia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="30F74F3AEA72FFBF011970DAB0BCFD5A" box="[856,966,691,716]" italics="true" pageId="27" pageNumber="28">R. guerini</emphasis>
</taxonomicName>
, and
<taxonomicName id="C583E8ABEA72FFBF064070DAB7CEFD5B" box="[1025,1204,691,717]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andamanicus">
<emphasis id="30F74F3AEA72FFBF064070DAB7CEFD5B" box="[1025,1204,691,717]" italics="true" pageId="27" pageNumber="28">R. andamanicus</emphasis>
</taxonomicName>
by the general shape of the body, and the characteristic subapical projection of the penis guide. Bielawski (1972) illustrated the habitus and male genitalia of this species.
<taxonomicName id="C583E8ABEA72FFBF005D7092B7EAFC83" authority="(Mader, 1939)" baseAuthorityName="Mader" baseAuthorityYear="1939" box="[540,1168,763,789]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="variety" species="quadrimaculatus" variety="sexmaculatus">
<emphasis id="30F74F3AEA72FFBF005D7092B09DFC83" box="[540,999,763,789]" italics="true" pageId="27" pageNumber="28">Novius quadrimaculatus v. sexmaculatus</emphasis>
(
<bibRefCitation id="6612EED9EA72FFBF01B77092B7FDFC83" author="Mader, L." box="[1014,1159,763,789]" pageId="27" pageNumber="28" pagination="41 - 51" refId="ref15137" refString="Mader, L. (1939) Neue Coleopteren aus China. Entomologisches Nachrichtenblatt (Troppau), 13, 41 - 51." type="journal article" year="1939">Mader, 1939</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="9AB88FADEA72FFBF06E17092B78AFC83" box="[1184,1264,763,789]" captionStart="FIGURE 19" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[288,1298,181,1918]" captionTargetId="figure-21@26.[288,1299,181,1919]" captionTargetPageId="26" captionText="FIGURE 19. Novius octoguttatus (Weise):a.adult, dorsal view (BMNH); e. adult(from Rajasthan, India); f.abdominal postcoxal line; g–i. male genitalia: g. tegmen, lateral view; h. tegmen, inner view; i. penis; b–d. Novius quadrimaculatus (Mader): b. adult, dorsal view; c. adult, lateral view; d. adult, frontal view (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.8089969" httpUri="https://zenodo.org/record/8089969/files/figure.png" pageId="27" pageNumber="28">Fig. 19</figureCitation>
b-d, BMNH) is treated as a synonym of
<taxonomicName id="C583E8ABEA72FFBF03EF7176B120FCAE" box="[430,602,799,824]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="30F74F3AEA72FFBF03EF7176B120FCAE" box="[430,602,799,824]" italics="true" pageId="27" pageNumber="28">N. octoguttatus</emphasis>
</taxonomicName>
in
<bibRefCitation id="6612EED9EA72FFBF00C17176B038FCAF" author="Ren, S. X. &amp; Wang, X. M. &amp; Pang, H. &amp; Peng, Z. Q. &amp; Zeng, T." box="[640,834,799,825]" pageId="27" pageNumber="28" refId="ref15983" refString="Ren, S. X., Wang, X. M., Pang, H., Peng, Z. Q. &amp; Zeng, T. (2009) Colored Pictorial Handbook of Ladybird Beetles in China. Science Press, Beijing, 336 pp." type="book" year="2009">
Ren
<emphasis id="30F74F3AEA72FFBF00F47176B192FCAF" box="[693,744,799,825]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. (2009)
</bibRefCitation>
but listed as a valid species in the world checklist of
<taxonomicName id="C583E8ABEA72FFBF02D6712AB39FFCCA" authorityName="Mulsant" authorityYear="1846" box="[151,229,835,860]" class="Insecta" family="Coccinellidae" genus="Novius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="30F74F3AEA72FFBF02D6712AB39FFCCA" box="[151,229,835,860]" italics="true" pageId="27" pageNumber="28">Novius</emphasis>
</taxonomicName>
given by
<bibRefCitation id="6612EED9EA72FFBF0315712AB15AFCCB" author="Pang, H. &amp; Tang, X. F. &amp; Booth, R. G. &amp; Vandenberg, N. &amp; Forrester, J. &amp; McHugh, J. &amp; Slipinski, A." box="[340,544,835,861]" pageId="27" pageNumber="28" pagination="1 - 24" refId="ref15453" refString="Pang, H., Tang, X. F., Booth, R. G., Vandenberg, N., Forrester, J., McHugh, J. &amp; Slipinski, A. (2020) Revision of the Australian Coccinellidae (Coleoptera). Genus Novius Mulsant of tribe Noviini. Annales Zoologici, 70 (1), 1 - 24. https: // doi. org / 10.3161 / 00034541 ANZ 2020.70.1.001" type="journal article" year="2020">
Pang
<emphasis id="30F74F3AEA72FFBF03D2712AB2BEFCCB" box="[403,452,835,861]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. (2020)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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