157 lines
20 KiB
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157 lines
20 KiB
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<document ID-CLB-Dataset="32500" ID-DOI="http://dx.doi.org/10.3897/zookeys.688.12590" ID-GBIF-Dataset="5c79e002-b20e-4f1f-a5ea-24e6db9cf93d" ID-PMC="PMC5672582" ID-Pensoft-Pub="1313-2970-688-49" ID-Pensoft-UUID="4A72FF87A05DF179FFE8FF852A295E52" ID-PubMed="29118592" ID-Zenodo-Dep="3484841" ID-ZooBank="9FD88178828543D89A6E560287FE0199" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-688-49" ModsDocOrigin="ZooKeys 688" ModsDocTitle="A review of the genus Lordiphosa Basden in India, with descriptions of four new species from the Himalayan region (Diptera, Drosophilidae)" checkinTime="1502744385809" checkinUser="pensoft" docAuthor="Fartyal, Rajendra S., Sati, Pradeep C., Pradhan, Sushmika, Kandpal, Mukul C., Toda, Masanori J., Chatterjee, Rabindra N., Singh, Birendra K. & Bhardwai, Asha" docDate="2017" docId="87BACB7EBB0D9A160C5B0BB81C2542B1" docLanguage="en" docName="ZooKeys 688: 49-79" docOrigin="ZooKeys 688" docPubDate="2017-08-08" docSource="http://dx.doi.org/10.3897/zookeys.688.12590" docTitle="Lordiphosa Basden 1961" docType="treatment" docVersion="7" id="4A72FF87A05DF179FFE8FF852A295E52" lastPageNumber="49" masterDocId="4A72FF87A05DF179FFE8FF852A295E52" masterDocTitle="A review of the genus Lordiphosa Basden in India, with descriptions of four new species from the Himalayan region (Diptera, Drosophilidae)" masterLastPageNumber="79" masterPageNumber="49" pageNumber="49" updateTime="1705949741233" updateUser="pensoft">
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<mods:title id="19AF2E24D892BF065C1C810FCD19DB1B">A review of the genus Lordiphosa Basden in India, with descriptions of four new species from the Himalayan region (Diptera, Drosophilidae)</mods:title>
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<mods:namePart id="2ACF654B74EE32E2E762B6CB705FE943">Fartyal, Rajendra S.</mods:namePart>
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<mods:affiliation id="67DB30614860211CC967EDA1528E3315">Systematics, Cytogenetics and Molecular Laboratory, Department of Zoology and Biotechnology, Srinagar-Garhwal, Uttarakhand, India</mods:affiliation>
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<mods:nameIdentifier id="E25C3C316982470D9D940FDAE1EF60F0" type="email">fartyalrs@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E43E636507A39265D29A1A55FA0BF889">Sati, Pradeep C.</mods:namePart>
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<mods:affiliation id="08EB4F597C4D7C65ED8D977E7D9FBEFC">Systematics, Cytogenetics and Molecular Laboratory, Department of Zoology and Biotechnology, Srinagar-Garhwal, Uttarakhand, India</mods:affiliation>
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<mods:namePart id="C413D057A0D0E59E1C92A40E63420249">Pradhan, Sushmika</mods:namePart>
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<mods:affiliation id="3EFF5C60D401173F3DD0479289AA9C04">P. G. Department of Zoology, Darjeeling Government College, Darjeeling, West Bengal, India & Genetics Research Unit, Department of Zoology, University of Calcutta, West Bengal, India</mods:affiliation>
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<mods:namePart id="FC9D444371A13F98D1B8B73741646AEB">Kandpal, Mukul C.</mods:namePart>
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<mods:affiliation id="5A8A34D416B1329C4EA3E5387F3DD029">Cytogenetics Laboratory, Department of Zoology, Kumaun University, Nainital, Uttarakhand, India</mods:affiliation>
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<mods:namePart id="55FE39E1326F5F1B78829C27DB0EDED9">Toda, Masanori J.</mods:namePart>
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<mods:affiliation id="A12C0CFD70F417AF7CEAD33214ABB27D">Hokkaido University Museum, Hokkaido University, N 10, W 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="FB38F24688075ED4044ED2E7C47C8668">Chatterjee, Rabindra N.</mods:namePart>
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<mods:affiliation id="4946A05AAA8D0D94A2C1AA82E4B21ADF">Genetics Research Unit, Department of Zoology, University of Calcutta, West Bengal, India</mods:affiliation>
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<mods:namePart id="F146112634F4FBC5FF137B247EAD8B8E">Singh, Birendra K.</mods:namePart>
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<mods:affiliation id="F9473F2EAB7A10913835225DA8BE491A">Cytogenetics Laboratory, Department of Zoology, Kumaun University, Nainital, Uttarakhand, India</mods:affiliation>
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<mods:namePart id="4F715C810980C7E6DEBD843994462BFF">Bhardwai, Asha</mods:namePart>
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<mods:affiliation id="7A93BF3A7DEA5CD2962F818E203E2A5A">Systematics, Cytogenetics and Molecular Laboratory, Department of Zoology and Biotechnology, Srinagar-Garhwal, Uttarakhand, India</mods:affiliation>
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<mods:date id="ABE97E9420DE519D38A3B1B85627FA76">2017</mods:date>
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<mods:number id="1BE7F1E6C23567F8C0AEF7D8CDB44029">2017-08-08</mods:number>
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<mods:number id="15E9DE24F8F2A8ED72C48D45F6070E86">688</mods:number>
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<treatment id="87BACB7EBB0D9A160C5B0BB81C2542B1" ID-GBIF-Taxon="132648609" LSID="urn:lsid:plazi:treatment:87BACB7EBB0D9A160C5B0BB81C2542B1" httpUri="http://treatment.plazi.org/id/87BACB7EBB0D9A160C5B0BB81C2542B1" lastPageNumber="49" pageId="0" pageNumber="49">
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<subSubSection id="A31810946860A2D12ED6F5C218A8D25C" pageId="0" pageNumber="49" type="nomenclature">
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<paragraph id="88258B013AE3742E549532653EDBE78E" pageId="0" pageNumber="49">
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Genus
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<taxonomicName id="2EF13B9DF073EF35BBC8A7C04D60DCB5" ID-CoL="8VTHS" ID-ENA="7331" LSID="525DE0B8-C337-59E4-A576-BB7C4DA8B129" authority="Basden" authorityName="Basden" authorityYear="1961" class="Insecta" family="Drosophilidae" genus="Lordiphosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lordiphosa" order="Diptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Lordiphosa Basden</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0AA92D6F335622012A0D543B33392329" pageId="0" pageNumber="49" type="reference_group">
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<paragraph id="AF2B481AA3500A382BFAFF302A7EE9DB" pageId="0" pageNumber="49">
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<taxonomicName id="49210D73E8F15F8D304CC0F89136AE1A" authorityName="Basden" authorityYear="1961" class="Insecta" family="Drosophilidae" genus="Lordiphosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lordiphosa" order="Diptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Lordiphosa</taxonomicName>
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Basden, 1961: 186 (as a subgenus of
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<taxonomicName id="225598FE763937B9FD710D726F765E1A" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drosophila" order="Diptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Drosophila</taxonomicName>
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);
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<bibRefCitation id="A6495F0B4C4D57A05E83C616F74AA09E" author="Lastovka, P" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="19" pageNumber="68" pagination="404 - 420" refId="B24" refString="Lastovka, P, Maca, J, 1978. European species of the Drosophila subgenus Lordiphosa (Diptera, Drosophilidae). Acta Entomologica Bohemoslovaca 75: 404 - 420" title="European species of the Drosophila subgenus Lordiphosa (Diptera, Drosophilidae)." volume="75" year="1978">
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<normalizedToken id="D3D9A90A4C9C6EDAE7A7B7EE1453F2B5" originalValue="Laštovka">Lastovka</normalizedToken>
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and
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<normalizedToken id="F1327687CC647464CB82B7BC593BFAF7" originalValue="Máca">Maca</normalizedToken>
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1978
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</bibRefCitation>
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: 404;
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<bibRefCitation id="A8291A1C4C609B120BBAFACF06CB8F8B" author="Okada, T" journalOrPublisher="Kontyu" pageId="20" pageNumber="69" pagination="565 - 575" refId="B35" refString="Okada, T, 1984. New or little known species of Drosophila (Lordiphosa) with taximatrical analysis (Diptera: Drosophilidae). Kontyu 52: 565 - 575" title="New or little known species of Drosophila (Lordiphosa) with taximatrical analysis (Diptera: Drosophilidae)." volume="52" year="1984">Okada 1984</bibRefCitation>
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: 571. Type species:
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<taxonomicName id="398A5702709668558A32CF0F5CEF1F2B" authorityName="Fallen" authorityYear="1823" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drosophila fenestrarum" order="Diptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="fenestrarum">Drosophila fenestrarum</taxonomicName>
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<normalizedToken id="5AA80E53CD95CECA1DD0A9FAF07B6BD3" originalValue="Fallén">Fallen</normalizedToken>
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, 1823.
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</paragraph>
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<paragraph id="436DAC13D9AF0ED424E0F6523651B14F" pageId="0" pageNumber="49">
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<taxonomicName id="CB0D7F9A1BBDAE26EE2D8E2C5BD3363E" authorityName="Basden" authorityYear="1961" class="Insecta" family="Drosophilidae" genus="Lordiphosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lordiphosa" order="Diptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">Lordiphosa</taxonomicName>
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: Grimaldi, 1990: 121 (new status as genus);
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<bibRefCitation id="56F7656D5DD11F91CE4B05F2EF7FF6CA" author="De, A" journalOrPublisher="Senckenbergiana Biologica" pageId="18" pageNumber="67" pagination="129 - 133" refId="B8" refString="De, A, Gupta, JP, 1996. Records of drosophilid species from West Bengal with description of one new and two previously unrecorded species from India (Insecta: Diptera: Drosophilidae). Senckenbergiana Biologica 76: 129 - 133" title="Records of drosophilid species from West Bengal with description of one new and two previously unrecorded species from India (Insecta: Diptera: Drosophilidae)." volume="76" year="1996">De and Gupta 1996</bibRefCitation>
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: 131;
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<bibRefCitation id="4E879246A271FF543B41A87F7A49E7CA" author="Baechli, G" journalOrPublisher="Evolution & Development" pageId="18" pageNumber="67" refId="B3" refString="Baechli, G, Vilela, CR, Andersson, Escher S, Saura, A, 2004. The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica Volume 39. Brill Leiden, 362 pp." title="The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica Volume 39. Brill Leiden, 362 pp." year="2004">
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<normalizedToken id="3314577727681E2DCF72724B4B138840" originalValue="Bächli">Baechli</normalizedToken>
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et al. 2004
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</bibRefCitation>
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: 250.
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</paragraph>
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</subSubSection>
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<subSubSection id="2E43FFF24D86B50BBDEA9236DB5C2E8E" pageId="0" pageNumber="49" type="diagnosis">
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<paragraph id="9563146207D0D40AC52612DDE8A15FDE" pageId="0" pageNumber="49">Diagnosis.</paragraph>
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<paragraph id="DC59561F5BB631BF0DF9D31C76271D77" pageId="0" pageNumber="49">
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Prementum thicker in ventral than in dorsal portion from lateral view (Figs
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<figureCitation id="9942A64C2E5A158DF17A387674039531" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lordiphosa curva Fartyal & Toda, sp. n. (♂ ♀ paratypes from Chopta Forest, Uttarakhand, India): A occiput: dorsolateral arms (dl a) and dorsomedial arm (dm a) of tentorial apodeme and supracervical setae (spc s) B, C proboscis: prementum (premnt) (B lateral view C posterior view) D foreleg tarsus E, F periphallic organs (E caudolateral view F caudal view) G, H phallic organs (G ventral view H lateral view) I ♀ terminalia J spermathecae. Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure2" httpUri="https://binary.pensoft.net/fig/149245" pageId="0" pageNumber="49">2B</figureCitation>
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,
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<figureCitation id="D086F93C188574DE01F94B1711651F33" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lordiphosa antillaria (Okada, 1984) (♂ ♀ from Bagdogra, Assam, West Bengal, India): A ♂ left lateral habitus B antenna C mouthparts (lateral view) D periphallic organs (caudolateral view) E, F phallic organs (aedeagal apodeme broken) (E ventral view F ventrolateral view) G ♀ terminalia H spermatheca. Scale bars 0.5 mm A; 0.1 mm B-H." figureDoi="10.3897/zookeys.688.12590.figure3" httpUri="https://binary.pensoft.net/fig/149246" pageId="0" pageNumber="49">3C</figureCitation>
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,
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<figureCitation id="8D7AF708189873DE68EA2CB4CE3BDA8B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lordiphosa ayarpathaensis Kandpal & Singh, sp. n. (♂ ♀ paratypes from Ayarpatha, Nainital district, Kumaon, Uttarakhand, India): A, B proboscis (A lateral view B posterior view) C, D periphallic organs (C caudolateral view D caudoventral view) E, F phallic organs: aedeagal basal processes (aed b p) (E ventral view F lateral view) G, H ♀ terminalia: perineal plate (per pl) (G lateral view H ventral view) I spermatheca. Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure4" httpUri="https://binary.pensoft.net/fig/149247" pageId="0" pageNumber="49">4A</figureCitation>
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), ventrally more or less expanded in posterior view (Figs
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<figureCitation id="3BC45B1F4067131F5B004F73CE7429ED" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lordiphosa curva Fartyal & Toda, sp. n. (♂ ♀ paratypes from Chopta Forest, Uttarakhand, India): A occiput: dorsolateral arms (dl a) and dorsomedial arm (dm a) of tentorial apodeme and supracervical setae (spc s) B, C proboscis: prementum (premnt) (B lateral view C posterior view) D foreleg tarsus E, F periphallic organs (E caudolateral view F caudal view) G, H phallic organs (G ventral view H lateral view) I ♀ terminalia J spermathecae. Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure2" httpUri="https://binary.pensoft.net/fig/149245" pageId="0" pageNumber="49">2C</figureCitation>
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,
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<figureCitation id="6285D84A78027BE0937DCE25924A93BE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lordiphosa ayarpathaensis Kandpal & Singh, sp. n. (♂ ♀ paratypes from Ayarpatha, Nainital district, Kumaon, Uttarakhand, India): A, B proboscis (A lateral view B posterior view) C, D periphallic organs (C caudolateral view D caudoventral view) E, F phallic organs: aedeagal basal processes (aed b p) (E ventral view F lateral view) G, H ♀ terminalia: perineal plate (per pl) (G lateral view H ventral view) I spermatheca. Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure4" httpUri="https://binary.pensoft.net/fig/149247" pageId="0" pageNumber="49">4B</figureCitation>
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). Paramere ventrobasally articulated with hypandrium (Figs
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<figureCitation id="8C109F92C7F9E9F24F7D7544256DB32C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lordiphosa neokurokawai (Singh & Gupta, 1981) (♂ ♀ from Kunming, Yunnan, China): A cibarium: anterior sensilla (a s), medial sensilla (m s), posterior sensilla (p s) and sensilla campaniformia (s c) (dorsal view) B foreleg tarsus C periphallic organs: epandrium (epand), cercus (cerc) and surstylus (sur) (caudal view) D, E phallic organs: aedeagus (aed), aedeagal apodeme (aed a), gonopods (gon), hypandrium (hypd) and paramere (pm) (D ventral view E lateral view) F ♀ terminalia: tergite VIII (8 T), epiproct (epiprct), hypoproct (hyprct) and oviscapt (ov) G spermatheca. Arrowheads indicate the diagnostic characters. Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure1" httpUri="https://binary.pensoft.net/fig/149244" pageId="0" pageNumber="49">1D</figureCitation>
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,
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<figureCitation id="9B2161FFBCA4229F808160815DFDFA19" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lordiphosa curva Fartyal & Toda, sp. n. (♂ ♀ paratypes from Chopta Forest, Uttarakhand, India): A occiput: dorsolateral arms (dl a) and dorsomedial arm (dm a) of tentorial apodeme and supracervical setae (spc s) B, C proboscis: prementum (premnt) (B lateral view C posterior view) D foreleg tarsus E, F periphallic organs (E caudolateral view F caudal view) G, H phallic organs (G ventral view H lateral view) I ♀ terminalia J spermathecae. Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure2" httpUri="https://binary.pensoft.net/fig/149245" pageId="0" pageNumber="49">2H</figureCitation>
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,
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<figureCitation id="63A1B7FABD3E4FF32A55AB69A163E7CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lordiphosa antillaria (Okada, 1984) (♂ ♀ from Bagdogra, Assam, West Bengal, India): A ♂ left lateral habitus B antenna C mouthparts (lateral view) D periphallic organs (caudolateral view) E, F phallic organs (aedeagal apodeme broken) (E ventral view F ventrolateral view) G ♀ terminalia H spermatheca. Scale bars 0.5 mm A; 0.1 mm B-H." figureDoi="10.3897/zookeys.688.12590.figure3" httpUri="https://binary.pensoft.net/fig/149246" pageId="0" pageNumber="49">3E</figureCitation>
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<figureCitation id="5B299A7EF9DBDB1082B848D0B1230FC7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lordiphosa ayarpathaensis Kandpal & Singh, sp. n. (♂ ♀ paratypes from Ayarpatha, Nainital district, Kumaon, Uttarakhand, India): A, B proboscis (A lateral view B posterior view) C, D periphallic organs (C caudolateral view D caudoventral view) E, F phallic organs: aedeagal basal processes (aed b p) (E ventral view F lateral view) G, H ♀ terminalia: perineal plate (per pl) (G lateral view H ventral view) I spermatheca. Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure4" httpUri="https://binary.pensoft.net/fig/149247" pageId="0" pageNumber="49">4E, F</figureCitation>
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<figureCitation id="BBDC436023A0CB7E136E12015633AB34" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lordiphosa srinagarensis Sati & Fartyal, sp. n. (♂ paratype from HNB Garhwal University, Srinagar, Garhwal, Uttarakhand, India): A head (frontal view) B periphallic organs (caudolateral view) C, D phallic organs (C ventral view D lateral view). Scale bars 0.1 mm." figureDoi="10.3897/zookeys.688.12590.figure7" httpUri="https://binary.pensoft.net/fig/149250" pageId="0" pageNumber="49">7D</figureCitation>
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). Hypandrium lacking paramedian setae.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |