treatments-xml/data/81/5F/2A/815F2A49FFB8B448FF5CFB73C5CBFC21.xml

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<mods:title id="1444F0228FB6BF6A531078F1FED976D2">Updates on the taxonomy and distribution of Gastroserica Brenske, 1897, Microserica Brenske, 1894, and Moronoserica Ahrens, Lukic &amp; Liu, 2023 (Coleoptera: Scarabaeidae: Sericinae: Sericini)</mods:title>
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<mods:namePart id="096C6A08F53BBB7EFB14930ABCFA2201">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="D10CEACFF7FC1AB01A976FF224B395F3">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany. ahrens. dirk _ col @ gmx. de, https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
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<mods:namePart id="A124CA03D84A60BE09F5158499E9038C">Pham, Phu</mods:namePart>
<mods:affiliation id="17BB983D57A47D1D808D80DC1AFEC20A">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<subSubSection id="41ECC8D4FFB8B449FF5CFB73C3F5FB5E" box="[151,931,1183,1210]" pageId="4" pageNumber="555" type="nomenclature">
<paragraph id="09499B5FFFB8B449FF5CFB73C3F5FB5E" blockId="4.[151,931,1183,1210]" box="[151,931,1183,1210]" pageId="4" pageNumber="555">
<heading id="52012C33FFB8B449FF5CFB73C3F5FB5E" bold="true" box="[151,931,1183,1210]" fontSize="11" level="1" pageId="4" pageNumber="555" reason="1">
<emphasis id="3B82474DFFB8B449FF5CFB73C3F5FB5E" bold="true" box="[151,931,1183,1210]" pageId="4" pageNumber="555">
Updated key to 
<taxonomicName id="CEF6E0DCFFB8B449FE96FB73C2CAFB5E" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[349,668,1183,1210]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="subGenus" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FE96FB73C1A0FB5E" bold="true" box="[349,502,1184,1210]" italics="true" pageId="4" pageNumber="555">Gastroserica</emphasis>
(
<emphasis id="3B82474DFFB8B449FDCCFB4CC2C7FB5E" bold="true" box="[519,657,1183,1210]" italics="true" pageId="4" pageNumber="555">Helioserica</emphasis>
)
</taxonomicName>
species (males only):
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="41ECC8D4FFB8B448FF5CFB36C5CBFC21" lastPageId="5" lastPageNumber="556" pageId="4" pageNumber="555" type="key">
<key id="7D67CC6EFFB8B448FF5CFB36C5CBFC21" lastPageId="5" lastPageNumber="556" pageId="4" pageNumber="555">
<keyStep id="B20283FAFFB8B449FF5CFB36C5CAFAFC" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CFB36C5CAFB1E" blockId="4.[151,1437,1253,2038]" box="[151,1436,1253,1275]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFB36C5CAFB1E" box="[151,1436,1253,1275]" pageId="4" pageNumber="555">1 Dorsal surface mostly dull or dull-iridescent................................................................ 2</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CFAD0C5CAFAFC" blockId="4.[151,1437,1253,2038]" box="[151,1436,1282,1304]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFAD0C5CAFAFC" box="[151,1436,1282,1304]" pageId="4" pageNumber="555">- Dorsal surface completely simply shiny, or shiny-iridescent................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CFAF3C5CAFAB6" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CFAF3C5CBFAD1" blockId="4.[151,1437,1253,2038]" box="[151,1437,1311,1334]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFAF3C5CBFAD1" box="[151,1437,1311,1334]" pageId="4" pageNumber="555">
2 Elytra with multiple dark spots and darker striae....................................... 
<taxonomicName id="CEF6E0DCFFB8B449FB4AFACCC5CBFAD1" authority="(Frey, 1972)" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1153,1437,1311,1333]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="cognata" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FB4AFACCC5CBFAD1" bold="true" box="[1153,1437,1311,1333]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FB4AFACCC4CFFAD0" bold="true" box="[1153,1177,1311,1332]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FB6CFAF3C496FAD1" bold="true" box="[1191,1216,1312,1333]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FB06FAF3C54AFAD1" bold="true" box="[1229,1308,1312,1333]" italics="true" pageId="4" pageNumber="555">cognata</emphasis>
(Frey, 1972)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CFAEEC5CAFAB6" blockId="4.[151,1437,1253,2038]" box="[151,1436,1341,1363]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFAEEC5CAFAB6" box="[151,1436,1341,1363]" pageId="4" pageNumber="555">- Elytra without multiple dark spots and striae of the same colour as intervals....................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CFA89C5CBFA69" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CFA89C5CAFA8B" blockId="4.[151,1437,1253,2038]" box="[151,1436,1370,1392]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFA89C5CAFA8B" box="[151,1436,1370,1392]" pageId="4" pageNumber="555">3 Left paramere with a distinct basal lobe.................................................................... 4</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CFAABC5CBFA69" blockId="4.[151,1437,1253,2038]" box="[151,1437,1399,1422]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFAABC5CBFA69" box="[151,1437,1399,1422]" pageId="4" pageNumber="555">
- Left paramere without a distinct basal lobe......................................... 
<taxonomicName id="CEF6E0DCFFB8B449FBAFFAA4C5CBFA69" authority="(Frey, 1972)" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1124,1437,1399,1421]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="quateorum" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FBAFFAA4C5CBFA69" bold="true" box="[1124,1437,1399,1421]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FBAFFAA4C42DFA68" bold="true" box="[1124,1147,1399,1420]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FB42FAABC4F4FA69" bold="true" box="[1161,1186,1400,1421]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FB7BFAABC54AFA69" bold="true" box="[1200,1308,1400,1421]" italics="true" pageId="4" pageNumber="555">quateorum</emphasis>
(Frey, 1972)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CFA46C5CAFA2C" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CFA46C5CBFA4F" blockId="4.[151,1437,1253,2038]" box="[151,1437,1428,1451]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFA46C5CBFA4F" box="[151,1437,1428,1451]" pageId="4" pageNumber="555">
4 Left, distal portion of phallobase strongly widened (lateral view)........................ 
<taxonomicName id="CEF6E0DCFFB8B449FBA7FA46C5CBFA4F" authority="(Brenske, 1897)" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[1132,1437,1428,1451]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="hiulca" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FBA7FA46C5CBFA4F" bold="true" box="[1132,1437,1428,1451]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FBA7FA46C4D5FA4E" bold="true" box="[1132,1155,1429,1450]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FB5AFA46C4FCFA4E" bold="true" box="[1169,1194,1429,1450]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FB73FA47C4A1FA4E" bold="true" box="[1208,1271,1428,1450]" italics="true" pageId="4" pageNumber="555">hiulca</emphasis>
(Brenske, 1897)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CFA60C5CAFA2C" blockId="4.[151,1437,1253,2038]" box="[151,1436,1458,1480]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFA60C5CAFA2C" box="[151,1436,1458,1480]" pageId="4" pageNumber="555">- Distal portion of phallobase at both sides narrowed towards apex (lateral view)..................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CFA1CC5CAF9E6" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CFA1CC5CAFA02" blockId="4.[151,1437,1253,2038]" box="[151,1436,1487,1510]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFA1CC5CAFA02" box="[151,1436,1487,1510]" pageId="4" pageNumber="555">5 Left paramere bilobed in distal part....................................................................... 6</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CFA3EC5CAF9E6" blockId="4.[151,1437,1253,2038]" box="[151,1436,1517,1539]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CFA3EC5CAF9E6" box="[151,1436,1517,1539]" pageId="4" pageNumber="555">- Left paramere simple in distal part (with just one lobe or branch)............................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CF9D9C5CBF9D9" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CF9D9C5CAF9C4" blockId="4.[151,1437,1253,2038]" box="[151,1436,1546,1568]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF9D9C5CAF9C4" box="[151,1436,1546,1568]" pageId="4" pageNumber="555">6 Phallobase with a ventral lamellose extension on the left side.................................................. 7</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CF9FBC5CBF9D9" blockId="4.[151,1437,1253,2038]" box="[151,1437,1575,1598]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF9FBC5CBF9D9" box="[151,1437,1575,1598]" pageId="4" pageNumber="555">
- Phallobase without a ventral lamellose extension on the left side..................... 
<taxonomicName id="CEF6E0DCFFB8B449FB85F9F4C5CBF9D9" authority="(Nomura, 1974)" baseAuthorityName="Nomura" baseAuthorityYear="1974" box="[1102,1437,1575,1597]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="bisignata" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FB85F9F4C5CBF9D9" bold="true" box="[1102,1437,1575,1597]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FB85F9F4C433F9D8" bold="true" box="[1102,1125,1575,1596]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FBBFF9FBC4DAF9D9" bold="true" box="[1140,1164,1576,1597]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FB51F9F4C4A3F9D9" bold="true" box="[1178,1269,1575,1597]" italics="true" pageId="4" pageNumber="555">bisignata</emphasis>
(Nomura, 1974)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CF996C5CAF956" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CF996C5CBF99C" blockId="4.[151,1437,1253,2038]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF996C5CBF99C" pageId="4" pageNumber="555">
7 Left paramere robust, as wide as apical phallobase. Both distal lobes of left paramere subequal in length to the right paramere................................................................. 
<taxonomicName id="CEF6E0DCFFB8B449FB8DF9B1C5CBF99C" authority="(Nomura, 1974)" baseAuthorityName="Nomura" baseAuthorityYear="1974" box="[1094,1437,1634,1656]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="nitidipyga" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FB8DF9B1C5CBF99C" bold="true" box="[1094,1437,1634,1656]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FB8DF9B1C40BF993" bold="true" box="[1094,1117,1634,1655]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FBA7F9B0C4D2F99C" bold="true" box="[1132,1156,1635,1656]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FB59F9B1C4A3F99C" bold="true" box="[1170,1269,1634,1656]" italics="true" pageId="4" pageNumber="555">nitidipyga</emphasis>
(Nomura, 1974)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CF953C5CAF956" blockId="4.[151,1437,1253,2038]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF953C5CAF956" pageId="4" pageNumber="555">
- Left paramere narrower, distinctly more narrow than apical phallobase. External distal lobes of left paramere half of length to the right paramere............................................... 
<taxonomicName id="CEF6E0DCFFB8B449FC07F94EC5CAF956" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[972,1436,1692,1715]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="yuebaensis" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FC07F94EC5CAF956" bold="true" box="[972,1436,1692,1715]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FC07F94EC3B5F956" bold="true" box="[972,995,1693,1714]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FC3AF94EC45CF956" bold="true" box="[1009,1034,1693,1714]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FBD3F94FC4D2F956" bold="true" box="[1048,1156,1692,1714]" italics="true" pageId="4" pageNumber="555">yuebaensis</emphasis>
<bibRefCitation id="6D67E6AEFFB8B449FB41F94EC5CAF956" author="Ahrens, D. &amp; Liu, W. G. &amp; Lukic, D. &amp; Bai, M." box="[1162,1436,1693,1715]" pageId="4" pageNumber="555" pagination="1 - 115" refId="ref18504" refString="Ahrens, D., Liu, W. G., Lukic, D. &amp; Bai, M. (2023) A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae, Sericini). Zootaxa, 5241 (1), 1 - 115. https: // doi. org / 10.11646 / zootaxa. 5241.1.1" type="journal article" year="2023">Ahrens, Lukic &amp; Liu, 2023</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CF969C5CAF909" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CF969C5CAF92B" blockId="4.[151,1437,1253,2038]" box="[151,1436,1722,1744]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF969C5CAF92B" box="[151,1436,1722,1744]" pageId="4" pageNumber="555">8 Right side of apical phallobase with a small granulate raster area................................................ 9</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CF90BC5CAF909" blockId="4.[151,1437,1253,2038]" box="[151,1436,1752,1774]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF90BC5CAF909" box="[151,1436,1752,1774]" pageId="4" pageNumber="555">- Right side of apical phallobase without a granulate raster area................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CF926C5CAF886" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CF926C5CAF8C3" blockId="4.[151,1437,1253,2038]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF926C5CAF8C3" pageId="4" pageNumber="555">
9 Left paramere basally wider (lateral view). Right paramere before apex almost straight............................................................................................... 
<taxonomicName id="CEF6E0DCFFB8B449FC23F8C1C5CAF8C3" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[1000,1436,1810,1832]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="fumaria" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FC23F8C1C5CAF8C3" bold="true" box="[1000,1436,1810,1832]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FC23F8C1C3A9F8C3" bold="true" box="[1000,1023,1810,1831]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FBC6F8C0C470F8CC" bold="true" box="[1037,1062,1811,1832]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FBFFF8C1C4D2F8CC" bold="true" box="[1076,1156,1810,1832]" italics="true" pageId="4" pageNumber="555">fumaria</emphasis>
<bibRefCitation id="6D67E6AEFFB8B449FB41F8C1C5CAF8C3" author="Ahrens, D. &amp; Liu, W. G. &amp; Lukic, D. &amp; Bai, M." box="[1162,1436,1810,1832]" pageId="4" pageNumber="555" pagination="1 - 115" refId="ref18504" refString="Ahrens, D., Liu, W. G., Lukic, D. &amp; Bai, M. (2023) A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae, Sericini). Zootaxa, 5241 (1), 1 - 115. https: // doi. org / 10.11646 / zootaxa. 5241.1.1" type="journal article" year="2023">Ahrens, Lukic &amp; Liu, 2023</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CF8E3C5CAF886" blockId="4.[151,1437,1253,2038]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF8E3C5CAF886" pageId="4" pageNumber="555">
- Left paramere basally more narrow (lateral view). Right paramere before apex strongly bent externally.............................................................................. 
<taxonomicName id="CEF6E0DCFFB8B449FC74F89EC5CAF886" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[959,1436,1868,1891]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="phukradung" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FC74F89EC5CAF886" bold="true" box="[959,1436,1868,1891]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FC74F89EC380F886" bold="true" box="[959,982,1869,1890]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FC2FF89EC3ABF886" bold="true" box="[996,1021,1869,1890]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FBC0F89FC4D0F886" bold="true" box="[1035,1158,1868,1890]" italics="true" pageId="4" pageNumber="555">phukradung</emphasis>
<bibRefCitation id="6D67E6AEFFB8B449FB41F89EC5CAF886" author="Ahrens, D. &amp; Liu, W. G. &amp; Lukic, D. &amp; Bai, M." box="[1162,1436,1869,1891]" pageId="4" pageNumber="555" pagination="1 - 115" refId="ref18504" refString="Ahrens, D., Liu, W. G., Lukic, D. &amp; Bai, M. (2023) A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae, Sericini). Zootaxa, 5241 (1), 1 - 115. https: // doi. org / 10.11646 / zootaxa. 5241.1.1" type="journal article" year="2023">Ahrens, Lukic &amp; Liu, 2023</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B449FF5CF8B9C5CAF879" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CF8B9C5CAF89B" blockId="4.[151,1437,1253,2038]" box="[151,1436,1898,1920]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF8B9C5CAF89B" box="[151,1436,1898,1920]" pageId="4" pageNumber="555">10 Right paramere in basal two thirds strongly widened (lateral view)............................................. 12</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B449FF5CF85BC5CAF879" blockId="4.[151,1437,1253,2038]" box="[151,1436,1928,1950]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF85BC5CAF879" box="[151,1436,1928,1950]" pageId="4" pageNumber="555">- Right paramere in basal half not particularly widened (lateral view)............................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB8B448FF5CF876C5CBFF49" lastPageId="5" lastPageNumber="556" pageId="4" pageNumber="555">
<paragraph id="09499B5FFFB8B449FF5CF876C5CBF83C" blockId="4.[151,1437,1253,2038]" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B449FF5CF876C5CBF83C" pageId="4" pageNumber="555">
11 Right paramere more than half as long as phallobase. Basal lobe of long left paramere much shorter than the rest of the paramere..................................................................... 
<taxonomicName id="CEF6E0DCFFB8B449FBBDF811C5CBF83C" authority="(Moser, 1915)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1142,1437,1986,2008]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="varians" subGenus="Helioserica">
<emphasis id="3B82474DFFB8B449FBBDF811C5CBF83C" bold="true" box="[1142,1437,1986,2008]" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FBBDF811C4DBF833" bold="true" box="[1142,1165,1986,2007]" italics="true" pageId="4" pageNumber="555">G.</emphasis>
(
<emphasis id="3B82474DFFB8B449FB50F810C4E2F83C" bold="true" box="[1179,1204,1987,2008]" italics="true" pageId="4" pageNumber="555">H.</emphasis>
) 
<emphasis id="3B82474DFFB8B449FB09F811C55DF833" bold="true" box="[1218,1291,1986,2007]" italics="true" pageId="4" pageNumber="555">varians</emphasis>
(Moser, 1915)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB8B448FF5CF833C5CBFF49" blockId="4.[151,1437,1253,2038]" lastBlockId="5.[151,1437,151,965]" lastPageId="5" lastPageNumber="556" pageId="4" pageNumber="555">
<keyLead id="B207386AFFB8B448FF5CF833C5CBFF49" lastPageId="5" lastPageNumber="556" pageId="4" pageNumber="555">
<emphasis id="3B82474DFFB8B449FF5CF833C0C9F811" bold="true" box="[151,159,2016,2037]" pageId="4" pageNumber="555">-</emphasis>
Right paramere half as long as phallobase. Basal lobe of short left paramere as long as the rest of the paramere............................................................................. 
<emphasis id="3B82474DFFB9B448FC39FF44C5CBFF49" bold="true" box="[1010,1437,151,173]" pageId="5" pageNumber="556">
<taxonomicName id="CEF6E0DCFFB9B448FC39FF44C54DFF49" authority="Ahrens &amp; Pham" authorityName="Ahrens &amp; Pham" authorityYear="2023" box="[1010,1307,151,173]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="horii" status="new species" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC39FF44C45FFF48" bold="true" box="[1010,1033,151,172]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBDCFF4BC466FF49" bold="true" box="[1047,1072,152,173]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBF5FF44C438FF49" bold="true" box="[1086,1134,151,173]" italics="true" pageId="5" pageNumber="556">horii</emphasis>
Ahrens &amp; Pham
</taxonomicName>
, 
<taxonomicNameLabel id="20B1FA36FFB9B448FAEDFF4BC5CBFF49" box="[1318,1437,151,173]" pageId="5" pageNumber="556" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFF66C5CBFF0C" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFF66C5CAFF2E" blockId="5.[151,1437,151,965]" box="[151,1436,181,203]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFF66C5CAFF2E" box="[151,1436,181,203]" pageId="5" pageNumber="556">12 Left paramere ventrally strongly excavated and bent......................................................... 13</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFF00C5CBFF0C" blockId="5.[151,1437,151,965]" box="[151,1437,210,232]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFF00C5CBFF0C" box="[151,1437,210,232]" pageId="5" pageNumber="556">
- Left paramere ventrally not excavated and straight....................... 
<taxonomicName id="CEF6E0DCFFB9B448FC2FFF01C5CBFF0C" authority="(Fabrizi &amp; Ahrens, 2016)" baseAuthorityName="Fabrizi &amp; Ahrens" baseAuthorityYear="2016" box="[996,1437,210,232]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="roingensis" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC2FFF01C5CBFF0C" bold="true" box="[996,1437,210,232]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FC2FFF01C3ADFF03" bold="true" box="[996,1019,210,231]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBC2FF01C474FF03" bold="true" box="[1033,1058,210,231]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBFBFF01C4C1FF03" bold="true" box="[1072,1175,210,231]" italics="true" pageId="5" pageNumber="556">roingensis</emphasis>
(Fabrizi &amp; 
<bibRefCitation id="6D67E6AEFFB9B448FAC5FF01C5C3FF0C" author="Ahrens, D. &amp; Fabrizi, S." box="[1294,1429,210,232]" pageId="5" pageNumber="556" pagination="1 - 355" refId="ref17716" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens, 2016</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFF23C5CBFEC7" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFF23C5CAFEE1" blockId="5.[151,1437,151,965]" box="[151,1436,239,261]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFF23C5CAFEE1" box="[151,1436,239,261]" pageId="5" pageNumber="556">13 Right paramere at apex strongly bent externally (dorsal view).................................................. 14</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFEDEC5CBFEC7" blockId="5.[151,1437,151,965]" box="[151,1437,268,291]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFEDEC5CBFEC7" box="[151,1437,268,291]" pageId="5" pageNumber="556">
- Right paramere at apex weakly bent externally (dorsal view)................. 
<taxonomicName id="CEF6E0DCFFB9B448FC33FEDEC5CBFEC7" authority="(Ahrens &amp; Fabrizi, 2009)" baseAuthorityName="Ahrens &amp; Fabrizi" baseAuthorityYear="2009" box="[1016,1437,268,291]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="dohertyi" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC33FEDEC5CBFEC7" bold="true" box="[1016,1437,268,291]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FC33FEDEC459FEC6" bold="true" box="[1016,1039,269,290]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBD6FEDEC460FEC6" bold="true" box="[1053,1078,269,290]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FB8FFEDFC4C3FEC6" bold="true" box="[1092,1173,268,290]" italics="true" pageId="5" pageNumber="556">dohertyi</emphasis>
(Ahrens &amp; Fabrizi, 2009)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFEF9C5CAFEB8" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFEF9C5CAFEDB" blockId="5.[151,1437,151,965]" box="[151,1436,298,320]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFEF9C5CAFEDB" box="[151,1436,298,320]" pageId="5" pageNumber="556">
14 Elytra colour yellow to dark brown. Basal lobe of left paramere short...................... 
<taxonomicName id="CEF6E0DCFFB9B448FB4CFEF9C5CAFEDB" authority="Frey, 1972" authorityName="Frey" authorityYear="1972" box="[1159,1436,298,320]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="fukiensis" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FB4CFEF9C5CAFEDB" bold="true" box="[1159,1436,298,320]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FB4CFEF9C4C8FEDB" bold="true" box="[1159,1182,298,319]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FB67FEF9C493FEDB" bold="true" box="[1196,1221,298,319]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FB18FEF9C57AFEA4" bold="true" box="[1235,1324,298,320]" italics="true" pageId="5" pageNumber="556">fukiensis</emphasis>
Frey, 1972
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFE9BC5CAFEB8" blockId="5.[151,1437,151,965]" box="[151,1436,327,349]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFE9BC5CAFEB8" box="[151,1436,327,349]" pageId="5" pageNumber="556">
- Elytra colour blackish, pronotum reddish. Basal lobe of left paramere long... 
<taxonomicName id="CEF6E0DCFFB9B448FC06FE94C5CAFEB8" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[973,1436,327,349]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="simaoensis" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC06FE94C5CAFEB8" bold="true" box="[973,1436,327,349]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FC06FE94C3B2FEB8" bold="true" box="[973,996,327,348]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FC38FE9BC45AFEB9" bold="true" box="[1011,1036,328,349]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBD2FE94C4D3FEB8" bold="true" box="[1049,1157,327,348]" italics="true" pageId="5" pageNumber="556">simaoensis</emphasis>
<bibRefCitation id="6D67E6AEFFB9B448FB41FE94C5CAFEB8" author="Ahrens, D. &amp; Liu, W. G. &amp; Lukic, D. &amp; Bai, M." box="[1162,1436,327,349]" pageId="5" pageNumber="556" pagination="1 - 115" refId="ref18504" refString="Ahrens, D., Liu, W. G., Lukic, D. &amp; Bai, M. (2023) A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae, Sericini). Zootaxa, 5241 (1), 1 - 115. https: // doi. org / 10.11646 / zootaxa. 5241.1.1" type="journal article" year="2023">Ahrens, Lukic &amp; Liu, 2023</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFEB6C5CAFE73" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFEB6C5CAFE9F" blockId="5.[151,1437,151,965]" box="[151,1436,357,379]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFEB6C5CAFE9F" box="[151,1436,357,379]" pageId="5" pageNumber="556">15 Phallobase ventrally with prominent lateral elevations....................................................... 16</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFE50C5CAFE73" blockId="5.[151,1437,151,965]" box="[151,1436,386,408]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFE50C5CAFE73" box="[151,1436,386,408]" pageId="5" pageNumber="556">- Phallobase ventrally without prominent lateral elevations..................................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFE4CC5CAFE36" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFE4CC5CAFE51" blockId="5.[151,1437,151,965]" box="[151,1436,415,437]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFE4CC5CAFE51" box="[151,1436,415,437]" pageId="5" pageNumber="556">16 Prominent lateral elevations of phallobase at apex.......................................................... 17</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFE6EC5CAFE36" blockId="5.[151,1437,151,965]" box="[151,1436,445,467]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFE6EC5CAFE36" box="[151,1436,445,467]" pageId="5" pageNumber="556">- Prominent lateral elevations of phallobase at middle......................................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFE09C5CAFDA3" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFE09C5CAFDE8" blockId="5.[151,1437,151,965]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFE09C5CAFDE8" pageId="5" pageNumber="556">
17 Right paramere more than half as long as phallobase. Eyes moderately large, ratio diameter/interocular width: 0.64................................................................... 
<taxonomicName id="CEF6E0DCFFB9B448FC64FE24C5CAFDE8" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[943,1436,503,525]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="piceocoerulea" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC64FE24C5CAFDE8" bold="true" box="[943,1436,503,525]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FC64FE24C390FDE8" bold="true" box="[943,966,503,524]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FC1FFE2BC3BBFDE9" bold="true" box="[980,1005,504,525]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FC30FE24C4D2FDE9" bold="true" box="[1019,1156,503,525]" italics="true" pageId="5" pageNumber="556">piceocoerulea</emphasis>
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</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFDC6C5CAFDA3" blockId="5.[151,1437,151,965]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFDC6C5CAFDA3" pageId="5" pageNumber="556">
- Right paramere less than half as long as phallobase. Eyes very large, ratio diameter/interocular width: 0.93.................................................................................... 
<taxonomicName id="CEF6E0DCFFB9B448FBDEFDE1C5CAFDA3" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[1045,1436,562,584]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="loei" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FBDEFDE1C5CAFDA3" bold="true" box="[1045,1436,562,584]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FBDEFDE1C47AFDA3" bold="true" box="[1045,1068,562,583]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBF0FDE1C402FDA3" bold="true" box="[1083,1108,562,583]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBAAFDE1C4D3FDAC" bold="true" box="[1121,1157,562,584]" italics="true" pageId="5" pageNumber="556">loei</emphasis>
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</emphasis>
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</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFD83C5CBFD44" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFD83C5CAFD81" blockId="5.[151,1437,151,965]" box="[151,1436,591,613]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFD83C5CAFD81" box="[151,1436,591,613]" pageId="5" pageNumber="556">18 Left paramere with a distinct and subequal ventral and dorsal/basal lobe......................................... 19</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFDBEC5CBFD44" blockId="5.[151,1437,151,965]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFDBEC5CBFD44" pageId="5" pageNumber="556">
- Left paramere without a ventral and dorsal/basal lobe, the latter basally reduced in length and developed short and medially..................................................................... 
<taxonomicName id="CEF6E0DCFFB9B448FBC2FD59C5CBFD44" authority="(Ahrens &amp; Fabrizi, 2009)" baseAuthorityName="Ahrens &amp; Fabrizi" baseAuthorityYear="2009" box="[1033,1437,650,672]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="lucens" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FBC2FD59C5CBFD44" bold="true" box="[1033,1437,650,672]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FBC2FD59C477FD7B" bold="true" box="[1033,1057,650,671]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBE4FD59C41EFD7B" bold="true" box="[1071,1096,650,671]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FB9EFD59C4C3FD44" bold="true" box="[1109,1173,650,672]" italics="true" pageId="5" pageNumber="556">lucens</emphasis>
(Ahrens &amp; Fabrizi, 2009)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFD7BC5CAFD13" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFD7BC5CAFD58" blockId="5.[151,1437,151,965]" box="[151,1436,679,701]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFD7BC5CAFD58" box="[151,1436,679,701]" pageId="5" pageNumber="556">
19 Dorsal/basal lobe shorter than the ventral one, sharply pointed, simple........... 
<taxonomicName id="CEF6E0DCFFB9B448FBDFFD74C5CAFD58" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[1044,1436,679,701]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="ivoi" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FBDFFD74C5CAFD58" bold="true" box="[1044,1436,679,701]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FBDFFD74C47DFD58" bold="true" box="[1044,1067,679,700]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBF2FD7BC404FD59" bold="true" box="[1081,1106,680,701]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBABFD74C4D2FD58" bold="true" box="[1120,1156,679,700]" italics="true" pageId="5" pageNumber="556">ivoi</emphasis>
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</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFD16C5CAFD13" blockId="5.[151,1437,151,965]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFD16C5CAFD13" pageId="5" pageNumber="556">
- Dorsal/basal lobe as long as the ventral one, rounded at apex, tubuliform with a membranous opening at median apex............................................................... 
<taxonomicName id="CEF6E0DCFFB9B448FC5AFD31C5CAFD13" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[913,1436,738,760]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="lucidomarginalis" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC5AFD31C5CAFD13" bold="true" box="[913,1436,738,760]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FC5AFD31C3FEFD13" bold="true" box="[913,936,738,759]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FC7CFD31C399FD13" bold="true" box="[951,975,738,759]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FC16FD31C4D3FD1C" bold="true" box="[989,1157,738,760]" italics="true" pageId="5" pageNumber="556">lucidomarginalis</emphasis>
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</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFCD3C5CAFCAB" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFCD3C5CAFCD6" blockId="5.[151,1437,151,965]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFCD3C5CAFCD6" pageId="5" pageNumber="556">
20 Left paramere with a distinct and subequal ventral and dorsal lobe, dorsal (=basal) lobe directed distally............................................................................... 
<taxonomicName id="CEF6E0DCFFB9B448FC06FCCEC5CAFCD6" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[973,1436,796,819]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="rubropicea" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC06FCCEC5CAFCD6" bold="true" box="[973,1436,796,819]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FC06FCCEC3B2FCD6" bold="true" box="[973,996,797,818]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FC38FCCEC45AFCD6" bold="true" box="[1011,1036,797,818]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBD2FCCFC4D3FCD6" bold="true" box="[1049,1157,796,818]" italics="true" pageId="5" pageNumber="556">rubropicea</emphasis>
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</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFCE8C5CAFCAB" blockId="5.[151,1437,151,965]" box="[151,1436,826,848]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFCE8C5CAFCAB" box="[151,1436,826,848]" pageId="5" pageNumber="556">- Left paramere dorsal lobe shorter and developed medially.................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="B20283FAFFB9B448FF5CFC8BC5CBFC21" pageId="5" pageNumber="556">
<paragraph id="09499B5FFFB9B448FF5CFC8BC5CAFC6E" blockId="5.[151,1437,151,965]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFC8BC5CAFC6E" pageId="5" pageNumber="556">
21 Right paramere straight (lateral view), widened at apex (dorsal view). Basal lobe of left paramere mesobasally directed..................................................................... 
<taxonomicName id="CEF6E0DCFFB9B448FC26FCA6C5CAFC6E" authority="Ahrens, Lukic &amp; Liu, 2023" authorityName="Ahrens, Lukic &amp; Liu" authorityYear="2023" box="[1005,1436,884,907]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="bannok" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC26FCA6C5CAFC6E" bold="true" box="[1005,1436,884,907]" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FC26FCA6C452FC6E" bold="true" box="[1005,1028,885,906]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBD8FCA6C47AFC6E" bold="true" box="[1043,1068,885,906]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBF2FCA7C4D2FC6E" bold="true" box="[1081,1156,884,906]" italics="true" pageId="5" pageNumber="556">bannok</emphasis>
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</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="09499B5FFFB9B448FF5CFC41C5CBFC21" blockId="5.[151,1437,151,965]" pageId="5" pageNumber="556">
<keyLead id="B207386AFFB9B448FF5CFC41C5CBFC21" pageId="5" pageNumber="556">
<emphasis id="3B82474DFFB9B448FF5CFC41C0C9FC43" bold="true" box="[151,159,914,935]" pageId="5" pageNumber="556">-</emphasis>
Right paramere weakly curved ventrally (lateral view), not widened at apex (dorsal view). Basal lobe of left paramere mesally directed......................................................... 
<emphasis id="3B82474DFFB9B448FC15FC7CC5CBFC21" bold="true" box="[990,1437,943,965]" pageId="5" pageNumber="556">
<taxonomicName id="CEF6E0DCFFB9B448FC15FC7CC54DFC21" authority="Ahrens &amp; Pham" authorityName="Ahrens &amp; Pham" authorityYear="2023" box="[990,1307,943,965]" class="Insecta" family="Scarabaeidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="kachin" status="new species" subGenus="Helioserica">
<emphasis id="3B82474DFFB9B448FC15FC7CC3A3FC20" bold="true" box="[990,1013,943,964]" italics="true" pageId="5" pageNumber="556">G.</emphasis>
(
<emphasis id="3B82474DFFB9B448FBC8FC63C44AFC21" bold="true" box="[1027,1052,944,965]" italics="true" pageId="5" pageNumber="556">H.</emphasis>
) 
<emphasis id="3B82474DFFB9B448FBE1FC7CC438FC21" bold="true" box="[1066,1134,943,965]" italics="true" pageId="5" pageNumber="556">kachin</emphasis>
Ahrens &amp; Pham
</taxonomicName>
, 
<taxonomicNameLabel id="20B1FA36FFB9B448FAEDFC63C5CBFC21" box="[1318,1437,943,965]" pageId="5" pageNumber="556" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
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