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<document id="1C420BBBC1162F760D07E16A1203B13C" ID-CLB-Dataset="21840" ID-DOI="10.5852/ejt.2012.13" ID-GBIF-Dataset="9691a997-b6e4-4d47-9082-bbcba7347a3e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3857775" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590516738498" checkinUser="carolina" docAuthor="Ryvkin, Alexandr B." docDate="2012" docId="803F87FDC01DFFFEFF1CAE3EFAA299CB" docLanguage="en" docName="ejt-2012-13.pdf.imf" docOrigin="European Journal of Taxonomy 13" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Stenus canaliculatus Gyllenhal 1827" docType="key" docVersion="11" lastPageNumber="52" masterDocId="7C06FF85C02CFFCDFFA1AD48FFD59832" masterDocTitle="New species and records of Stenus (Nestus) of the canaliculatus group, with the erection of a new species group (Insecta: Coleoptera: Staphylinidae: Steninae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="50" updateTime="1698841419189" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9ACD5153841061452718950658AA9D91">New species and records of Stenus (Nestus) of the canaliculatus group, with the erection of a new species group (Insecta: Coleoptera: Staphylinidae: Steninae)</mods:title>
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Key to the Palaearctic species of the
<taxonomicName id="CF964D68C01DFFFCFD25AE3EFCF29BA3" ID-CoL="5297K" authorityName="Gyllenhal" authorityYear="1827" box="[644,807,886,913]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C01DFFFCFD25AE3EFCF29BA3" bold="true" box="[644,807,886,913]" italics="true" pageId="49" pageNumber="50">canaliculatus</emphasis>
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group
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1. Front with deep longitudinal impressions and high median elevation in between. Seventh abdominal tergite varnish shining, without ground-sculpture between punctures. Puncturation of pronotum and elytra fairly deep and coarse, average diameter of punctures on pronotum larger than basal cross of antennal segment 3. Disk of pronotum with deep longitudinal furrow rather broad and not lineshaped throughout. Internal tooth of each mandible shifted distally, directed mediodorsally of the main plane. Abdominal sternite 9 of both males and females (valvifera) with posterolateral teeth very short. Aedeagus small (see
<figureCitation id="90AD2A6EC01DFFFCFDC7A930FD6B9CA0" box="[614,702,1144,1170]" captionStart="Fig" captionStartId="41.[189,232,1803,1829]" captionTargetBox="[239,1339,1025,1762]" captionTargetId="figure@41.[206,1398,1001,1771]" captionTargetPageId="41" captionText="Fig. 7. ♂ copulative structures. AI, K: Aedeagus, ventral view. J: Abdominal sternite 9. A. Stenus (Nestus) delitor sp. nov. (holotype). B. S. (N.) illusor Ryvkin, 1987 (Russia: Amur Area: Angelokit River). CH: S. (N.) latipennis J. Sahlberg, 1880. C. (Russia: SW Taimyr: Ladannakh Lake). D. (Russia: Taimyr: Kotuy River). E. (Russia: Putorana: Ayan Lake). FG. (Russia: Evenkia: Krutenkiy Stream). H. (Russia: Magadan Area: Aborigen). IJ. S. (N.) sphaerops Casey, 1884 (U.S.A.: New York: Albany Co.). K. S. (N.) melanopus Marsham, 1802 (Ukraine: Odessa Area: Krinichnoye). Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857789" httpUri="https://zenodo.org/record/3857789/files/figure.png" pageId="49" pageNumber="50">Fig. 7K</figureCitation>
).
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……………………………………………… ……………………………………………………[
<taxonomicName id="CF964D68C01DFFFCFCE7A9D3FC109C87" ID-CoL="529GC" authorityName="Marsham" authorityYear="1802" box="[838,965,1179,1205]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="melanopus">
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<emphasis id="3AE2EAF9C01DFFFCFB82A9D3FB779C87" box="[1059,1186,1179,1205]" italics="true" pageId="49" pageNumber="50">melanopus</emphasis>
(
<bibRefCitation id="6C074B1AC01DFFFCFB13A9D3FABB9C87" author="Marsham" box="[1202,1390,1179,1205]" firstAuthor="Marsham" pageId="49" pageNumber="50" refId="ref42860" refString="Marsham T. 1802. Entomologia britannica, sistens insecta, Britanniae indigena, secundum methodum Linnaeanum disposita. Tomus I. Coleoptera: I-XXXI + 1 - 549. Wilks &amp; Taylor, Londini." type="book" year="1802">Marsham, 1802</bibRefCitation>
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If front with deep longitudinal impressions and 7
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abdominal tergite between punctures without regular ground sculpture, puncturation of pronotum very fine and dense with average diameter of punctures distinctly smaller than basal cross of antennal segment 3. Longitudinal median furrow of pronotum more or less deep but always long, line-shaped. Internal tooth of each mandible placed a little before the middle, directed only slightly dorsally of the main plane. Abdominal sternite 9 of both males and females (valvifera) with posterolateral teeth normally developed (see
<figureCitation id="90AD2A6EC01DFFFCFB4BA8C4FAFB9D95" box="[1258,1326,1420,1447]" captionStart="Fig" captionStartId="32.[189,232,1851,1877]" captionTargetBox="[249,1336,263,1808]" captionTargetId="figure@32.[244,1344,255,1810]" captionTargetPageId="32" captionText="Fig. 5. ♂ copulative structures. AE: Abdominal sternite 9. A. Stenus (Nestus) latipennis J. Sahlberg, 1880 (Russia: Krasnoyarsk Territory: Mirnoye). B. S. (N.) illusor Ryvkin, 1987 (Russia: Magadan Area: 2 km W of Splavnaya). C. S. (N.) delitor sp. nov. (holotype). D. S. (N.) canosus sp. nov. (holotype). E. S. (N.) canalis sp. nov. (holotype). F. S. (N.) caseyi Puthz, 1972 (U.S.A.: Michigan: Berrien Co.). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857785" httpUri="https://zenodo.org/record/3857785/files/figure.png" pageId="49" pageNumber="50">Fig. 5</figureCitation>
AE). Aedeagus large ………………………………………………………………[
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<emphasis id="3AE2EAF9C01DFFFCFBD3A8F8FAC59DF8" box="[1138,1296,1456,1482]" italics="true" pageId="49" pageNumber="50">canaliculatus</emphasis>
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group] 2
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<keyLead id="B36795DEC01DFFFCFF1CA8B9FAA29E39" box="[189,1399,1521,1548]" pageId="49" pageNumber="50">2. Front with distinct lateral impressions and median elevation in between …………………………3</keyLead>
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<paragraph id="082936EBC01DFFFCFF1CAB5DFAA29E60" blockId="49.[189,1399,1521,1618]" pageId="49" pageNumber="50">
<keyLead id="B36795DEC01DFFFCFF1CAB5DFAA29E60" pageId="49" pageNumber="50"> Front feebly convex or feebly concave, with median elevation and lateral depressions rather vague or entirely absent ……………………………………………………………………………8</keyLead>
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<keyStep id="B3622E4EC01DFFFCFF1CAB31FAA29F54" pageId="49" pageNumber="50">
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3. Median elevation of front broad and knob-like. Puncturation of pronotum very fine and dense, average diameter of punctures distinctly smaller than basal cross of antennal segment 3. Disk of abdominal tergite 7 without regular ground sculpture or with reticulation very feeble and vague. Aedeagus as in
<figureCitation id="90AD2A6EC01DFFFCFEB1ABAAFEBF9ECF" box="[272,362,1762,1789]" captionStart="Fig" captionStartId="8.[189,232,1899,1925]" captionTargetBox="[214,1367,264,1867]" captionTargetId="figure@8.[189,1383,238,1873]" captionTargetPageId="8" captionText="Fig. 1. AD: Stenus (Nestus) nitens Stephens, 1833: ♂ genitalia, ventral view. AB: (Russia: Moscow Area: Gorenki). CD: (Russia: Evenkia: Stolbovaya River). A, C. Aedeagus. B, D. Apical part of median lobe. E. S. (N.) labilis Erichson, 1840: ♂ (Russia: S Yamal: nr. Salekhard). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857777" httpUri="https://zenodo.org/record/3857777/files/figure.png" pageId="49" pageNumber="50">Fig. 1E</figureCitation>
.
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………………………………………………………
<taxonomicName id="CF964D68C01DFFFCFBD2ABAAFAA29ECF" ID-CoL="6ZNTT" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[1139,1399,1762,1789]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="labilis">
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<keyLead id="B36795DEC01DFFFCFF1CAA4EFAA29F54" pageId="49" pageNumber="50"> Median elevation of front ridge-shaped, angular in its cross section, flanked by broader and less sharp lateral impressions. Puncturation of pronotum coarser, average diameter of punctures about as large as or larger than basal cross of antennal segment 3 ………………………………………4</keyLead>
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4. Apical abdominal tergites without ground sculpture. For aedeagus, see
<bibRefCitation id="6C074B1AC01DFFFCFB95AAC5FB6C9F95" author="Puthz" box="[1076,1209,1933,1959]" firstAuthor="Puthz" pageId="49" pageNumber="50" pagination="171 - 196" refId="ref44527" refString="Puthz V. 2006. Ein Dutzend neuer palaarktischer Stenus - Arten (Coleoptera, Staphylinidae). 290. Beitrag zur Kenntnis der Steninen. Entomologische Blatter fur Biologie und Systematik der Kafer 101 (2005) (2 - 3): 171 - 196." type="journal article" year="2006">Puthz 2006</bibRefCitation>
(fig. 4). 2.94.0 mm …………………………………………………………………………
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<emphasis id="3AE2EAF9C01DFFFCFBFCAAFAFB369FF9" box="[1117,1251,1970,1995]" italics="true" pageId="49" pageNumber="50">immigratus</emphasis>
<bibRefCitation id="6C074B1AC01DFFFCFB4BAAF9FAA29FF9" author="Puthz" box="[1258,1399,1969,1995]" firstAuthor="Puthz" pageId="49" pageNumber="50" pagination="171 - 196" refId="ref44527" refString="Puthz V. 2006. Ein Dutzend neuer palaarktischer Stenus - Arten (Coleoptera, Staphylinidae). 290. Beitrag zur Kenntnis der Steninen. Entomologische Blatter fur Biologie und Systematik der Kafer 101 (2005) (2 - 3): 171 - 196." type="journal article" year="2006">Puthz, 2006</bibRefCitation>
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At least tergites 710 distinctly reticulated …………………………………………………………5
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<paragraph id="082936EBC01EFFFFFF1CAC05FAA299B8" blockId="50.[189,1399,333,430]" pageId="50" pageNumber="51">
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5. Elytral suture about 1/41/5 longer than pronotum. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFBEBAC05FB409955" box="[1098,1173,333,359]" captionStart="Fig" captionStartId="41.[189,232,1803,1829]" captionTargetBox="[239,1339,1025,1762]" captionTargetId="figure@41.[206,1398,1001,1771]" captionTargetPageId="41" captionText="Fig. 7. ♂ copulative structures. AI, K: Aedeagus, ventral view. J: Abdominal sternite 9. A. Stenus (Nestus) delitor sp. nov. (holotype). B. S. (N.) illusor Ryvkin, 1987 (Russia: Amur Area: Angelokit River). CH: S. (N.) latipennis J. Sahlberg, 1880. C. (Russia: SW Taimyr: Ladannakh Lake). D. (Russia: Taimyr: Kotuy River). E. (Russia: Putorana: Ayan Lake). FG. (Russia: Evenkia: Krutenkiy Stream). H. (Russia: Magadan Area: Aborigen). IJ. S. (N.) sphaerops Casey, 1884 (U.S.A.: New York: Albany Co.). K. S. (N.) melanopus Marsham, 1802 (Ukraine: Odessa Area: Krinichnoye). Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857789" httpUri="https://zenodo.org/record/3857789/files/figure.png" pageId="50" pageNumber="51">Fig. 7</figureCitation>
CH. 4.0
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…………………………………………………………………………
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J.
<bibRefCitation id="6C074B1AC01EFFFFFB64AC38FAA299B8" author="Sahlberg" box="[1221,1399,368,394]" firstAuthor="Sahlberg" pageId="50" pageNumber="51" pagination="1 - 115" refId="ref45272" refString="Sahlberg J. R. 1880. Bidrag till Nordvestra Sibiriens Insektfauna. Coleoptera. Insamlade under Expeditionerna till Obi och Jenissej 1876 och 1877. I. Cicindelidae, Carabidae, Dytiscidae, Hydrophilidae, Gyrinidae, [...] Staphylinidae och Micropepeplidae. Kongliga Svenska Vetenskaps- Akademiens Handlingar 17 (4): 1 - 115." type="journal article" year="1880">Sahlberg, 1880</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="082936EBC01EFFFFFF1CACDCFAA2999F" blockId="50.[189,1399,333,430]" box="[189,1399,403,430]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CACDCFAA2999F" box="[189,1399,403,430]" pageId="50" pageNumber="51"> Elytral suture not more than l/7 longer than pronotum ……………………………………………6</keyLead>
</paragraph>
</keyStep>
<keyStep id="B3622E4EC01EFFFFFF1CAC9CFAA29A20" pageId="50" pageNumber="51">
<paragraph id="082936EBC01EFFFFFF1CAC9CFAA299DD" blockId="50.[189,1399,468,530]" box="[189,1399,468,495]" pageId="50" pageNumber="51">
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6. Elytral suture a bit shorter than pronotum.Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFC23AC9DFC3299DC" box="[898,999,468,495]" captionStart="Fig" captionStartId="38.[189,232,1956,1982]" captionTargetBox="[177,1408,266,1913]" captionTargetId="figure@38.[161,1420,248,1934]" captionTargetPageId="38" captionText="Fig. 6. Aedeagus, ventral view. A. Stenus (Nestus) delitor sp. nov. (holotype). B. S. (N.) sphaerops Casey, 1884 (U.S.A.: New York: Albany Co.). Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857787" httpUri="https://zenodo.org/record/3857787/files/figure.png" pageId="50" pageNumber="51">Figs. 6A</figureCitation>
,
<figureCitation id="90AD2A6EC01EFFFFFC51AC9DFBC299DD" box="[1008,1047,469,495]" captionStart="Fig" captionStartId="41.[189,232,1803,1829]" captionTargetBox="[239,1339,1025,1762]" captionTargetId="figure@41.[206,1398,1001,1771]" captionTargetPageId="41" captionText="Fig. 7. ♂ copulative structures. AI, K: Aedeagus, ventral view. J: Abdominal sternite 9. A. Stenus (Nestus) delitor sp. nov. (holotype). B. S. (N.) illusor Ryvkin, 1987 (Russia: Amur Area: Angelokit River). CH: S. (N.) latipennis J. Sahlberg, 1880. C. (Russia: SW Taimyr: Ladannakh Lake). D. (Russia: Taimyr: Kotuy River). E. (Russia: Putorana: Ayan Lake). FG. (Russia: Evenkia: Krutenkiy Stream). H. (Russia: Magadan Area: Aborigen). IJ. S. (N.) sphaerops Casey, 1884 (U.S.A.: New York: Albany Co.). K. S. (N.) melanopus Marsham, 1802 (Ukraine: Odessa Area: Krinichnoye). Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857789" httpUri="https://zenodo.org/record/3857789/files/figure.png" pageId="50" pageNumber="51">7A</figureCitation>
. 4.0
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<taxonomicName id="CF964D68C01EFFFFFB6DAC9DFACF99DD" ID-CoL="8Y9ZD" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[1228,1306,469,495]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="delitor" status="sp. nov.">
<emphasis id="3AE2EAF9C01EFFFFFB6DAC9DFACF99DD" box="[1228,1306,469,495]" italics="true" pageId="50" pageNumber="51">delitor</emphasis>
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</paragraph>
<paragraph id="082936EBC01EFFFFFF1CACB0FAA29A20" blockId="50.[189,1399,468,530]" box="[189,1399,504,530]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CACB0FAA29A20" box="[189,1399,504,530]" pageId="50" pageNumber="51"> Elytral suture distinctly longer than pronotum ……………………………………………………7</keyLead>
</paragraph>
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7. Elytra more than 1/4 broader than head. Puncturation less coarse, much more regular; interspaces between punctures along the midline of anterior visible tergites distinctly wider than average diameter of punctures. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFD13AFC8FCDF9AA8" box="[690,778,640,666]" captionStart="Fig" captionStartId="13.[189,232,1852,1878]" captionTargetBox="[244,1294,261,1809]" captionTargetId="figure@13.[219,1366,255,1824]" captionTargetPageId="13" captionText="Fig. 2. AB: Stenus (Nestus) raddei Ryvkin, 1987: ♂ (Russia: Magadan Area: Sibit-Tyhellakh). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. CD: S. (N.) illotulus Puthz, 1972: ♂ (Russia: Amur Area: Zeyskiy Nature Reserve). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. Right paramere removed. E. S. (N.) confusus J. Sahlberg, 1876: ♂ (Russia: Kalinin Area: Putilovo). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857779" httpUri="https://zenodo.org/record/3857779/files/figure.png" pageId="50" pageNumber="51">Fig. 2E</figureCitation>
. 3.44.0 mm ……………
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<emphasis id="3AE2EAF9C01EFFFFFB9BAFC8FB749AA8" box="[1082,1185,640,666]" italics="true" pageId="50" pageNumber="51">confusus</emphasis>
J.
<bibRefCitation id="6C074B1AC01EFFFFFB67AFC8FAA29AA8" author="Sahlberg" box="[1222,1399,640,666]" firstAuthor="Sahlberg" pageId="50" pageNumber="51" pagination="1 - 248" refId="ref45221" refString="Sahlberg J. R. 1876. Enumeratio Coleopterorum Brachelytrorum Fenniae. Systematisk forteckning ofver de inom Finlands naturalhistoriska omrade hittils funna Coleoptera Brachelytra jemte uppgift om arternas utbredning och &lt;...&gt;. I. Staphylinidae. Acta Societatis pro Fauna et Flora Fennica 1: 1 - 248." type="journal article" year="1876">Sahlberg, 1876</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="082936EBC01EFFFFFF1CAFEBFAA29B31" blockId="50.[189,1399,569,772]" pageId="50" pageNumber="51">
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Elytra by 1/41/5 broader than head. Puncturation much coarser, distinctly irregular; interspaces between punctures along the midline of anterior visible tergites smaller than average diameter of punctures. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFD8BAFA1FD519B36" box="[554,644,745,772]" captionStart="Fig" captionStartId="41.[189,232,1803,1829]" captionTargetBox="[239,1339,1025,1762]" captionTargetId="figure@41.[206,1398,1001,1771]" captionTargetPageId="41" captionText="Fig. 7. ♂ copulative structures. AI, K: Aedeagus, ventral view. J: Abdominal sternite 9. A. Stenus (Nestus) delitor sp. nov. (holotype). B. S. (N.) illusor Ryvkin, 1987 (Russia: Amur Area: Angelokit River). CH: S. (N.) latipennis J. Sahlberg, 1880. C. (Russia: SW Taimyr: Ladannakh Lake). D. (Russia: Taimyr: Kotuy River). E. (Russia: Putorana: Ayan Lake). FG. (Russia: Evenkia: Krutenkiy Stream). H. (Russia: Magadan Area: Aborigen). IJ. S. (N.) sphaerops Casey, 1884 (U.S.A.: New York: Albany Co.). K. S. (N.) melanopus Marsham, 1802 (Ukraine: Odessa Area: Krinichnoye). Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857789" httpUri="https://zenodo.org/record/3857789/files/figure.png" pageId="50" pageNumber="51">Fig. 7B</figureCitation>
.
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………………………………
<taxonomicName id="CF964D68C01EFFFFFB25AFA1FAA29B31" ID-CoL="529DC" authority="Ryvkin, 1987" authorityName="Ryvkin" authorityYear="1987" box="[1156,1399,745,771]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="illusor">
<emphasis id="3AE2EAF9C01EFFFFFB25AFA1FB049B31" box="[1156,1233,745,771]" italics="true" pageId="50" pageNumber="51">illusor</emphasis>
<bibRefCitation id="6C074B1AC01EFFFFFB79AFA1FAA29B31" author="Ryvkin" box="[1240,1399,745,771]" firstAuthor="Ryvkin" pageId="50" pageNumber="51" pagination="149 - 161" refId="ref44999" refString="Ryvkin A. B. 1987. New species of Stenus Latreille 1796 from the Far East of the Soviet Union (Insecta: Coleoptera: Staphylinidae). Senckenbergiana biologica 68 (1 / 3): 149 - 161." type="journal article" year="1987">Ryvkin, 1987</bibRefCitation>
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<paragraph id="082936EBC01EFFFFFF1CAE63FAA29BB9" blockId="50.[189,1399,811,943]" pageId="50" pageNumber="51">
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8. Elytra not longer than pronotum, nearly as broad as head. Abdominal tergites with very dense and deep ground-sculpture. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFCF7AE06FC089B5A" box="[854,989,846,872]" captionStart="Fig" captionStartId="13.[189,232,1852,1878]" captionTargetBox="[244,1294,261,1809]" captionTargetId="figure@13.[219,1366,255,1824]" captionTargetPageId="13" captionText="Fig. 2. AB: Stenus (Nestus) raddei Ryvkin, 1987: ♂ (Russia: Magadan Area: Sibit-Tyhellakh). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. CD: S. (N.) illotulus Puthz, 1972: ♂ (Russia: Amur Area: Zeyskiy Nature Reserve). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. Right paramere removed. E. S. (N.) confusus J. Sahlberg, 1876: ♂ (Russia: Kalinin Area: Putilovo). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857779" httpUri="https://zenodo.org/record/3857779/files/figure.png" pageId="50" pageNumber="51">Fig. 2C, D</figureCitation>
.
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(the latter value for specimens with the abdomen extended) ……………………………………
<taxonomicName id="CF964D68C01EFFFFFBDBAE39FAA29BB9" ID-CoL="529DB" authority="Puthz, 1972" authorityName="Puthz" authorityYear="1972" box="[1146,1399,881,907]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="illotulus">
<emphasis id="3AE2EAF9C01EFFFFFBDBAE39FB0F9BB9" box="[1146,1242,881,907]" italics="true" pageId="50" pageNumber="51">illotulus</emphasis>
<bibRefCitation id="6C074B1AC01EFFFFFB47AE39FAA29BB9" author="Puthz V." box="[1254,1399,881,907]" pageId="50" pageNumber="51" pagination="239 - 292" refId="ref43931" refString="Puthz V. 1972 a. Zur Staphylinidenfauna des Balkans: die bisher aus Jugoslawien und angrenzenden Landern bekannten Steninen (Coleoptera. Staphylinidae). 63. [83!] Beitrag zur Kenntnis der Steninen. Wissenschafliche Mitteilungen des Bosnisch-herzegovinischen Landesmuseums 1 (C): 239 - 292." type="book chapter" year="1972">Puthz, 1972</bibRefCitation>
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<keyLead id="B36795DEC01EFFFFFF1CAEDDFAA29B9C" box="[189,1399,916,943]" pageId="50" pageNumber="51"> Elytra longer than pronotum, broader than head …………………………………………………9</keyLead>
</paragraph>
</keyStep>
<keyStep id="B3622E4EC01EFFFFFF1CAE9EFAA39C6B" pageId="50" pageNumber="51">
<paragraph id="082936EBC01EFFFFFF1CAE9EFAA29C04" blockId="50.[189,1399,982,1114]" pageId="50" pageNumber="51">
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9. Front as a whole feebly concave, with median elevation very vague, nearly flat, situated below internal margin level of eye; the upper surface except abdominal tergite 10 without ground sculpture. For aedeagus, see
<bibRefCitation id="6C074B1AC01EFFFFFE6AA954FD8D9C04" author="Puthz" box="[459,600,1052,1078]" firstAuthor="Puthz" pageId="50" pageNumber="51" pagination="45 - 48" refId="ref44353" refString="Puthz V. 1987. Ein neuer Stenus (Col. Staphylinidae) aus Japan. 208. Beitrag zur Kenntnis der Steninen. Mitteilungen des Internationalen Entomologischen Vereins, Frankfurt a. M. 11 (2 / 3): 45 - 48." type="journal article" year="1987">Puthz, 1987</bibRefCitation>
(Abb. 1).
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…………………………
<taxonomicName id="CF964D68C01EFFFFFB2CA954FAA29C04" ID-CoL="8YB4G" authority="Puthz, 1987" authorityName="Puthz" authorityYear="1987" box="[1165,1399,1052,1078]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="shogun">
<emphasis id="3AE2EAF9C01EFFFFFB2CA954FB379C04" box="[1165,1250,1052,1078]" italics="true" pageId="50" pageNumber="51">shogun</emphasis>
<bibRefCitation id="6C074B1AC01EFFFFFB4BA954FAA29C04" author="Puthz" box="[1258,1399,1052,1078]" firstAuthor="Puthz" pageId="50" pageNumber="51" pagination="45 - 48" refId="ref44353" refString="Puthz V. 1987. Ein neuer Stenus (Col. Staphylinidae) aus Japan. 208. Beitrag zur Kenntnis der Steninen. Mitteilungen des Internationalen Entomologischen Vereins, Frankfurt a. M. 11 (2 / 3): 45 - 48." type="journal article" year="1987">Puthz, 1987</bibRefCitation>
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<paragraph id="082936EBC01EFFFFFF1CA908FAA39C6B" blockId="50.[189,1399,982,1114]" box="[189,1398,1087,1114]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CA908FAA39C6B" box="[189,1398,1087,1114]" pageId="50" pageNumber="51"> If front as a whole concave, at least abdominal tergites 78 with evident ground sculpture ……10</keyLead>
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<keyStep id="B3622E4EC01EFFFFFF1CA9C9FAA39D1A" pageId="50" pageNumber="51">
<paragraph id="082936EBC01EFFFFFF1CA9C9FAA29CD3" blockId="50.[189,1399,1153,1320]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CA9C9FAA29CD3" pageId="50" pageNumber="51">
10. Front with median elevation and lateral depressions rather feeble but evident. Median parts of anterior abdominal tergites with puncturation very small, scattered, and shallow. For aedeagus, see
<bibRefCitation id="6C074B1AC01EFFFFFE44A98FFD7A9CD0" author="Renkonen" box="[485,687,1223,1250]" firstAuthor="Renkonen" pageId="50" pageNumber="51" pagination="27 - 32" refId="ref44719" refString="Renkonen O. 1935. Zwei neue Arten der Gattung Stenus (Col., Staphylinidae) aus Finnland. Annales Entomologici Fennici 1 (1): 27 - 32." type="journal article" year="1935">Renkonen, 1935</bibRefCitation>
(fig. 1). 3.0
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……………
<taxonomicName id="CF964D68C01EFFFFFBCBA98FFAA29CD3" ID-CoL="529Q5" authority="Casey, 1884" authorityName="Casey" authorityYear="1884" box="[1130,1399,1223,1249]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vinnulus">
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<bibRefCitation id="6C074B1AC01EFFFFFB7FA98FFAA29CD3" author="Casey" box="[1246,1399,1223,1249]" firstAuthor="Casey" pageId="50" pageNumber="51" refId="ref40664" refString="Casey Th. L. 1884. Revision of the Stenini of America north of Mexico. Insects of the family Staphylinidae, order Coleoptera. Collins Printing House, Philadelphia. 206 pp. http: // dx. doi. org / 10.5962 / bhl. title. 9208" type="book" year="1884">Casey, 1884</bibRefCitation>
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Front with median elevation entirely absent or, occasionally (in
<taxonomicName id="CF964D68C01EFFFFFB8FA9A3FABC9D36" ID-CoL="529HH" authority="Stephens, 1833" authorityName="Stephens" authorityYear="1833" box="[1070,1385,1258,1285]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C01EFFFFFB8FA9A3FB4C9D36" box="[1070,1177,1259,1285]" italics="true" pageId="50" pageNumber="51">S. nitens</emphasis>
<bibRefCitation id="6C074B1AC01EFFFFFB08A9A2FABC9D36" author="Stephens" box="[1193,1385,1258,1284]" firstAuthor="Stephens" pageId="50" pageNumber="51" refId="ref46202" refString="Stephens J. F. 1833. Illustrations of British Entomology; or a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V: 241 - 304. Baldwin and Cradock, London. http: // dx. doi. org / 10.5962 / bhl. title. 8133" type="book" year="1833">Stephens, 1833</bibRefCitation>
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), represented by narrow, smooth longitudinal strip ………………………………………………11
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<paragraph id="082936EBC01EFFFFFF1CA807FAA29DBE" blockId="50.[189,1399,1359,1491]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CA807FAA29DBE" pageId="50" pageNumber="51">11. Elytra with fairly small and dense puncturation. Anterior margin of labrum with a small median notch ………………………………………………………………………………………………12</keyLead>
</paragraph>
<paragraph id="082936EBC01EFFFFFF1CA8DEFAA29DE1" blockId="50.[189,1399,1359,1491]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CA8DEFAA29DE1" pageId="50" pageNumber="51"> Elytra with puncturation larger and more or less sparse. Anterior margin of labrum without distinct median notch, at most feebly sinuate …………………………………………………15</keyLead>
</paragraph>
</keyStep>
<keyStep id="B3622E4EC01EFFFFFF1CA8B2FAA29E69" pageId="50" pageNumber="51">
<paragraph id="082936EBC01EFFFFFF1CA8B2FAA29E0A" blockId="50.[189,1399,1530,1627]" pageId="50" pageNumber="51">
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12. Pronotum 1/71/8 longer than broad. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFC22A8B2FBDE9E27" box="[899,1035,1530,1557]" captionStart="Fig" captionStartId="23.[189,232,1887,1913]" captionTargetBox="[213,1378,257,1865]" captionTargetId="figure@23.[189,1388,238,1875]" captionTargetPageId="23" captionText="Fig. 3. AC: Stenus (Nestus) alopex sp. nov.: ♂. A. Aedeagus, ventral view (holotype). B. Apical part of aedeagus, ventral view (holotype). C. Abdominal sternite 9 (paratype). DE: S. (N.) canaliculatus Gyllenhal, 1827: ♂. D. Aedeagus, ventral view (Russia: Rostov Area: Nedvigovka). E. Abdominal sternite 9 (Russia: Taimyr: Maymecha River). Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857781" httpUri="https://zenodo.org/record/3857781/files/figure.png" pageId="50" pageNumber="51">Fig. 3A, B</figureCitation>
.
<quantity id="CF6E9B0EC01EFFFFFBBBA8B2FB7E9E27" box="[1050,1195,1530,1557]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.9" metricValueMin="3.2" pageId="50" pageNumber="51" unit="mm" value="3.55" valueMax="3.9" valueMin="3.2">3.23.9 mm</quantity>
(the latter value for specimens with abdomen extended) …………………………………………
<taxonomicName id="CF964D68C01EFFFFFB1BAB55FADD9E05" ID-CoL="8Y9Z4" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[1210,1288,1565,1591]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="alopex" status="sp. nov.">
<emphasis id="3AE2EAF9C01EFFFFFB1BAB55FADD9E05" box="[1210,1288,1565,1591]" italics="true" pageId="50" pageNumber="51">alopex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C01EFFFFFAB7AB56FAA29E0A" box="[1302,1399,1566,1592]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="082936EBC01EFFFFFF1CAB09FAA29E69" blockId="50.[189,1399,1530,1627]" box="[189,1399,1600,1627]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CAB09FAA29E69" box="[189,1399,1600,1627]" pageId="50" pageNumber="51"> Pronotum not, or only a little, longer than broad …………………………………………………13</keyLead>
</paragraph>
</keyStep>
<keyStep id="B3622E4EC01EFFFFFF1CABCAFAA39F34" pageId="50" pageNumber="51">
<paragraph id="082936EBC01EFFFFFF1CABCAFAA29ED0" blockId="50.[189,1399,1666,1798]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CABCAFAA29ED0" pageId="50" pageNumber="51">
13. Entire body with ground sculpture more or less visible. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFBD3ABCAFB019EAE" box="[1138,1236,1666,1692]" captionStart="Fig" captionStartId="23.[189,232,1887,1913]" captionTargetBox="[213,1378,257,1865]" captionTargetId="figure@23.[189,1388,238,1875]" captionTargetPageId="23" captionText="Fig. 3. AC: Stenus (Nestus) alopex sp. nov.: ♂. A. Aedeagus, ventral view (holotype). B. Apical part of aedeagus, ventral view (holotype). C. Abdominal sternite 9 (paratype). DE: S. (N.) canaliculatus Gyllenhal, 1827: ♂. D. Aedeagus, ventral view (Russia: Rostov Area: Nedvigovka). E. Abdominal sternite 9 (Russia: Taimyr: Maymecha River). Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857781" httpUri="https://zenodo.org/record/3857781/files/figure.png" pageId="50" pageNumber="51">Fig. 3D</figureCitation>
. 3.0
<quantity id="CF6E9B0EC01EFFFFFAB9ABCAFAA29EAE" box="[1304,1399,1666,1693]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="50" pageNumber="51" unit="mm" value="4.5">4.5 mm</quantity>
(the latter value for the American specimen with abdomen extended, the maximum value for Palaearctic specimens known to me:
<quantity id="CF6E9B0EC01EFFFFFD02AB80FD2B9ED1" box="[675,766,1736,1763]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="50" pageNumber="51" unit="mm" value="4.3">4.3 mm</quantity>
) ………………………
<taxonomicName id="CF964D68C01EFFFFFBB4AB80FAA29ED0" ID-CoL="5297K" authority="Gyllenhal, 1827" authorityName="Gyllenhal" authorityYear="1827" box="[1045,1399,1736,1762]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="3AE2EAF9C01EFFFFFBB4AB80FB669ED0" box="[1045,1203,1736,1762]" italics="true" pageId="50" pageNumber="51">canaliculatus</emphasis>
<bibRefCitation id="6C074B1AC01EFFFFFB1BAB80FAA29ED0" author="Gyllenhal" box="[1210,1399,1736,1762]" firstAuthor="Gyllenhal" pageId="50" pageNumber="51" refId="ref41615" refString="Gyllenhal L. 1827. Insecta Suecica descripta. Classis I: Coleoptera sive Eleuterata. Vol. 1. ps. IV: i-viii + 1 - 762. Friederichum Fleischer, Lipsiae. http: // dx. doi. org / 10.5962 / bhl. title. 8767" type="book" year="1827">Gyllenhal, 1827</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="082936EBC01EFFFFFF1CABA4FAA39F34" blockId="50.[189,1399,1666,1798]" box="[189,1398,1771,1798]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CABA4FAA39F34" box="[189,1398,1771,1798]" pageId="50" pageNumber="51"> At least median part of abdominal tergites 37 without traces of microsculpture ………………14</keyLead>
</paragraph>
</keyStep>
<keyStep id="B3622E4EC01EFFFFFF1CAA65FAA29FE6" pageId="50" pageNumber="51">
<paragraph id="082936EBC01EFFFFFF1CAA65FAA29F59" blockId="50.[189,1399,1837,2005]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CAA65FAA29F59" pageId="50" pageNumber="51">
14. Elytra between punctures with ground sculpture throughout. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFB15AA65FA959F75" box="[1204,1344,1837,1863]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[228,1365,265,1852]" captionTargetId="figure@28.[217,1370,253,1873]" captionTargetPageId="28" captionText="Fig. 4. AB: Stenus (Nestus) canosus sp. nov.: ♂ (holotype). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. CD: S. (N.) canalis sp. nov.: ♂ (holotype). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. [Note that the difference in shape of the apical lobe compared to that in C is due to the level of magnification.] Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857783" httpUri="https://zenodo.org/record/3857783/files/figure.png" pageId="50" pageNumber="51">Fig. 4A, B</figureCitation>
. 4.0 mm (with abdomen extended) …………………………………………………
<taxonomicName id="CF964D68C01EFFFFFB06AA19FAD39F58" ID-CoL="8Y9Z9" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[1191,1286,1873,1898]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="canosus" status="sp. nov.">
<emphasis id="3AE2EAF9C01EFFFFFB06AA19FAD39F58" box="[1191,1286,1873,1898]" italics="true" pageId="50" pageNumber="51">canosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C01EFFFFFAB5AA19FAA29F59" box="[1300,1399,1873,1899]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="082936EBC01EFFFFFF1CAA3CFAA29FE6" blockId="50.[189,1399,1837,2005]" pageId="50" pageNumber="51">
<keyLead id="B36795DEC01EFFFFFF1CAA3CFAA29FE6" pageId="50" pageNumber="51">
Elytra between punctures with ground sculpture mainly in humeral impressions and near suture. Aedeagus as in
<figureCitation id="90AD2A6EC01EFFFFFDB4AADEFD4A9F83" box="[533,671,1942,1969]" captionStart="Fig" captionStartId="28.[189,232,1895,1921]" captionTargetBox="[228,1365,265,1852]" captionTargetId="figure@28.[217,1370,253,1873]" captionTargetPageId="28" captionText="Fig. 4. AB: Stenus (Nestus) canosus sp. nov.: ♂ (holotype). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. CD: S. (N.) canalis sp. nov.: ♂ (holotype). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. [Note that the difference in shape of the apical lobe compared to that in C is due to the level of magnification.] Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857783" httpUri="https://zenodo.org/record/3857783/files/figure.png" pageId="50" pageNumber="51">Fig. 4C, D</figureCitation>
.
<quantity id="CF6E9B0EC01EFFFFFD11AADFFC969F83" box="[688,835,1943,1969]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.7" metricValueMin="2.8" pageId="50" pageNumber="51" unit="mm" value="3.25" valueMax="3.7" valueMin="2.8">2.83.7 mm</quantity>
(the latter value for specimen with abdomen extended) …………………………………………………………………………
<taxonomicName id="CF964D68C01EFFFFFB0DAAF2FAD49FE6" ID-CoL="8Y9Z7" authority="Ryvkin, 2012" authorityName="Ryvkin" authorityYear="2012" box="[1196,1281,1978,2004]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="canalis" status="sp. nov.">
<emphasis id="3AE2EAF9C01EFFFFFB0DAAF2FAD49FE6" box="[1196,1281,1978,2004]" italics="true" pageId="50" pageNumber="51">canalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21D15782C01EFFFFFAB3AAF2FAA29FE6" box="[1298,1399,1978,2004]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B3622E4EC01FFFFEFF1CAC43FAA299CB" pageId="51" pageNumber="52">
<paragraph id="082936EBC01FFFFEFF1CAC43FAA299BD" blockId="51.[189,1399,267,505]" pageId="51" pageNumber="52">
<keyLead id="B36795DEC01FFFFEFF1CAC43FAA299BD" pageId="51" pageNumber="52">
15. Middle of front punctured much sparser than lateral parts, occasionally with a small longitudinal strip or spot broader than the average diameter of punctures. Elytra only a bit (l/14th to l/15th) broader than head, being punctured larger and sparser. Aedeagus as in
<figureCitation id="90AD2A6EC01FFFFEFBF2AC1AFB43995E" box="[1107,1174,338,364]" captionStart="Fig" captionStartId="8.[189,232,1899,1925]" captionTargetBox="[214,1367,264,1867]" captionTargetId="figure@8.[189,1383,238,1873]" captionTargetPageId="8" captionText="Fig. 1. AD: Stenus (Nestus) nitens Stephens, 1833: ♂ genitalia, ventral view. AB: (Russia: Moscow Area: Gorenki). CD: (Russia: Evenkia: Stolbovaya River). A, C. Aedeagus. B, D. Apical part of median lobe. E. S. (N.) labilis Erichson, 1840: ♂ (Russia: S Yamal: nr. Salekhard). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857777" httpUri="https://zenodo.org/record/3857777/files/figure.png" pageId="51" pageNumber="52">Fig. 1</figureCitation>
AD. 3.44.0 mm (the latter value for specimens with the abdomen extended) …………………
<taxonomicName id="CF964D68C01FFFFEFBD6AC3EFAA299BD" ID-CoL="529HH" authority="Stephens, 1833" authorityName="Stephens" authorityYear="1833" box="[1143,1399,373,399]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="3AE2EAF9C01FFFFEFBD6AC3EFB6899BD" box="[1143,1213,374,399]" italics="true" pageId="51" pageNumber="52">nitens</emphasis>
<bibRefCitation id="6C074B1AC01FFFFEFB65AC3DFAA299BD" author="Stephens" box="[1220,1399,373,399]" firstAuthor="Stephens" pageId="51" pageNumber="52" refId="ref46202" refString="Stephens J. F. 1833. Illustrations of British Entomology; or a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V: 241 - 304. Baldwin and Cradock, London. http: // dx. doi. org / 10.5962 / bhl. title. 8133" type="book" year="1833">Stephens, 1833</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="082936EBC01FFFFEFF1CACD1FAA299CB" blockId="51.[189,1399,267,505]" pageId="51" pageNumber="52">
<keyLead id="B36795DEC01FFFFEFF1CACD1FAA299CB" pageId="51" pageNumber="52">
Front with puncturation more regular, without evident smooth strips or spots. Elytra l/7 to l/10 broader than head, being smaller and denser punctured. Aedeagus as in
<figureCitation id="90AD2A6EC01FFFFEFBF4ACF3FB0C99E4" box="[1109,1241,443,470]" captionStart="Fig" captionStartId="13.[189,232,1852,1878]" captionTargetBox="[244,1294,261,1809]" captionTargetId="figure@13.[219,1366,255,1824]" captionTargetPageId="13" captionText="Fig. 2. AB: Stenus (Nestus) raddei Ryvkin, 1987: ♂ (Russia: Magadan Area: Sibit-Tyhellakh). A. Aedeagus, ventral view. B. Apical part of aedeagus, ventral view. CD: S. (N.) illotulus Puthz, 1972: ♂ (Russia: Amur Area: Zeyskiy Nature Reserve). C. Aedeagus, ventral view. D. Apical part of aedeagus, ventral view. Right paramere removed. E. S. (N.) confusus J. Sahlberg, 1876: ♂ (Russia: Kalinin Area: Putilovo). Aedeagus, ventral view. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3857779" httpUri="https://zenodo.org/record/3857779/files/figure.png" pageId="51" pageNumber="52">Fig. 2A, B</figureCitation>
.
<quantity id="CF6E9B0EC01FFFFEFB46ACF3FAA299E4" box="[1255,1399,443,470]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.1" metricValueMin="3.1" pageId="51" pageNumber="52" unit="mm" value="3.5999999999999996" valueMax="4.1" valueMin="3.1">3.14.1 mm</quantity>
(the latter value for specimens with the abdomen extended) ………………
<taxonomicName id="CF964D68C01FFFFEFBDFAC97FAA299CB" ID-CoL="8Y9ZZ" authority="Ryvkin, 1987" authorityName="Ryvkin" authorityYear="1987" box="[1150,1399,478,505]" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="raddei">
<emphasis id="3AE2EAF9C01FFFFEFBDFAC97FB1E99CB" box="[1150,1227,479,505]" italics="true" pageId="51" pageNumber="52">raddei</emphasis>
<bibRefCitation id="6C074B1AC01FFFFEFB74AC96FAA299CB" author="Ryvkin" box="[1237,1399,478,505]" firstAuthor="Ryvkin" pageId="51" pageNumber="52" pagination="149 - 161" refId="ref44999" refString="Ryvkin A. B. 1987. New species of Stenus Latreille 1796 from the Far East of the Soviet Union (Insecta: Coleoptera: Staphylinidae). Senckenbergiana biologica 68 (1 / 3): 149 - 161." type="journal article" year="1987">Ryvkin, 1987</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>