treatments-xml/data/7C/78/3A/7C783A27FFE90C44FF78F8C2ED418615.xml

<document id="C590EF93E15A9E5A62923E81A354C387" ID-CLB-Dataset="29388" ID-DOI="10.11646/zootaxa.4462.4.10" ID-GBIF-Dataset="c2abe59c-39ee-449f-9585-ede35c9f91fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441797" ID-ZooBank="8463393B-B671-4851-9C64-60A36D0C0182" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538404794315" checkinUser="plazi" docAuthor="Nguyen, Lien Thi Phuong, Nguyen, Ha Thi Thu &amp; Bozdoğan, Hakan" docDate="2018" docId="7C783A27FFE90C44FF78F8C2ED418615" docLanguage="en" docName="zootaxa.4462.4.10.pdf" docOrigin="Zootaxa 4462 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Okinawepipona yty Nguyen &amp; Nguyen &amp; Bozdoğan 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="595" masterDocId="8041425FFFE90C47FFEFFF92E9138151" masterDocTitle="Contribution to the genus Okinawepipona Yamane (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species" masterLastPageNumber="596" masterPageNumber="592" pageNumber="592" updateTime="1698561220577" updateUser="ExternalLinkService">
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<mods:title id="B9CE9F14EE42C3C9832200B0E80D8794">Contribution to the genus Okinawepipona Yamane (Hymenoptera: Vespidae: Eumeninae) from Vietnam, with description of a new species</mods:title>
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<mods:namePart id="562FD9DD1526AA873C3F56CAFE5CF725">Nguyen, Lien Thi Phuong</mods:namePart>
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<mods:namePart id="BC2C1A1077BCB3359314273789F9A929">Nguyen, Ha Thi Thu</mods:namePart>
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<subSubSection id="BCCBD8BAFFE90C47FF78F8C2E81986DA" pageId="0" pageNumber="592" type="nomenclature">
<paragraph id="F46E8B31FFE90C47FF78F8C2EB41863B" blockId="0.[151,594,1872,1932]" box="[151,594,1872,1898]" pageId="0" pageNumber="592">
<heading id="AF263C5DFFE90C47FF78F8C2EB41863B" bold="true" box="[151,594,1872,1898]" fontSize="11" level="1" pageId="0" pageNumber="592" reason="1">
<emphasis id="C6A55723FFE90C47FF78F8C2EB41863B" bold="true" box="[151,594,1872,1898]" pageId="0" pageNumber="592">
<taxonomicName id="33D1F0B2FFE90C47FF78F8C2E8FB863B" authority="Nguyen" authorityName="Nguyen &amp; Nguyen &amp; Bozdoğan" authorityYear="2018" box="[151,488,1872,1898]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="592" phylum="Arthropoda" rank="species" species="yty" status="sp. nov.">
<emphasis id="C6A55723FFE90C47FF78F8C2E86C863B" bold="true" box="[151,383,1872,1898]" italics="true" pageId="0" pageNumber="592">Okinawepipona yty</emphasis>
Nguyen
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, 
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(
<figureCitation id="6CEA97B4FFE90C47FF70F8E1E81286DA" box="[159,257,1907,1932]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–9. 7" captionStartId-0="2.[151,264,1948,1971]" captionStartId-1="3.[151,264,1233,1256]" captionTargetBox-0="[150,1436,267,1925]" captionTargetBox-1="[151,1436,193,1224]" captionTargetId-0="figure@2.[150,1436,267,1925]" captionTargetId-1="figure@3.[151,1436,193,1224]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." captionText-1="FIGURES 7–9. 7. Tergum I &amp; II in lateral view in Okinawepipona yty sp. nov., holotype, female. 8. O. yty, female inside the nest. 9. Completed nest of O. nigra Nguyen &amp; Xu (2014)." httpUri-0="https://zenodo.org/record/1441799/files/figure.png" httpUri-1="https://zenodo.org/record/1441801/files/figure.png" pageId="0" pageNumber="592">Figs 1–8</figureCitation>
)
</paragraph>
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<paragraph id="F46E8B31FFE90C47FF78F829EC8E8682" blockId="0.[151,1437,1979,2004]" box="[151,1437,1979,2004]" pageId="0" pageNumber="592">
<materialsCitation id="44B9816CFFE90C47FF78F829EC8E8682" ID-GBIF-Occurrence="1913893131" box="[151,1437,1979,2004]" collectionCode="IEBR" collectorName="Y Ty &amp; Bat Xat" country="Vietnam" latitude="22.6205" location="Lao Cai" pageId="0" pageNumber="592" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="holotype">
<emphasis id="C6A55723FFE90C47FF78F829E8938685" bold="true" box="[151,384,1979,2004]" pageId="0" pageNumber="592">Material examined.</emphasis>
<typeStatus id="2B6A3593FFE90C47FE69F829E8E28685" box="[390,497,1979,2004]" pageId="0" pageNumber="592" type="holotype">Holotype</typeStatus>
, 
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, pinned (deposited in 
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), 
<collectingCountry id="8CC6CBA1FFE90C47FCB0F829EAF08682" box="[863,995,1979,2003]" name="Vietnam" pageId="0" pageNumber="592">VIETNAM</collectingCountry>
, 
<collectorName id="5924EEE7FFE90C47FC01F82EED348685" box="[1006,1063,1980,2004]" pageId="0" pageNumber="592">Y Ty</collectorName>
, 
<collectorName id="5924EEE7FFE90C47FBDEF82EED998682" box="[1073,1162,1980,2003]" pageId="0" pageNumber="592">Bat Xat</collectorName>
, 
<collectingRegion id="361545D3FFE90C47FB7AF82EEDFD8685" box="[1173,1262,1979,2004]" country="Vietnam" name="Lao Cai" pageId="0" pageNumber="592">Lao Cai</collectingRegion>
(
<geoCoordinate id="91E5EDF6FFE90C47FB12F829EC848682" box="[1277,1431,1979,2003]" direction="north" orientation="latitude" pageId="0" pageNumber="592" precision="1" value="22.6205">22°37'13.8''N</geoCoordinate>
,
</materialsCitation>
</paragraph>
<paragraph id="F46E8B31FFE80C46FF78FF0AED9D8184" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="593">
<materialsCitation id="44B9816CFFE80C46FF78FF0AED9D8184" ID-GBIF-Occurrence="1913893132" collectingDate="2017-08-14" collectionCode="IEBR, VNMN" collectorName="L. T. P. Nguyen &amp; C. Q. Nguyen" country="Vietnam" elevation="1850" latitude="22.6205" location="Lao Cai" longitude="103.62353" pageId="1" pageNumber="593" specimenCount="6" specimenCount-female="6" stateProvince="Lao Cai" typeStatus="paratype">
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), alt. 
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, 
<collectingDate id="902B5419FFE80C46FE0DFF05EB7F81E1" box="[482,620,151,176]" pageId="1" pageNumber="593" value="2017-08-14">
<date id="806FADF1FFE80C46FE0DFF05EB7F81E1" box="[482,620,151,176]" pageId="1" pageNumber="593" value="2017-08-14">14.viii.2017</date>
</collectingDate>
, 
<collectorName id="5924EEE7FFE80C46FD97FF0AEA0D81E1" box="[632,798,152,176]" pageId="1" pageNumber="593">L.T.P. Nguyen</collectorName>
, 
<collectorName id="5924EEE7FFE80C46FCC5FF0AEAD381E1" box="[810,960,152,176]" pageId="1" pageNumber="593">C.Q. Nguyen</collectorName>
&amp; Q.H. Bui. 
<typeStatus id="2B6A3593FFE80C46FBB3FF0AEDDD81E1" box="[1116,1230,152,176]" pageId="1" pageNumber="593" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="8CC6CBA1FFE80C46FB36FF0AEC4B81E1" box="[1241,1368,152,176]" name="Vietnam" pageId="1" pageNumber="593">VIETNAM</collectingCountry>
: 
<specimenCount id="E2D740B8FFE80C46FA85FF05EC8681FE" box="[1386,1429,151,176]" pageId="1" pageNumber="593" type="female">6 ♀</specimenCount>
, same data as holotype (3 deposited in 
<collectionCode id="92C013F4FFE80C46FDADFF2FEB968185" box="[578,645,189,212]" pageId="1" pageNumber="593">IEBR</collectionCode>
, 3 deposited in 
<collectionCode id="92C013F4FFE80C46FCDBFF2FEA9A8185" box="[820,905,189,212]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="1" pageNumber="593">VNMN</collectionCode>
(
<collectionCode id="92C013F4FFE80C46FC77FF2EEAE18184" box="[920,1010,188,213]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="1" pageNumber="593">VNMN</collectionCode>
.0104–0106)).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BCCBD8BAFFE80C46FF28FF4DEDF180D9" pageId="1" pageNumber="593" type="diagnosis">
<paragraph id="F46E8B31FFE80C46FF28FF4DEDF180D9" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="593">
<emphasis id="C6A55723FFE80C46FF28FF4DE85281A9" bold="true" box="[199,321,223,248]" pageId="1" pageNumber="593">Diagnosis.</emphasis>
This species can be distinguished from other species of genus 
<taxonomicName id="33D1F0B2FFE80C46FBCDFF72EDC081A9" authorityName="Nguyen &amp; Nguyen &amp; Bozdoğan" authorityYear="2018" box="[1058,1235,224,248]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="C6A55723FFE80C46FBCDFF72EDC081A9" box="[1058,1235,224,248]" italics="true" pageId="1" pageNumber="593">Okinawepipona</emphasis>
</taxonomicName>
by the following combination of characters: propodeum with posterior surface dull, rugose, with oblique striae along the median carina; T 
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lateral view strongly convex anteriorly and clearly concave dorsally; metasoma black except a narrow orange band at the apical margins of metasomal tergum I and II (incised in the middle); and clypeus with three yellow spots, two at lateral basal margin and the other at the lower middle, near the apical margin.
</paragraph>
</subSubSection>
<subSubSection id="BCCBD8BAFFE80C44FF28FE06E8968719" lastPageId="3" lastPageNumber="595" pageId="1" pageNumber="593" type="description">
<paragraph id="F46E8B31FFE80C46FF28FE06E82B8299" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="593">
<emphasis id="C6A55723FFE80C46FF28FE06E84A80FC" bold="true" box="[199,345,404,429]" pageId="1" pageNumber="593">Description.</emphasis>
<emphasis id="C6A55723FFE80C46FE8DFE06E8A580FD" box="[354,438,404,428]" italics="true" pageId="1" pageNumber="593">Female</emphasis>
. Body length 
<quantity id="332926D4FFE80C46FDB2FE06EBC480FD" box="[605,727,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="1" pageNumber="593" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
(
<typeStatus id="2B6A3593FFE80C46FD07FE06EA4380FC" box="[744,848,404,429]" pageId="1" pageNumber="593" type="holotype">holotype</typeStatus>
: 
<quantity id="332926D4FFE80C46FCB2FE06EAD480FD" box="[861,967,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="1" pageNumber="593" unit="mm" value="11.8">11.8 mm</quantity>
); fore wing length 
<quantity id="332926D4FFE80C46FB4AFE06EC3380FD" box="[1189,1312,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="1" pageNumber="593" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
(
<typeStatus id="2B6A3593FFE80C46FADEFE06EC8A80FC" box="[1329,1433,404,429]" pageId="1" pageNumber="593" type="holotype">holotype</typeStatus>
: 
<quantity id="332926D4FFE80C46FF78FE2AE8108081" box="[151,259,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="593" unit="mm" value="11.7">11.7 mm</quantity>
). Head in frontal view subcircular, wider than high, about 1.1× as wide as high (
<figureCitation id="6CEA97B4FFE80C46FB2EFE25EDED8081" box="[1217,1278,439,464]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig.1</figureCitation>
). Vertex with cephalic foveae small, bearing dense pubescence, situated very close to each other, almost touching to each other; depression for cephalic foveae conspicuous (
<figureCitation id="6CEA97B4FFE80C46FD63FE6DEBC28349" box="[652,721,511,536]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 2</figureCitation>
). Distance from posterior ocelli to apical margin of vertex more than 1.7× distance from posterior ocelli to inner eye margin (
<figureCitation id="6CEA97B4FFE80C46FCADFDB6EA95836D" box="[834,902,548,573]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 2</figureCitation>
). Gena much narrower than eye, in lateral view about 0.7× as wide as eye. Occipital carina complete, present along entire length of gena, but dorsally somewhat weak. Inner eye margins strongly convergent ventrally; in frontal view about 1.3× further apart from each other at vertex than at clypeus. Disc of clypeus in lateral view weakly convex at basal half, then straight to near apical margin; in frontal view about as wide as high (
<figureCitation id="6CEA97B4FFE80C46FD5FFD26EBE0839D" box="[688,755,692,717]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 1</figureCitation>
), with basal margin slightly convex medially and distinctly separated from antennal sockets; apical margin deeply emarginate medially, forming sharp tooth on each lateral side (
<figureCitation id="6CEA97B4FFE80C46FF35FD6EE8318245" box="[218,290,764,789]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 1</figureCitation>
), with two faint carina running from the tooth at apical point to the base direction; width of emargination slightly less than 1/3 width of clypeus between inner eye margins. Mandible with prominent teeth, second and third teeth with inner side produced with round margin, fourth tooth pointed apically. Antennal scape about 3.7× as long as its maximum width; flagellomere I about 1.5× as long as wide, flagellomere II slightly longer than wide, III-IV as wide as long, V–IX wider than long, terminal flagellomere bullet-shaped, as long as its basal width (
<figureCitation id="6CEA97B4FFE80C46FF09FC3DE8398299" box="[230,298,943,968]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="F46E8B31FFE80C46FF28FC47E875845D" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="593">
Mesosoma longer than wide in dorsal view. Pronotal carina slightly raised, produced as humeral angles, reaching ventral corner of pronotum. Mesoscutum weakly convex, slightly longer than wide between tegulae; anterior margin broadly rounded. Disc of scutellum almost flat, in profile slightly higher than level of apical margin of mesoscutum (
<figureCitation id="6CEA97B4FFE80C46FEB4FBADE8B78509" box="[347,420,1087,1112]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 5</figureCitation>
). Metanotum convex, sloping down to apical margin. Propodeum excavated medially, posterior surface dull, rugose, basal triangular area with basal fovea, at lower end of which median carina runs to apical margin, basal fovea greater than 1/3 of length of median carina, median carina developed for entire length of propodeum (
<figureCitation id="6CEA97B4FFE80C46FEC6FB3EE8618595" box="[297,370,1196,1221]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 4</figureCitation>
); upper part of propodeum slightly produced; dorsal surface of propodeum strongly rugose; dorsal and posterior surfaces bordered by a blunt edge. Hind coxa with distinct carina, which is elevated to form triangular process.
</paragraph>
<paragraph id="F46E8B31FFE80C46FF28FA85EB1584B5" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="593">
Metasomal tergum I narrower than tergum II, truncate at base (
<figureCitation id="6CEA97B4FFE80C46FC4EFA85EAFB8461" box="[929,1000,1303,1328]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 6</figureCitation>
); anterior vertical surface of tergum I strongly convex, with sparse shallow punctures, clearly separated from posterior horizontal surface, but not by a carina; posterior horizontal surface of tergum I in lateral view clearly concave dorsaly (
<figureCitation id="6CEA97B4FFE80C46FB95FACDEDD38429" box="[1146,1216,1375,1400]" captionStart="FIGURES 7–9. 7" captionStartId="3.[151,264,1233,1256]" captionTargetBox="[151,1436,193,1224]" captionTargetId="figure@3.[151,1436,193,1224]" captionTargetPageId="3" captionText="FIGURES 7–9. 7. Tergum I &amp; II in lateral view in Okinawepipona yty sp. nov., holotype, female. 8. O. yty, female inside the nest. 9. Completed nest of O. nigra Nguyen &amp; Xu (2014)." httpUri="https://zenodo.org/record/1441801/files/figure.png" pageId="1" pageNumber="593">Fig. 7</figureCitation>
); tergum I laterally divided by sharp carina into upper and lower parts. Tergum I in dorsal view slightly greater than twice as wide as long; tergum II in dorsal view nearly as wide as long (
<figureCitation id="6CEA97B4FFE80C46FCFCFA35EA498491" box="[787,858,1447,1472]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1948,1971]" captionTargetBox="[150,1436,267,1925]" captionTargetId="figure@2.[150,1436,267,1925]" captionTargetPageId="2" captionText="FIGURES 1–6. Okinawepipona yty sp. nov., holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm." httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="1" pageNumber="593">Fig. 6</figureCitation>
); sternum II in lateral view gradually and slightly convex to apical margin (
<figureCitation id="6CEA97B4FFE80C46FE5BFA5EE8EB84B5" box="[436,504,1484,1509]" captionStart="FIGURES 7–9. 7" captionStartId="3.[151,264,1233,1256]" captionTargetBox="[151,1436,193,1224]" captionTargetId="figure@3.[151,1436,193,1224]" captionTargetPageId="3" captionText="FIGURES 7–9. 7. Tergum I &amp; II in lateral view in Okinawepipona yty sp. nov., holotype, female. 8. O. yty, female inside the nest. 9. Completed nest of O. nigra Nguyen &amp; Xu (2014)." httpUri="https://zenodo.org/record/1441801/files/figure.png" pageId="1" pageNumber="593">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="F46E8B31FFE80C45FF28FA7DEACF8185" blockId="1.[151,1437,151,2013]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="594" pageId="1" pageNumber="593">Body covered with long silvery setae except metasoma covered with short silvery setae. Clypeus with dense, large, flat-bottomed punctures, each bearing silver bristle, punctures at center larger than at sides, space between punctures larger than puncture diameter. Mandible with several shallow small punctures. Frons densely covered with coarse punctures, interspaces between punctures slightly raised to form reticulation. Vertex and gena with coarse punctures. Pronotum with punctures slightly coarser than those on frons, spaces between punctures narrow, strongly raised to form reticulation. Mesocutum densely and coarsely covered with flat-bottomed punctures, punctures near apical margin with interspaces showing tendency to run into irregular longitudinal striae. Punctures on scutellum similar to those near apical margin of mesoscutum. Punctures on metanotum denser and smaller than those on scutellum. Mesepisternum with deep and large punctures posterodorsally, shallow and small punctures anteroventrally; border between posterodorsal and anteroventral parts indistinct. Metapleuron almost smooth in dorsal area, with sparse shallow punctures in ventral area. Propodeum with punctures on dorsal surface very coarse, spaces between punctures strongly raised to form reticulation, lateral faces with weak striae, posterior surface dull, rugose and with some weak and short oblique striae along median carina. Metasomal terga I–II densely covered with punctures, punctures on tergum I coarser and denser than those on tergum II, punctures on terga III–V much smaller and weaker than those on terga I–II; tergum VI with minute punctures; punctures on sternum II stronger and larger than those on sternum III–V, sternum VI with minute punctures.</paragraph>
<caption id="A0AEDBB9FFEB0C45FF78F80EEC4F8683" httpUri="https://zenodo.org/record/1441799/files/figure.png" pageId="2" pageNumber="594" startId="2.[151,264,1948,1971]" targetBox="[150,1436,267,1925]" targetPageId="2">
<paragraph id="F46E8B31FFEB0C45FF78F80EEC4F8683" blockId="2.[151,1437,1948,2002]" pageId="2" pageNumber="594">
<emphasis id="C6A55723FFEB0C45FF78F80EE82986E3" bold="true" box="[151,314,1948,1971]" pageId="2" pageNumber="594">FIGURES 1–6.</emphasis>
<taxonomicName id="33D1F0B2FFEB0C45FED0F80FEB1086E2" authority="Nguyen &amp; Nguyen &amp; Bozdoğan, 2018" authorityName="Nguyen &amp; Nguyen &amp; Bozdoğan" authorityYear="2018" box="[319,515,1949,1971]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="594" phylum="Arthropoda" rank="species" species="yty" status="sp. nov.">
<emphasis id="C6A55723FFEB0C45FED0F80FEB1086E2" box="[319,515,1949,1971]" italics="true" pageId="2" pageNumber="594">Okinawepipona yty</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD96EA58FFEB0C45FDE6F80CEB4A86E2" box="[521,601,1950,1971]" pageId="2" pageNumber="594" rank="species">
<emphasis id="C6A55723FFEB0C45FDE6F80CEB4A86E2" bold="true" box="[521,601,1950,1971]" pageId="2" pageNumber="594">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female.1. Head in frontal view. 2.Vertex in dorsal view. 3. Left antenna. 4. Metanotum and propodeum in dorsal view. 5. Thorax in lateral view. 6. Tergum I &amp; II in dorsal view. Scale bar: 1mm.
</paragraph>
</caption>
<caption id="A0AEDBB9FFEA0C44FF78FB43EBF18457" httpUri="https://zenodo.org/record/1441801/files/figure.png" pageId="3" pageNumber="595" startId="3.[151,264,1233,1256]" targetBox="[151,1436,193,1224]" targetPageId="3">
<paragraph id="F46E8B31FFEA0C44FF78FB43EBF18457" blockId="3.[151,1436,1233,1286]" pageId="3" pageNumber="595">
<emphasis id="C6A55723FFEA0C44FF78FB43E82885B6" bold="true" box="[151,315,1233,1256]" pageId="3" pageNumber="595">FIGURES 7–9.</emphasis>
7. Tergum I &amp; II in lateral view in 
<taxonomicName id="33D1F0B2FFEA0C44FD48FB40EA7F85B9" authority="Nguyen &amp; Nguyen &amp; Bozdoğan, 2018" authorityName="Nguyen &amp; Nguyen &amp; Bozdoğan" authorityYear="2018" box="[679,876,1234,1256]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="595" phylum="Arthropoda" rank="species" species="yty" status="sp. nov.">
<emphasis id="C6A55723FFEA0C44FD48FB40EA7F85B9" box="[679,876,1234,1256]" italics="true" pageId="3" pageNumber="595">Okinawepipona yty</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD96EA58FFEA0C44FC9CFB40EAD785B6" box="[883,964,1234,1255]" pageId="3" pageNumber="595" rank="species">
<emphasis id="C6A55723FFEA0C44FC9CFB40EAD785B6" bold="true" box="[883,964,1234,1255]" pageId="3" pageNumber="595">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female. 8. 
<taxonomicName id="33D1F0B2FFEA0C44FB4BFB40EDF385B9" authorityName="Nguyen &amp; Nguyen &amp; Bozdoğan" authorityYear="2018" box="[1188,1248,1234,1256]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="595" phylum="Arthropoda" rank="species" species="yty">
<emphasis id="C6A55723FFEA0C44FB4BFB40EDF385B9" box="[1188,1248,1234,1256]" italics="true" pageId="3" pageNumber="595">O. yty</emphasis>
</taxonomicName>
, female inside the nest. 9. Completed nest of 
<taxonomicName id="33D1F0B2FFEA0C44FE4AFB63EBCE8457" authority="Nguyen &amp; Xu (2014)" authorityName="Nguyen &amp; Xu" authorityYear="2014" box="[421,733,1264,1286]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="595" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="C6A55723FFEA0C44FE4AFB63E8E88457" box="[421,507,1265,1286]" italics="true" pageId="3" pageNumber="595">O. nigra</emphasis>
Nguyen &amp; Xu (2014)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="F46E8B31FFEA0C44FF28FAA1E8C78775" blockId="3.[151,1436,1331,1860]" pageId="3" pageNumber="595">
<emphasis id="C6A55723FFEA0C44FF28FAA1E814841A" box="[199,263,1331,1355]" italics="true" pageId="3" pageNumber="595">Color</emphasis>
. Black; following parts yellow: large spots on upper lateral corner and a large spot on lower middle of clypeus; narrow band along inner eye margin extending from bottom of frons nearly to ocular sinus, large spot between antennal sockets, mandible except margins, antennal scape beneath; following parts orange: thick band along pronotal carina in dorsal part of pronotum, narrow band at apical margin of terga I-II (incised in the middle, and narrowed toward lateral side of tergum I), a small spot at the apical corners of sternum II. Legs black except stripes on outer surfaces of fore femur yellow. Propodeal valvulae dark brown. Wings dark brown, slightly infuscate, veins dark brown.
</paragraph>
<paragraph id="F46E8B31FFEA0C44FF28F9BDE8968719" blockId="3.[151,1436,1331,1860]" box="[199,389,1583,1608]" pageId="3" pageNumber="595">
<emphasis id="C6A55723FFEA0C44FF28F9BDE8128716" box="[199,257,1583,1607]" italics="true" pageId="3" pageNumber="595">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="BCCBD8BAFFEA0C44FF28F9C1EB53873D" box="[199,576,1619,1644]" pageId="3" pageNumber="595" type="distribution">
<paragraph id="F46E8B31FFEA0C44FF28F9C1EB53873D" blockId="3.[151,1436,1331,1860]" box="[199,576,1619,1644]" pageId="3" pageNumber="595">
<emphasis id="C6A55723FFEA0C44FF28F9C1E873873D" bold="true" box="[199,352,1619,1644]" pageId="3" pageNumber="595">Distribution.</emphasis>
<collectingCountry id="8CC6CBA1FFEA0C44FE88F9C1E8D5873D" box="[359,454,1619,1644]" name="Vietnam" pageId="3" pageNumber="595">Vietnam</collectingCountry>
(
<collectingRegion id="361545D3FFEA0C44FE3AF9C6EB27873D" box="[469,564,1619,1644]" country="Vietnam" name="Lao Cai" pageId="3" pageNumber="595">Lao Cai</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="BCCBD8BAFFEA0C44FF28F9E5E8B287E5" pageId="3" pageNumber="595" type="etymology">
<paragraph id="F46E8B31FFEA0C44FF28F9E5E8B287E5" blockId="3.[151,1436,1331,1860]" pageId="3" pageNumber="595">
<emphasis id="C6A55723FFEA0C44FF28F9E5E85D87C1" bold="true" box="[199,334,1655,1680]" pageId="3" pageNumber="595">Etymology.</emphasis>
The specific name refers to the 
<typeStatus id="2B6A3593FFEA0C44FD5BF9EAEBF787C1" box="[692,740,1656,1680]" pageId="3" pageNumber="595">type</typeStatus>
locality, Y Ty, 
<collectingRegion id="361545D3FFEA0C44FC61F9EAED4287C1" box="[910,1105,1655,1680]" country="Vietnam" name="Lao Cai" pageId="3" pageNumber="595">Lao Cai province</collectingRegion>
of 
<collectingCountry id="8CC6CBA1FFEA0C44FB9AF9E5EDC487C1" box="[1141,1239,1655,1680]" name="Vietnam" pageId="3" pageNumber="595">Vietnam</collectingCountry>
; it is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="BCCBD8BAFFEA0C44FF28F92DED418615" pageId="3" pageNumber="595" type="biology_ecology">
<paragraph id="F46E8B31FFEA0C44FF28F92DED418615" blockId="3.[151,1436,1331,1860]" pageId="3" pageNumber="595">
<emphasis id="C6A55723FFEA0C44FF28F92DE80C8789" bold="true" box="[199,287,1727,1752]" pageId="3" pageNumber="595">Biology</emphasis>
. The species made its nest inside a hole of a woody stick (
<figureCitation id="6CEA97B4FFEA0C44FC41F92DEAE28786" box="[942,1009,1727,1752]" captionStart="FIGURES 7–9. 7" captionStartId="3.[151,264,1233,1256]" captionTargetBox="[151,1436,193,1224]" captionTargetId="figure@3.[151,1436,193,1224]" captionTargetPageId="3" captionText="FIGURES 7–9. 7. Tergum I &amp; II in lateral view in Okinawepipona yty sp. nov., holotype, female. 8. O. yty, female inside the nest. 9. Completed nest of O. nigra Nguyen &amp; Xu (2014)." httpUri="https://zenodo.org/record/1441801/files/figure.png" pageId="3" pageNumber="595">Fig. 8</figureCitation>
), which was used to make a house for drying medicine plants in the forest. The same character was observed in 
<taxonomicName id="33D1F0B2FFEA0C44FC30F971EC6C87AA" authority="Nguyen &amp; Xu" authorityName="Nguyen &amp; Xu" authorityYear="2014" box="[991,1407,1763,1788]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="595" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="C6A55723FFEA0C44FC30F971EDC587AA" box="[991,1238,1763,1787]" italics="true" pageId="3" pageNumber="595">Okinawepipona nigra</emphasis>
Nguyen &amp; Xu
</taxonomicName>
at Bat Dai Son natural reserve, 
<collectingRegion id="361545D3FFEA0C44FE0DF89AEBA98671" box="[482,698,1799,1824]" country="Vietnam" name="Ha Giang" pageId="3" pageNumber="595">Ha Giang province</collectingRegion>
of 
<collectingCountry id="8CC6CBA1FFEA0C44FD0DF895EA568671" box="[738,837,1799,1824]" name="Vietnam" pageId="3" pageNumber="595">Vietnam</collectingCountry>
. This species made its nest inside a hole of a woody door. After completing the nest, 
<taxonomicName id="33D1F0B2FFEA0C44FDEDF8BEEB728612" authorityName="Nguyen &amp; Xu" authorityYear="2014" box="[514,609,1836,1859]" class="Insecta" family="Vespidae" genus="Okinawepipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="595" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="C6A55723FFEA0C44FDEDF8BEEB728612" box="[514,609,1836,1859]" italics="true" pageId="3" pageNumber="595">O. nigra</emphasis>
</taxonomicName>
closed the hole with white material (
<figureCitation id="6CEA97B4FFEA0C44FBEFF8B9ED578615" box="[1024,1092,1835,1860]" captionStart="FIGURES 7–9. 7" captionStartId="3.[151,264,1233,1256]" captionTargetBox="[151,1436,193,1224]" captionTargetId="figure@3.[151,1436,193,1224]" captionTargetPageId="3" captionText="FIGURES 7–9. 7. Tergum I &amp; II in lateral view in Okinawepipona yty sp. nov., holotype, female. 8. O. yty, female inside the nest. 9. Completed nest of O. nigra Nguyen &amp; Xu (2014)." httpUri="https://zenodo.org/record/1441801/files/figure.png" pageId="3" pageNumber="595">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>