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<mods:title id="3F79A82E840B104E2E0AFA171BACF8CC">Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)</mods:title>
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<mods:namePart id="6D580CC436D57F531C3F19AA81B80FB5">Hutchinson, Paul M.</mods:namePart>
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<treatment id="79268507FFD8FFB3CBE5D247FEE3FD9D" ID-DOI="http://doi.org/10.5281/zenodo.5750358" ID-GBIF-Taxon="190874932" ID-Zenodo-Dep="5750358" LSID="urn:lsid:plazi:treatment:79268507FFD8FFB3CBE5D247FEE3FD9D" httpUri="http://treatment.plazi.org/id/79268507FFD8FFB3CBE5D247FEE3FD9D" lastPageId="22" lastPageNumber="461" pageId="21" pageNumber="460">
<subSubSection id="B995679AFFD8FFB0CBE5D247FD85FDDE" box="[151,524,611,638]" pageId="21" pageNumber="460" type="nomenclature">
<paragraph id="F1303411FFD8FFB0CBE5D247FD85FDDE" blockId="21.[151,524,611,638]" box="[151,524,611,638]" pageId="21" pageNumber="460">
<heading id="AA78837DFFD8FFB0CBE5D247FD85FDDE" bold="true" box="[151,524,611,638]" fontSize="11" level="1" pageId="21" pageNumber="460" reason="1">
<taxonomicName id="368F4F92FFD8FFB0CBE5D247FD85FDDE" authority="Blackburn, 1889" authorityName="Blackburn" authorityYear="1889" box="[151,524,611,638]" class="Insecta" family="Dynastidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="460" phylum="Arthropoda" rank="genus">
<emphasis id="C3FBE803FFD8FFB0CBE5D247FD85FDDE" bold="true" box="[151,524,611,638]" pageId="21" pageNumber="460">
<emphasis id="C3FBE803FFD8FFB0CBE5D247FEBCFDDE" bold="true" box="[151,309,611,638]" italics="true" pageId="21" pageNumber="460">Adoryphorus</emphasis>
<bibRefCitation id="951E49E0FFD8FFB0CA4FD240FD85FDDE" author="Blackburn, T." box="[317,524,612,638]" pageId="21" pageNumber="460" pagination="1387 - 1506" refId="ref10419" refString="Blackburn, T. (1889) Further notes on Australian Coleoptera, with descriptions of new genera and species. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 1387 - 1506." type="journal article" year="1889">Blackburn, 1889</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="F1303411FFD8FFB0CBE5D28CFB5BFD61" blockId="21.[151,1234,680,705]" box="[151,1234,680,705]" pageId="21" pageNumber="460">
<treatmentCitation id="702E1200FFD8FFB0CBE5D28CFD87FD61" author="Blackburn, T." box="[151,526,680,705]" page="1412" pageId="21" pageNumber="460" year="1889">
<taxonomicName id="368F4F92FFD8FFB0CBE5D28CFD87FD61" authority="Blackburn, 1889: 1412" authorityName="Blackburn" authorityPageNumber="1412" authorityYear="1889" box="[151,526,680,705]" class="Insecta" family="Dynastidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="460" phylum="Arthropoda" rank="genus">
<emphasis id="C3FBE803FFD8FFB0CBE5D28CFE92FD60" box="[151,283,680,704]" italics="true" pageId="21" pageNumber="460">Adoryphorus</emphasis>
<bibRefCitation id="951E49E0FFD8FFB0CA50D28CFD87FD61" author="Blackburn, T." box="[290,526,680,705]" pageId="21" pageNumber="460" pagination="1387 - 1506" refId="ref10419" refString="Blackburn, T. (1889) Further notes on Australian Coleoptera, with descriptions of new genera and species. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 1387 - 1506." type="journal article" year="1889">Blackburn, 1889: 1412</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="2E348AB3FFD8FFB0C965D28DFDC0FD61" box="[535,585,681,705]" pageId="21" pageNumber="460">Type</typeStatus>
species:
<taxonomicName id="368F4F92FFD8FFB0C9D4D28CFBB2FD61" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[678,1083,680,705]" class="Insecta" family="Dynastidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="460" phylum="Arthropoda" rank="species" species="coulonii">
<emphasis id="C3FBE803FFD8FFB0C9D4D28CFC0DFD60" box="[678,900,680,704]" italics="true" pageId="21" pageNumber="460">Dasygnathus coulonii</emphasis>
Burmeister, 1847
</taxonomicName>
, by monotypy.
</paragraph>
<caption id="A5F06499FFD8FFB0CBE5D67CFB42F9D0" ID-DOI="http://doi.org/10.5281/zenodo.5749041" ID-Zenodo-Dep="5749041" box="[151,1227,1624,1648]" httpUri="https://zenodo.org/record/5749041/files/figure.png" pageId="21" pageNumber="460" startId="21.[151,264,1624,1648]" targetBox="[196,1373,738,1580]" targetPageId="21">
<paragraph id="F1303411FFD8FFB0CBE5D67CFB42F9D0" blockId="21.[151,1227,1624,1648]" box="[151,1227,1624,1648]" pageId="21" pageNumber="460">
<emphasis id="C3FBE803FFD8FFB0CBE5D67CFEDDF9D0" bold="true" box="[151,340,1624,1648]" pageId="21" pageNumber="460">FIGURES 64–65.</emphasis>
<taxonomicName id="368F4F92FFD8FFB0CA28D67CFD6CF9D0" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[346,741,1624,1648]" class="Insecta" family="Dynastidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="460" phylum="Arthropoda" rank="species" species="canei">
<emphasis id="C3FBE803FFD8FFB0CA28D67CFD95F9D0" box="[346,540,1624,1648]" italics="true" pageId="21" pageNumber="460">Adoryphorus canei</emphasis>
(Burmeister, 1847)
</taxonomicName>
.
<emphasis id="C3FBE803FFD8FFB0C99CD67CFC87F9D0" bold="true" box="[750,782,1624,1648]" pageId="21" pageNumber="460">64,</emphasis>
Male, dorsal view;
<emphasis id="C3FBE803FFD8FFB0C8AAD67CFC7EF9D0" bold="true" box="[984,1015,1624,1648]" pageId="21" pageNumber="460">65,</emphasis>
female, dorsal view.
</paragraph>
</caption>
<paragraph id="F1303411FFD8FFB3CBE5D6B3FB4AFF75" blockId="21.[151,1437,1687,2038]" lastBlockId="22.[151,1437,151,573]" lastPageId="22" lastPageNumber="461" pageId="21" pageNumber="460">
<emphasis id="C3FBE803FFD8FFB0CBE5D6B3FE38F911" bold="true" box="[151,433,1687,1713]" pageId="21" pageNumber="460">
Diagnosis (
<figureCitation id="69B42894FFD8FFB0CA6BD6B3FE2DF911" box="[281,420,1687,1713]" captionStart="FIGURES 64–65" captionStartId="21.[151,264,1624,1648]" captionTargetBox="[196,1373,738,1580]" captionTargetId="figure-173@21.[151,1435,725,1600]" captionTargetPageId="21" captionText="FIGURES 64–65. Adoryphorus canei (Burmeister, 1847). 64, Male, dorsal view; 65, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5749041" httpUri="https://zenodo.org/record/5749041/files/figure.png" pageId="21" pageNumber="460">Figs. 64–65</figureCitation>
).
</emphasis>
Body
<quantity id="367799F4FFD8FFB0CA8FD6BCFDE1F912" box="[509,616,1688,1714]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.7" metricValueMin="0.9" pageId="21" pageNumber="460" unit="mm" value="13.0" valueMax="17.0" valueMin="9.0">9–17 mm</quantity>
long, reddish black to black, ovate, low convex, head with at most a low tubercle on vertex, pronotum unarmed. Maxilla and galea reduced to small conical pieces, unarmed, terminated by pencils of sensory setae; mentum longitudinally convex, disc setose, ligula with apex rounded, palps globular and short; mandibles just visible at sides of clypeus, external margin rounded. Antennae with 9 or 10 antennomeres, club with 3 lamellae, shorter than length of shaft. Clypeus glabrous, sides straight, not or only slightly converging towards truncate apex, slightly emarginate; clypeofrontal suture slightly carinate and transverse. Pronotum evenly convex, lightly and finely punctate, anterior edge with narrow membrane not extending to anterior angles, basal ridge obsolete at least in median third, lateral margins unguttered and converging towards apex. Elytra deeply impressed and distinctly punctate, lateral margin unguttered, epipleural plane in anterior half vertical. Female penultimate abdominal ventrite with row of setae close to posterior edge. Protarsomere 5 about as long as tarsomere 1; proclaws of males strongly curved, sometimes slightly asymmetrical; metafemur slender; metatibia of female stout with apex dilated. Parameres with internal margin irregular or distinctly toothed or serrate, usually setose.
</paragraph>
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</subSubSection>
<subSubSection id="B995679AFFDBFFB3CBB5D0FAFEB1FE0D" pageId="22" pageNumber="461" type="distribution">
<paragraph id="F1303411FFDBFFB3CBB5D0FAFEB1FE0D" blockId="22.[151,1437,151,573]" pageId="22" pageNumber="461">
<emphasis id="C3FBE803FFDBFFB3CBB5D0FAFEE8FF58" bold="true" box="[199,353,222,248]" pageId="22" pageNumber="461">Distribution.</emphasis>
The genus contains three species (
<bibRefCitation id="951E49E0FFDBFFB3C987D0FBFC30FF59" author="Weir, T. A. &amp; Lawrence, J. F. &amp; Lemann, C. &amp; Gunter, N. L." box="[757,953,223,249]" pageId="22" pageNumber="461" pagination="1919" refId="ref11237" refString="Weir, T. A., Lawrence, J. F., Lemann, C. &amp; Gunter, N. L. (2019) Scarabaeidae: Dynastinae Macleay 1919 [sic]. In: Slipinski, A. &amp; Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book chapter" year="2019">
Weir
<emphasis id="C3FBE803FFDBFFB3C845D0FBFCFBFF59" box="[823,882,223,249]" italics="true" pageId="22" pageNumber="461">et al.</emphasis>
2019
</bibRefCitation>
):
<taxonomicName id="368F4F92FFDBFFB3C8BCD0FBFAA6FF59" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[974,1327,223,249]" class="Insecta" family="Dynastidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="coulonii">
<emphasis id="C3FBE803FFDBFFB3C8BCD0FBFBC5FF59" box="[974,1100,223,249]" italics="true" pageId="22" pageNumber="461">A. coulonii</emphasis>
(Burmeister, 1847)
</taxonomicName>
,
<taxonomicName id="368F4F92FFDBFFB3CE4CD0FBFEA9FEBD" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" class="Insecta" family="Dynastidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="canei">
<emphasis id="C3FBE803FFDBFFB3CE4CD0FBFA14FF58" box="[1342,1437,223,249]" italics="true" pageId="22" pageNumber="461">A. canei</emphasis>
<bibRefCitation id="951E49E0FFDBFFB3CBE5D127FEA9FEBD" author="Carne, P. B." box="[151,288,258,285]" pageId="22" pageNumber="461" refId="ref10459" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
</taxonomicName>
(
<figureCitation id="69B42894FFDBFFB3CA42D127FE3EFEBD" box="[304,439,258,285]" captionStart="FIGURES 64–65" captionStartId="21.[151,264,1624,1648]" captionTargetBox="[196,1373,738,1580]" captionTargetId="figure-173@21.[151,1435,725,1600]" captionTargetPageId="21" captionText="FIGURES 64–65. Adoryphorus canei (Burmeister, 1847). 64, Male, dorsal view; 65, female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5749041" httpUri="https://zenodo.org/record/5749041/files/figure.png" pageId="22" pageNumber="461">Figs. 64–65</figureCitation>
) and
<taxonomicName id="368F4F92FFDBFFB3CA87D127FD70FEBD" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[501,761,258,285]" class="Insecta" family="Dynastidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="mellori">
<emphasis id="C3FBE803FFDBFFB3CA87D127FDECFEBD" box="[501,613,259,285]" italics="true" pageId="22" pageNumber="461">A. mellori</emphasis>
<bibRefCitation id="951E49E0FFDBFFB3C91FD127FD70FEBD" author="Carne, P. B." box="[621,761,258,285]" pageId="22" pageNumber="461" refId="ref10459" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
</taxonomicName>
, that occur from southern Queensland to southeastern South Australia and northern Tasmania (
<bibRefCitation id="951E49E0FFDBFFB3C967D103FD15FEE1" author="Carne, P. B." box="[533,668,294,321]" pageId="22" pageNumber="461" refId="ref10459" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne 1957</bibRefCitation>
, Map 15), with
<taxonomicName id="368F4F92FFDBFFB3C83DD103FC42FEE1" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[847,971,295,321]" class="Insecta" family="Dynastidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="coulonii">
<emphasis id="C3FBE803FFDBFFB3C83DD103FC42FEE1" box="[847,971,295,321]" italics="true" pageId="22" pageNumber="461">A. coulonii</emphasis>
</taxonomicName>
having established in the South Island of
<collectingCountry id="89987481FFDBFFB3CBE5D16FFEA4FEC5" box="[151,301,331,357]" name="New Zealand" pageId="22" pageNumber="461">New Zealand</collectingCountry>
(
<bibRefCitation id="951E49E0FFDBFFB3CA4ED16FFDD8FEC5" author="Stufkens, M. W. &amp; Farrell, J. A." box="[316,593,331,357]" pageId="22" pageNumber="461" pagination="569 - 573" refId="ref11176" refString="Stufkens, M. W. &amp; Farrell, J. A. (1980) Life history and distribution of Adoryphorus couloni [sic] Burmeister (Coleoptera: Scarabaeidae) in Canterbury, New Zealand. New Zealand Journal of Agricultural Research, 23, 569 - 573. https: // doi. org / 10.1080 / 00288233.1980.10417884" type="journal article" year="1980">Stufkens &amp; Farrell 1980</bibRefCitation>
). Adults of all three species were adequately described and keyed by
<bibRefCitation id="951E49E0FFDBFFB3CE2BD16FFF6DFE29" author="Carne, P. B." pageId="22" pageNumber="461" refId="ref10459" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne (1957)</bibRefCitation>
, and
<bibRefCitation id="951E49E0FFDBFFB3CA52D14BFE7AFE29" author="Dechambre, R. - P." box="[288,499,366,393]" pageId="22" pageNumber="461" refId="ref10613" refString="Dechambre, R. - P. (2005) Dynastidae Australiens et Oceaniens. Les Coleopteres du Monde 30. Hillside Books, Canterbury, 132 pp." type="book" year="2005">Dechambre (2005)</bibRefCitation>
illustrated adults of
<taxonomicName id="368F4F92FFDBFFB3C9A8D14BFCBFFE28" authorityName="Carne" authorityYear="1957" box="[730,822,367,393]" class="Insecta" family="Dynastidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="canei">
<emphasis id="C3FBE803FFDBFFB3C9A8D14BFCBFFE28" box="[730,822,367,393]" italics="true" pageId="22" pageNumber="461">A. canei</emphasis>
</taxonomicName>
and
<emphasis id="C3FBE803FFDBFFB3C81FD14BFC66FE29" box="[877,1007,367,393]" italics="true" pageId="22" pageNumber="461">
<taxonomicName id="368F4F92FFDBFFB3C81FD14BFC62FE29" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[877,1003,367,393]" class="Insecta" family="Dynastidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="coulonii">A. coulonii</taxonomicName>
.
</emphasis>
<bibRefCitation id="951E49E0FFDBFFB3C884D14BFB48FE29" author="McQuillan, P. B." box="[1014,1217,366,393]" pageId="22" pageNumber="461" pagination="509 - 546" refId="ref11024" refString="McQuillan, P. B. (1985) The identification of root-feeding cockchafer larvae (Coleoptera: Scarabaeidae) found in pastures in Tasmania. Australian Journal of Zoology, 33, 509 - 546. https: // doi. org / 10.1071 / ZO 9850509" type="journal article" year="1985">McQuillan (1985)</bibRefCitation>
described the larva of
<emphasis id="C3FBE803FFDBFFB3CBC4D1B7FEB1FE0D" box="[182,312,403,429]" italics="true" pageId="22" pageNumber="461">
<taxonomicName id="368F4F92FFDBFFB3CBC4D1B7FEBDFE0D" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[182,308,403,429]" class="Insecta" family="Dynastidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="coulonii">A. coulonii</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFDBFFB3CBB5D193FEE3FD9D" pageId="22" pageNumber="461" type="discussion">
<paragraph id="F1303411FFDBFFB3CBB5D193FEE3FD9D" blockId="22.[151,1437,151,573]" pageId="22" pageNumber="461">
<emphasis id="C3FBE803FFDBFFB3CBB5D193FEB2FE71" bold="true" box="[199,315,439,465]" pageId="22" pageNumber="461">Remarks.</emphasis>
<bibRefCitation id="951E49E0FFDBFFB3CA36D193FE53FE71" author="Carne, P. B." box="[324,474,438,465]" pageId="22" pageNumber="461" refId="ref10459" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne (1957)</bibRefCitation>
named
<taxonomicName id="368F4F92FFDBFFB3C944D193FD1DFE70" authorityName="Carne" authorityYear="1957" box="[566,660,439,465]" class="Insecta" family="Dynastidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="461" phylum="Arthropoda" rank="species" species="canei">
<emphasis id="C3FBE803FFDBFFB3C944D193FD1DFE70" box="[566,660,439,465]" italics="true" pageId="22" pageNumber="461">A. canei</emphasis>
</taxonomicName>
for “Miss H.M. Cane (Mrs. B. Cameron)” and so should have used an -ae ending (
<bibRefCitation id="951E49E0FFDBFFB3CA30D1FFFC74FE55" author="International Commission on Zoological Nomenclature" box="[322,1021,475,501]" pageId="22" pageNumber="461" refId="ref10923" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature 1999</bibRefCitation>
, Article 31.1.2). Even though it was improperly formed, the original spelling must be preserved in line with Article 31.1.3 as it should not be corrected under Article 32.5.
</paragraph>
</subSubSection>
</treatment>
</document>
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