219 lines
26 KiB
XML
219 lines
26 KiB
XML
<document id="D212261642FE345B1A86A3272C9E2DB5" ID-DOI="10.11646/zootaxa.4750.3.8" ID-GBIF-Dataset="c26e3746-2064-4f3a-b080-0da987427e4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3707541" ID-ZooBank="4557810B-C28F-42FF-92C2-7FE13267020A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584002442594" checkinUser="plazi" docAuthor="Frolov, Andrey V. & Akhmetova, Lilia A." docDate="2020" docId="715787DEFFAE3F47FF2C01764ED7FEF6" docLanguage="en" docName="zootaxa.4750.3.8.pdf" docOrigin="Zootaxa 4750 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orphnus (Orphnus) brevialatus Frolov & Akhmetova 2020, new species" docType="treatment" docVersion="6" lastPageNumber="426" masterDocId="8D6EFFA6FFAF3F46FFBB03534F50F94D" masterDocTitle="Description of a new brachypterous scarab species, Orphnus brevialatus (Coleoptera: Scarabaeidae: Orphninae) from East Africa, with notes on flightlessness in the orphnines" masterLastPageNumber="431" masterPageNumber="425" pageNumber="426" updateTime="1698727262469" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="912200BFD5E9BA5BA3E48D392ADB45BC">
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<mods:title id="48A46EF5295D7276C410EEE0DF3D52E9">Description of a new brachypterous scarab species, Orphnus brevialatus (Coleoptera: Scarabaeidae: Orphninae) from East Africa, with notes on flightlessness in the orphnines</mods:title>
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<mods:name id="4BEAF3C69FE7B9529FBCEE0E58EF5895" type="personal">
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<mods:namePart id="3DA2F2B970892475B3D4373DCD3A2D2C">Frolov, Andrey V.</mods:namePart>
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<mods:name id="4F78330E68F7497FC366A03EE45B9196" type="personal">
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<mods:roleTerm id="C4CE62718E671FD0EEE2E9E1A7DB7038">Author</mods:roleTerm>
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<mods:namePart id="938C578B91602EE52504AEEC9E18242A">Akhmetova, Lilia A.</mods:namePart>
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<mods:typeOfResource id="4623395D02645CF30F5FD800FE5A6156">text</mods:typeOfResource>
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<mods:title id="EB84CE96BEA49E7E5A161422F048A997">Zootaxa</mods:title>
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<mods:part id="EE27C47F84B233B47C18E5E892A439A4">
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<mods:date id="EA41219AD99BB426882C71AE526AB66B">2020</mods:date>
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<mods:number id="E98AC7938B75113D5B16EF28C84438FE">2020-03-12</mods:number>
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<mods:number id="745B5EED7A3BAED28B926CE2BDA5C091">4750</mods:number>
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<mods:identifier id="993DC3FD38BF35C1E7073D053E638B7C" type="DOI">10.11646/zootaxa.4750.3.8</mods:identifier>
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<treatment id="715787DEFFAE3F47FF2C01764ED7FEF6" ID-DOI="http://doi.org/10.5281/zenodo.3717918" ID-GBIF-Taxon="162484033" ID-Zenodo-Dep="3717918" LSID="urn:lsid:plazi:treatment:715787DEFFAE3F47FF2C01764ED7FEF6" httpUri="http://treatment.plazi.org/id/715787DEFFAE3F47FF2C01764ED7FEF6" lastPageNumber="426" pageId="1" pageNumber="426">
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<subSubSection id="B1E46543FFAE3F47FF2C01764C95FB72" box="[151,965,549,576]" pageId="1" pageNumber="426" type="nomenclature">
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<paragraph id="F94136C8FFAE3F47FF2C01764C95FB72" blockId="1.[151,965,549,611]" box="[151,965,549,576]" pageId="1" pageNumber="426">
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<heading id="A20981A4FFAE3F47FF2C01764C95FB72" bold="true" box="[151,965,549,576]" fontSize="11" level="1" pageId="1" pageNumber="426" reason="1">
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<emphasis id="CB8AEADAFFAE3F47FF2C01764C95FB72" bold="true" box="[151,965,549,576]" pageId="1" pageNumber="426">
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<taxonomicName id="3EFE4D4BFFAE3F47FF2C01764C79FB72" authority="Frolov & Akhmetova" authorityName="Frolov & Akhmetova" authorityYear="2020" box="[151,809,549,576]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="species" species="brevialatus" status="sp. nov." subGenus="Orphnus">
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<emphasis id="CB8AEADAFFAE3F47FF2C01764E53FB0D" bold="true" box="[151,259,549,576]" italics="true" pageId="1" pageNumber="426">Orphnus</emphasis>
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(
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<emphasis id="CB8AEADAFFAE3F47FEAF01764ED0FB0D" bold="true" box="[276,384,549,576]" italics="true" pageId="1" pageNumber="426">Orphnus</emphasis>
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)
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<emphasis id="CB8AEADAFFAE3F47FE2A01764D47FB0D" bold="true" box="[401,535,549,576]" italics="true" pageId="1" pageNumber="426">brevialatus</emphasis>
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Frolov & Akhmetova
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</taxonomicName>
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,
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<taxonomicNameLabel id="D0B957A1FFAE3F47FC8E01764C95FB72" box="[821,965,549,575]" pageId="1" pageNumber="426" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1E46543FFAE3F47FF2C011A4E2FFB2E" box="[151,383,585,611]" pageId="1" pageNumber="426" type="description">
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<paragraph id="F94136C8FFAE3F47FF2C011A4E2FFB2E" blockId="1.[151,965,549,611]" box="[151,383,585,611]" pageId="1" pageNumber="426">
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(
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<figureCitation id="61C52A4DFFAE3F47FF24011A4FB8FB2E" box="[159,232,585,611]" captionStart="FIGURES 1" captionStartId="2.[151,264,1692,1717]" captionTargetBox="[172,1416,193,1657]" captionTargetId="figure@2.[159,1428,181,1667]" captionTargetPageId="2" captionText="FIGURES 1A–D. Orphnus brevialatus Frolov & Akhmetova, new species, habitus. A–C, holotype, male (abdomen dissected); B, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.3707543" httpUri="https://zenodo.org/record/3707543/files/figure.png" pageId="1" pageNumber="426">Figs. 1</figureCitation>
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A–D, 2A–D)
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</paragraph>
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</subSubSection>
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<subSubSection id="B1E46543FFAE3F47FF2C01C24C2BFB82" pageId="1" pageNumber="426" type="materials_examined">
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<paragraph id="F94136C8FFAE3F47FF2C01C24C2BFB82" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF2C01C24E16FBE6" bold="true" box="[151,326,656,683]" pageId="1" pageNumber="426">Type material.</emphasis>
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<materialsCitation id="49963C95FFAE3F47FEF501C24F8CFB82" ID-GBIF-Occurrence="2575868886" collectingDate="2006-12-16" collectionCode="ZIN" country="Tanzania" elevation="2500" location="Werner" municipality="Chunya" pageId="1" pageNumber="426" specimenCount="1" stateProvince="Mbeya" typeStatus="holotype">
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<typeStatus id="2645886AFFAE3F47FEF501C24EE9FBE6" box="[334,441,657,683]" pageId="1" pageNumber="426" type="holotype">Holotype</typeStatus>
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, male with the label “
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<collectingCountry id="81E97658FFAE3F47FD1501C24C6EFBE6" box="[686,830,657,683]" name="Tanzania" pageId="1" pageNumber="426">TANZANIA</collectingCountry>
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<date id="8D401008FFAE3F47FCFD01C24C9FFBE6" box="[838,975,657,683]" pageId="1" pageNumber="426" value="2006-12-16">
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<collectingDate id="9D04E9E0FFAE3F47FCFD01C24C9FFBE6" box="[838,975,657,683]" pageId="1" pageNumber="426" value="2006-12-16">16.XII.2006</collectingDate>
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</date>
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<collectingMunicipality id="1925ACB2FFAE3F47FC6C01C24B7EFBE6" box="[983,1070,657,683]" pageId="1" pageNumber="426">Chunya</collectingMunicipality>
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to
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<collectingRegion id="3B3AF82AFFAE3F47FBE801C24BF4FBE6" box="[1107,1188,657,683]" country="Tanzania" name="Mbeya" pageId="1" pageNumber="426">Mbeya</collectingRegion>
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,
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<quantity id="3E069B2DFFAE3F47FB0B01C34A56FBE6" box="[1200,1286,656,683]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="426" unit="m" value="2500.0">
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<elevation id="72D3D1FBFFAE3F47FB0B01C34A56FBE6" box="[1200,1286,656,683]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="426" unit="m" value="2500.0">2500 m</elevation>
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</quantity>
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<location id="FC216013FFAE3F47FAB601C24A0FFBE6" LSID="urn:lsid:plazi:treatment:715787DEFFAE3F47FF2C01764ED7FEF6:FC216013FFAE3F47FAB601C24A0FFBE6" box="[1293,1375,657,683]" country="Tanzania" municipality="Chunya" name="Werner" pageId="1" pageNumber="426" stateProvince="Mbeya">Werner</location>
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leg.” (
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<collectionCode id="9FEFAE0DFFAE3F47FF2401E64F83FB82" box="[159,211,693,719]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="426" type="Museum">ZIN</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="49963C95FFAE3F47FF5D01E64C27FB82" ID-GBIF-Occurrence="2575868885" box="[230,887,693,719]" collectingDate="2006-12-16" collectionCode="ZIN" country="Tanzania" elevation="2500" location="Werner" municipality="Chunya" pageId="1" pageNumber="426" specimenCount="1" stateProvince="Mbeya" typeStatus="paratype">
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<typeStatus id="2645886AFFAE3F47FF5D01E64E1BFB82" box="[230,331,693,719]" pageId="1" pageNumber="426" type="paratype">Paratype</typeStatus>
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, female with the same data as the holotype (
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<collectionCode id="9FEFAE0DFFAE3F47FC8001E64C3FFB82" box="[827,879,693,719]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="426" type="Museum">ZIN</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1E46543FFAE3F47FF7C018B4B9AFF3A" pageId="1" pageNumber="426" type="description">
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<paragraph id="F94136C8FFAE3F47FF7C018B4DD7FA5A" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF7C018B4E09FBBE" bold="true" box="[199,345,728,755]" pageId="1" pageNumber="426">Description.</emphasis>
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<typeStatus id="2645886AFFAE3F47FEDA018A4E9CFBBE" box="[353,460,729,755]" pageId="1" pageNumber="426" type="holotype">Holotype</typeStatus>
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, male (
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<figureCitation id="61C52A4DFFAE3F47FDA5018A4D38FBBE" box="[542,616,729,755]" captionStart="FIGURES 1" captionStartId="2.[151,264,1692,1717]" captionTargetBox="[172,1416,193,1657]" captionTargetId="figure@2.[159,1428,181,1667]" captionTargetPageId="2" captionText="FIGURES 1A–D. Orphnus brevialatus Frolov & Akhmetova, new species, habitus. A–C, holotype, male (abdomen dissected); B, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.3707543" httpUri="https://zenodo.org/record/3707543/files/figure.png" pageId="1" pageNumber="426">Figs. 1</figureCitation>
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A–C, 2A–C). Body elongate, convex, shiny; length
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<quantity id="3E069B2DFFAE3F47FB03018A4A4EFBBE" box="[1208,1310,729,755]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="1" pageNumber="426" unit="mm" value="11.8">11.8 mm</quantity>
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, maximum width
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<quantity id="3E069B2DFFAE3F47FF6501AF4E67FA5A" box="[222,311,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="426" unit="mm" value="5.8">5.8 mm</quantity>
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. Color uniformly dark brown.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C00724A23FACE" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF7C00724E54FA76" box="[199,260,801,827]" italics="true" pageId="1" pageNumber="426">Head</emphasis>
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: Clypeus wide, with convex anterior margin, rounded laterally, finely crenulate. Genae small, not protruding past eyes. Frontal suture indistinct. Clypeus with long, slender horn, slightly curved posteriad (
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<figureCitation id="61C52A4DFFAE3F47FB6F00164A05FA12" box="[1236,1365,837,863]" captionStart="FIGURES 1" captionStartId="2.[151,264,1692,1717]" captionTargetBox="[172,1416,193,1657]" captionTargetId="figure@2.[159,1428,181,1667]" captionTargetPageId="2" captionText="FIGURES 1A–D. Orphnus brevialatus Frolov & Akhmetova, new species, habitus. A–C, holotype, male (abdomen dissected); B, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.3707543" httpUri="https://zenodo.org/record/3707543/files/figure.png" pageId="1" pageNumber="426">Figs. 1B, C</figureCitation>
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). Dorsal surface of head almost impunctate. Labrum deeply sinuate in the middle, distinctly protruding past clypeus.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C00DD4BAEFD5E" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF7C00DD4E66FAEA" box="[199,310,910,935]" italics="true" pageId="1" pageNumber="426">Pronotum</emphasis>
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: with rounded sides, about 1.2 times wider than long, with deeply concave disc and large lateral ridges almost triangular in lateral view (
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<figureCitation id="61C52A4DFFAE3F47FDEA00E24DF1FA86" box="[593,673,945,971]" captionStart="FIGURES 1" captionStartId="2.[151,264,1692,1717]" captionTargetBox="[172,1416,193,1657]" captionTargetId="figure@2.[159,1428,181,1667]" captionTargetPageId="2" captionText="FIGURES 1A–D. Orphnus brevialatus Frolov & Akhmetova, new species, habitus. A–C, holotype, male (abdomen dissected); B, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.3707543" httpUri="https://zenodo.org/record/3707543/files/figure.png" pageId="1" pageNumber="426">Fig 1C</figureCitation>
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). Anterior angles acute; posterior angles rounded, indistinct in dorsal view. Pronotum bordered on anterior margin and base. Base of pronotum rugose. Sides with coarse, rounded punctures separated by about 0.5–1.0 puncture diameters. Lateral margins with long, sparse, brown setae.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C074E4C2AFD7A" blockId="1.[151,1437,656,1979]" box="[199,890,1053,1079]" pageId="1" pageNumber="426">
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<taxonomicName id="3EFE4D4BFFAE3F47FF7C074E4E65FD7A" box="[199,309,1053,1079]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
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<emphasis id="CB8AEADAFFAE3F47FF7C074E4E65FD7A" box="[199,309,1053,1079]" italics="true" pageId="1" pageNumber="426">Scutellum</emphasis>
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</taxonomicName>
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: wide, semicircular, about 1/16 the length of elytra.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C07124ADEFD32" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF7C07124E5DFD16" box="[199,269,1089,1115]" italics="true" pageId="1" pageNumber="426">Elytra</emphasis>
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: convex, about as wide as long, widest in the middle, with almost indistinct humeral umbones. Striae indistinct. Elytra with coarse, ocellate punctation; punctures denser near base and sparser on disc and in apical part.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C07D94DF4FDEE" blockId="1.[151,1437,656,1979]" box="[199,676,1161,1187]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF7C07D94E5BFDEE" box="[199,267,1162,1187]" italics="true" pageId="1" pageNumber="426">Wings</emphasis>
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: reduced, about 1/3 length of elytra.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C07FD4DABFC1A" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF7C07FD4FACFD8A" box="[199,252,1198,1223]" italics="true" pageId="1" pageNumber="426">Legs</emphasis>
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: Protibiae with 3 outer teeth. Lateral margin basad of outer teeth not crenulate. Apical spur of protibia absent. Protarsi of the
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<typeStatus id="2645886AFFAE3F47FEC307824E8AFDA6" box="[376,474,1233,1259]" pageId="1" pageNumber="426" type="holotype">holotype</typeStatus>
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broken. Middle legs and hind legs similar in shape; metafemora and metatibiae about 1/8 longer than the mesofemora and mesotibiae. Meso- and metatibiae somewhat triangular with 2 apical spurs, inner margin almost straight, with 1 transverse keel. Upper spur of tibiae as long as two basal tarsomeres. Claws 1/3 length of apical tarsomere. Femora almost impunctate.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C06324DA7FCD2" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
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<emphasis id="CB8AEADAFFAE3F47FF7C06324E61FC36" box="[199,305,1377,1403]" italics="true" pageId="1" pageNumber="426">Abdomen</emphasis>
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: Abdominal sternites irregularly punctate, pubescent, with sparse, long setae. Visible sternite 6 medially about 2 times longer than sternites 2–5 combined.
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</paragraph>
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<paragraph id="F94136C8FFAE3F47FF7C06FA4CF7FC8E" blockId="1.[151,1437,656,1979]" box="[199,935,1449,1475]" pageId="1" pageNumber="426">
|
||
<taxonomicName id="3EFE4D4BFFAE3F47FF7C06FA4EC7FC8E" authority=": Surface" authorityName="Surface" box="[199,407,1449,1475]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="426" phylum="Chordata" rank="genus">
|
||
<emphasis id="CB8AEADAFFAE3F47FF7C06FA4E62FC8E" box="[199,306,1449,1475]" italics="true" pageId="1" pageNumber="426">Pygidium</emphasis>
|
||
: Surface
|
||
</taxonomicName>
|
||
irregularly punctate with transverse punctures.
|
||
</paragraph>
|
||
<paragraph id="F94136C8FFAE3F47FF7C069E4D32FF62" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
|
||
<emphasis id="CB8AEADAFFAE3F47FF7C069E4E64FCAA" box="[199,308,1485,1511]" italics="true" pageId="1" pageNumber="426">Aedeagus</emphasis>
|
||
: with relatively long (0.6 length of phallobase), curved downwards parameres tapering apically (
|
||
<figureCitation id="61C52A4DFFAE3F47FAD4069E4FEBFF46" captionStart="FIGURES 2" captionStartId="3.[151,264,1744,1769]" captionTargetBox="[187,1401,194,1706]" captionTargetId="figure@3.[177,1410,181,1718]" captionTargetPageId="3" captionText="FIGURES 2A–D. Orphnus brevialatus Frolov & Akhmetova, new species. A–C, holotype, male. A, aedeagus in lateral view; B, parameres in dorsal view; C, endophallus; D, locality map (red circle—O. brevialatus new species, black circle—O. longicornis Frolov et Akhmetova, white circle—O. valeriae Frolov et Akhmetova; terrestrial ecoregions: 1—tropical and subtropical moist broadleaf forests, 2—tropical and subtropical grasslands, savannas, and shrublands, 3—flooded grasslands and savannas, 4—montane grasslands and shrublands, 5—mangroves, 6—lakes)." figureDoi="http://doi.org/10.5281/zenodo.3707545" httpUri="https://zenodo.org/record/3707545/files/figure.png" pageId="1" pageNumber="426">Fig. 2B</figureCitation>
|
||
). Parameres spear-shaped in dorsal view, with angulate lateral teeth. Endophallus with 2 groups of microspinules and a cluster of larger spinules (
|
||
<figureCitation id="61C52A4DFFAE3F47FE4605464D06FF62" box="[509,598,1557,1583]" captionStart="FIGURES 2" captionStartId="3.[151,264,1744,1769]" captionTargetBox="[187,1401,194,1706]" captionTargetId="figure@3.[177,1410,181,1718]" captionTargetPageId="3" captionText="FIGURES 2A–D. Orphnus brevialatus Frolov & Akhmetova, new species. A–C, holotype, male. A, aedeagus in lateral view; B, parameres in dorsal view; C, endophallus; D, locality map (red circle—O. brevialatus new species, black circle—O. longicornis Frolov et Akhmetova, white circle—O. valeriae Frolov et Akhmetova; terrestrial ecoregions: 1—tropical and subtropical moist broadleaf forests, 2—tropical and subtropical grasslands, savannas, and shrublands, 3—flooded grasslands and savannas, 4—montane grasslands and shrublands, 5—mangroves, 6—lakes)." figureDoi="http://doi.org/10.5281/zenodo.3707545" httpUri="https://zenodo.org/record/3707545/files/figure.png" pageId="1" pageNumber="426">Fig. 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F94136C8FFAE3F47FF7C056A4B9AFF3A" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
|
||
<emphasis id="CB8AEADAFFAE3F47FF7C056A4E4EFF1E" bold="true" box="[199,286,1593,1619]" pageId="1" pageNumber="426">Female</emphasis>
|
||
(
|
||
<figureCitation id="61C52A4DFFAE3F47FE95056A4ED5FF1E" box="[302,389,1593,1619]" captionStart="FIGURES 1" captionStartId="2.[151,264,1692,1717]" captionTargetBox="[172,1416,193,1657]" captionTargetId="figure@2.[159,1428,181,1667]" captionTargetPageId="2" captionText="FIGURES 1A–D. Orphnus brevialatus Frolov & Akhmetova, new species, habitus. A–C, holotype, male (abdomen dissected); B, paratype, female." figureDoi="http://doi.org/10.5281/zenodo.3707543" httpUri="https://zenodo.org/record/3707543/files/figure.png" pageId="1" pageNumber="426">Fig. 1D</figureCitation>
|
||
) differs from the male in having a relatively smaller pronotum without armature, frontoclypeus without process, presence of prothoracic spur. Body length of the female
|
||
<typeStatus id="2645886AFFAE3F47FC43050E4B08FF3A" box="[1016,1112,1629,1655]" pageId="1" pageNumber="426" type="paratype">paratype</typeStatus>
|
||
<quantity id="3E069B2DFFAE3F47FBE4050F4B96FF3A" box="[1119,1222,1628,1655]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="426" unit="mm" value="10.5">10.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1E46543FFAE3F47FF7C05D34DA9FE4A" pageId="1" pageNumber="426" type="diagnosis">
|
||
<paragraph id="F94136C8FFAE3F47FF7C05D34DA9FE4A" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
|
||
<emphasis id="CB8AEADAFFAE3F47FF7C05D34E12FFD6" bold="true" box="[199,322,1664,1691]" pageId="1" pageNumber="426">Diagnosis.</emphasis>
|
||
From the other flightless
|
||
<taxonomicName id="3EFE4D4BFFAE3F47FDCF05D24D86FFD6" authorityName="Macleay" authorityYear="1819" box="[628,726,1665,1691]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CB8AEADAFFAE3F47FDCF05D24D86FFD6" box="[628,726,1665,1691]" italics="true" pageId="1" pageNumber="426">Orphnus</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName id="3EFE4D4BFFAE3F47FCF905D24CB4FFD6" authority="Frolov & Akhmetova, 2020" authorityName="Frolov & Akhmetova" authorityYear="2020" box="[834,996,1665,1691]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="species" species="brevialatus" status="sp. nov.">
|
||
<emphasis id="CB8AEADAFFAE3F47FCF905D24CB4FFD6" box="[834,996,1665,1691]" italics="true" pageId="1" pageNumber="426">O. brevialatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0B957A1FFAE3F47FC5405D24B29FFD6" box="[1007,1145,1665,1691]" pageId="1" pageNumber="426" rank="species">new species</taxonomicNameLabel>
|
||
differs in having a welldeveloped pronotal armature with two long, acute lateral processes. In the shape of the head, pronotum, and male genitalia it is similar to
|
||
<taxonomicName id="3EFE4D4BFFAE3F47FE1F059A4C71FFAE" authority="Quedenfeldt, 1884" authorityName="Quedenfeldt" authorityYear="1884" box="[420,801,1737,1763]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="species" species="compactilis">
|
||
<emphasis id="CB8AEADAFFAE3F47FE1F059A4D18FFAE" box="[420,584,1737,1763]" italics="true" pageId="1" pageNumber="426">O. compactilis</emphasis>
|
||
Quedenfeldt, 1884
|
||
</taxonomicName>
|
||
but differs from it, in addition to the reduced wings and shorter body, in the shape of the endophallic armature.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1E46543FFAE3F47FF7C04434BEAFE3E" pageId="1" pageNumber="426" type="distribution">
|
||
<paragraph id="F94136C8FFAE3F47FF7C04434BEAFE3E" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
|
||
<emphasis id="CB8AEADAFFAE3F47FF7C04434E31FE66" bold="true" box="[199,353,1808,1835]" pageId="1" pageNumber="426">Distribution.</emphasis>
|
||
The new species is known from a single locality in Kipengere Range (Livingstone Mountains), the eastern escarpment of the East African Rift in East Africa. The label does not contain exact locality but suggests that the specimens were collected on the road between towns of Chunya and
|
||
<collectingRegion id="3B3AF82AFFAE3F47FC4D040A4B14FE3E" box="[1014,1092,1881,1907]" country="Tanzania" name="Mbeya" pageId="1" pageNumber="426">Mbeya</collectingRegion>
|
||
(
|
||
<figureCitation id="61C52A4DFFAE3F47FBE8040A4BFDFE3E" box="[1107,1197,1881,1907]" captionStart="FIGURES 2" captionStartId="3.[151,264,1744,1769]" captionTargetBox="[187,1401,194,1706]" captionTargetId="figure@3.[177,1410,181,1718]" captionTargetPageId="3" captionText="FIGURES 2A–D. Orphnus brevialatus Frolov & Akhmetova, new species. A–C, holotype, male. A, aedeagus in lateral view; B, parameres in dorsal view; C, endophallus; D, locality map (red circle—O. brevialatus new species, black circle—O. longicornis Frolov et Akhmetova, white circle—O. valeriae Frolov et Akhmetova; terrestrial ecoregions: 1—tropical and subtropical moist broadleaf forests, 2—tropical and subtropical grasslands, savannas, and shrublands, 3—flooded grasslands and savannas, 4—montane grasslands and shrublands, 5—mangroves, 6—lakes)." figureDoi="http://doi.org/10.5281/zenodo.3707545" httpUri="https://zenodo.org/record/3707545/files/figure.png" pageId="1" pageNumber="426">Fig. 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1E46543FFAE3F47FF7C042E4ED7FEF6" pageId="1" pageNumber="426" type="etymology">
|
||
<paragraph id="F94136C8FFAE3F47FF7C042E4ED7FEF6" blockId="1.[151,1437,656,1979]" pageId="1" pageNumber="426">
|
||
<emphasis id="CB8AEADAFFAE3F47FF7C042E4E1EFEDA" bold="true" box="[199,334,1917,1943]" pageId="1" pageNumber="426">Etymology.</emphasis>
|
||
From Latin adjectives in the nominative singular
|
||
<emphasis id="CB8AEADAFFAE3F47FCC6042E4C90FEDA" box="[893,960,1917,1943]" italics="true" pageId="1" pageNumber="426">brevis</emphasis>
|
||
(= short), and
|
||
<emphasis id="CB8AEADAFFAE3F47FBDE042E4BF9FEDA" box="[1125,1193,1917,1943]" italics="true" pageId="1" pageNumber="426">alatus</emphasis>
|
||
(= winged), referring to the reduced wings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |