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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.106698" ID-Pensoft-Pub="1313-2970-1184-301" ID-Pensoft-UUID="283A8869D1C3500A8A3C075E14584C90" ID-ZooBank="4EB4BEE3865E4CF29F7B6477D2DEF4AF" ModsDocID="1313-2970-1184-301" checkinTime="1700745464390" checkinUser="pensoft" docAuthor="Romero-Ortiz, Catalina, Sarmiento, Carlos E. &amp; Harvey, Mark S." docDate="2023" docId="710E2ECDB1E1545683F67D4D819225FC" docLanguage="en" docName="ZooKeys 1184: 301-326" docOrigin="ZooKeys 1184" docPubDate="2023-11-22" docSource="http://dx.doi.org/10.3897/zookeys.1184.106698" docTitle="Cystowithius florezi Romero-Ortiz &amp; Sarmiento &amp; Harvey 2023, sp. nov." docType="treatment" docUuid="F2D766B6-215E-40D9-908B-76416F859F85" docUuidSource="ZooBank" docVersion="2" id="283A8869D1C3500A8A3C075E14584C90" lastPageNumber="301" masterDocId="283A8869D1C3500A8A3C075E14584C90" masterDocTitle="A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia" masterLastPageNumber="326" masterPageNumber="301" pageNumber="301" updateTime="1700745769359" updateUser="ExternalLinkService">
<mods:mods id="A1880B6C36D2FBF4F8513F1EE0820C5E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7C599B5664EC19DA45D278DC98DDB418">
<mods:title id="80B8550F5ED575BC69F0073C3F77B89D">A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia</mods:title>
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<mods:namePart id="73560F2F04887E79A454ED3890365490">Romero-Ortiz, Catalina</mods:namePart>
<mods:nameIdentifier id="0B2CA3C3166E513A4AFE24F570976D9B" type="ORCID">https://orcid.org/0000-0001-8939-7814</mods:nameIdentifier>
<mods:affiliation id="D8C0F0CC409E0A6692624D3A649E5124">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
<mods:nameIdentifier id="7C0A8535BD362CDA2EA461876F0DEA61" type="email">icromeroo@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="CD834CC539DDA056C5E083D5E7F250B4" type="personal">
<mods:role id="3FF9E17E6F8B513D2F6B4E028F4D1621">
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<mods:namePart id="BEF28A35D2ED620D2635573A2293385D">Sarmiento, Carlos E.</mods:namePart>
<mods:nameIdentifier id="5DE5FD92131080E705C739720E24FB71" type="ORCID">https://orcid.org/0000-0003-4012-8108</mods:nameIdentifier>
<mods:affiliation id="2FCB82D614DFD14D75A1625DB9F4697D">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
</mods:name>
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<mods:namePart id="491B828550507C82674D11A5759EF1A0">Harvey, Mark S.</mods:namePart>
<mods:nameIdentifier id="CBD5575E97C83AA5983C8CBB46A47B57" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
<mods:affiliation id="03CD585A6DC9C5A46A53A356487A5363">Collections and Research, Western Australian Museum, Welshpool, Western Australia 6106, Australia &amp; School of Biological Sciences, The University of Western Australia, Crawley, Western Australia 6009, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="ED24E0737CAA5E9F725E15CC22BB295F">text</mods:typeOfResource>
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<mods:title id="784BC2F5AA0F0A9813E6034A5035156A">ZooKeys</mods:title>
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<mods:part id="9EBCF637B3159D9B9B7272E971BE0B48">
<mods:date id="BDD9F224DDCA87D4C7C5E7733E1F22B5">2023</mods:date>
<mods:detail id="EB66298ABCB7ADADCE6213035C6BE137" type="pubDate">
<mods:number id="E02E3DCD6CF787A78362F35903002723">2023-11-22</mods:number>
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<mods:detail id="5A7385F8AEFA06F93D5FAD676B483D80" type="volume">
<mods:number id="2CDBDD33D7E3578149C50FE5E26BEAEB">1184</mods:number>
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<mods:classification id="3052F144C28A7D8B7F60EB8FBB1D5BA7">journal article</mods:classification>
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<treatment id="710E2ECDB1E1545683F67D4D819225FC" LSID="urn:lsid:zoobank.org:act:F2D766B6-215E-40D9-908B-76416F859F85" httpUri="http://treatment.plazi.org/id/710E2ECDB1E1545683F67D4D819225FC" lastPageNumber="301" pageId="0" pageNumber="301">
<subSubSection id="011A5EA84E028AF4E614A557CC5C812C" pageId="0" pageNumber="301" type="nomenclature">
<paragraph id="06E82AC803AAA57745408A1DDCCA3610" pageId="0" pageNumber="301">
<taxonomicName id="20B7961811AF86DE489C18219513D09E" LSID="https://zoobank.org/F2D766B6-215E-40D9-908B-76416F859F85" authority="Romero-Ortiz &amp; Sarmiento &amp; Harvey, 2023" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius florezi" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="florezi" status="sp. nov.">Cystowithius florezi</taxonomicName>
<taxonomicNameLabel id="84A8E92D2F343439047EA6CDA5FB025D" pageId="0" pageNumber="301">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6DB0B73934E0482356A9D80F03C5C54A" pageId="0" pageNumber="301" type="description">
<paragraph id="6E5DD5B3E4DC68C01BF22F26705DB462" pageId="0" pageNumber="301">
<figureCitation id="4EE50B7C34EEC2AED88B05942439F780" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of the new species (males) A Paciwithius chimbilacus sp. nov. B Cystowithius florezi sp. nov. C Parawithius bromelicola sp. nov. D Oligowithius achagua sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.106698.figure1" httpUri="https://binary.pensoft.net/fig/940870" pageId="0" pageNumber="301">Figs 1B</figureCitation>
<figureCitation id="8D5AC0B6A632FB0BC4A5C57F0ED16987" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the new species and the new record of Withiidae from Colombia." figureDoi="10.3897/zookeys.1184.106698.figure2" httpUri="https://binary.pensoft.net/fig/940871" pageId="0" pageNumber="301">, 2</figureCitation>
<figureCitation id="4A9AA1BFDD6D798FF163EAF70C066931" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">, 5</figureCitation>
<figureCitation id="755D8934CF28542B34700816B9ECD481" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glandular setae patches of the new species A Paciwithius chimbilacus sp. nov. (holotype, ICN-Aps- 076) B Parawithius bromelicola sp. nov. (holotype, ICN-Aps- 082) C Paciwithius valduparensis sp. nov. (holotype, ICN-Aps- 597) D Cystowithius florezi sp. nov. (holotype, ICN-Aps- 07) E, F Oligowithius achagua sp. nov. E male paratype (ICN-Aps- 388) F male holotype (ICN-Aps- 388). Scale bars: 0.1 mm (A-C, E, F); 0.05 mm (D)." figureDoi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301">, 9D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1838429EA30B59E23119F41FC50BB6F8" pageId="0" pageNumber="301" type="materials_examined">
<paragraph id="6800EAFF983BBCD54B84B9940A0C4F68" pageId="0" pageNumber="301">Material examined.</paragraph>
<paragraph id="E5BD940C7666919D30C1690B250337BF" pageId="0" pageNumber="301">
<materialsCitation id="D39A30ECC632B5851EDA76B78F6070CE" collectingDate="2007-02-12" collectorName="C. Cortes" country="Colombia" elevation="1700" latitude="4.5561113" location="Reserva Natural Ibanasca" longLatPrecision="1" longitude="-75.32144" municipality="Juntas" specimenCount="♂" stateProvince="Tolima" typeStatus="Holotype">
<emphasis id="C2415A7ECB73CA0AA1ED4ED97B6D6B6C" bold="true" pageId="0" pageNumber="301">
<emphasis id="93DFF5498203488BF6F9F2E39E30144B" italics="true" pageId="0" pageNumber="301">
<typeStatus id="80ACD90E94762A6DDFCF48383DECD42B">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="2BFD4827177525CEFCEFE38EF0EE35F5" name="Colombia">Colombia</collectingCountry>
•
<specimenCount id="D8CF26EAE14516658D96E324183629AC">♂</specimenCount>
;
<collectingRegion id="2FD175597CF004D45B67E588564F9430" country="Colombia" name="Tolima">Tolima</collectingRegion>
,
<collectingMunicipality id="F2B9DF1A81D0773BE438D5F7480E0631">Juntas</collectingMunicipality>
,
<location id="5EE918090A02F66C0159271F06C57FA7" LSID="urn:lsid:plazi:treatment:710E2ECDB1E1545683F67D4D819225FC:5EE918090A02F66C0159271F06C57FA7" country="Colombia" latitude="4.5561113" longLatPrecision="1" longitude="-75.32144" municipality="Juntas" name="Reserva Natural Ibanasca" stateProvince="Tolima">Reserva Natural Ibanasca</location>
; [
<geoCoordinate id="23DBAC34D394819F9AFA7B79E5B79D22" degrees="4" direction="north" minutes="33" orientation="latitude" precision="1" seconds="22.0" value="4.5561113">4°33'22.0&quot;N</geoCoordinate>
,
<geoCoordinate id="D8EC54EA030E939635B55986706D2291" degrees="75" direction="west" minutes="19" orientation="longitude" precision="1" seconds="17.2" value="-75.32144">75°19'17.2&quot;W</geoCoordinate>
];
<quantity id="E7119EED3087E2EC36016E69B3538995" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="03ECF802EA85325D3458F10C88EE3E0D" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</elevation>
</quantity>
;
<collectingDate id="17E9D9CD59A360B78906C7C22190B0CF" value="2007-02-12">12 Feb. 2007</collectingDate>
;
<collectorName id="125D7904EA7D63649097045F4F041C77">C. Cortes</collectorName>
leg.;
<taxonomicName id="455487C92E1E5137D5409BE462977ECA" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="0" pageNumber="301" phylum="Coniferophyta" rank="genus">
<emphasis id="B314ECF8627CA15AA093C396268AE51D" italics="true" pageId="0" pageNumber="301">Pinus</emphasis>
</taxonomicName>
plantation; on low vegetation, manual capture; ICN-APs-077.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0561688E6AA8752B92902769F5B7AF3C" pageId="0" pageNumber="301" type="diagnosis">
<paragraph id="12C81830574F05A2DB42DC584FC367ED" pageId="0" pageNumber="301">Diagnosis.</paragraph>
<paragraph id="0B65C2B64E2D5FD5E42CE0083A79ABDA" pageId="0" pageNumber="301">
<taxonomicName id="23501006D5A27ACF70E7041F4518D127" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius florezi" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="6AD0F9C92BE3056955084F87DEA5DB23" italics="true" pageId="0" pageNumber="301">Cystowithius florezi</emphasis>
</taxonomicName>
can be separated from the other
<taxonomicName id="E2F3F89EC5918F1BD286B8B8BD13C32E" authorityName="Harvey" authorityYear="2004" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="6C460A632A2F9D6D96B713CB6E92F044" italics="true" pageId="0" pageNumber="301">Cystowithius</emphasis>
</taxonomicName>
species as follows.
<taxonomicName id="D6B06B073E00529BE33C074A4031B9D2" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius florezi" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="4E393C141DF49608A2285C47F1408302" italics="true" pageId="0" pageNumber="301">Cystowithius florezi</emphasis>
</taxonomicName>
sp. nov. is very similar to
<taxonomicName id="61DB6FFDACBD2E84CB153A1A91E6D127" lsidName="C. colombicus" pageId="0" pageNumber="301" rank="species" species="colombicus">
<emphasis id="AE3F891029B221DC4E8EDD7EF4DA3050" italics="true" pageId="0" pageNumber="301">C. colombicus</emphasis>
</taxonomicName>
Harvey, 2004 and
<taxonomicName id="E833C8DDABAF572699363B5C3757CF89" lsidName="C. ecuadoricus" pageId="0" pageNumber="301" rank="species" species="ecuadoricus">
<emphasis id="718D513424488531833C4BB95E38EA8E" italics="true" pageId="0" pageNumber="301">C. ecuadoricus</emphasis>
</taxonomicName>
Harvey, 2004, so we provide a detailed comparison for each one in Table
<tableCitation id="7BE3E2C67EE444DE3DCF2A6209EDDE2F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparative characters between three species of Cystowithius. Bold font denotes ratios." httpUri="http://table.plazi.org/id/817D765CB7F29FAE67DE1B4A1D46434B" pageId="0" pageNumber="301" tableUuid="817D765CB7F29FAE67DE1B4A1D46434B">1</tableCitation>
. In general,
<taxonomicName id="588A516277308B7A8AABF08E028BCF09" lsidName="C. colombicus" pageId="0" pageNumber="301" rank="species" species="colombicus">
<emphasis id="0366BD8CF272D57BD7CAEBD5A0FFB353" italics="true" pageId="0" pageNumber="301">C. colombicus</emphasis>
</taxonomicName>
has bigger pedipalp segments and the patches of glandular setae located in sternites VII-IX; in
<taxonomicName id="67567BE6DEDFC1D23EE66E9527FFCF55" lsidName="C. florezi" pageId="0" pageNumber="301" rank="species" species="florezi">
<emphasis id="A20770326E6868FE5DF5A58A14C1221B" italics="true" pageId="0" pageNumber="301">C. florezi</emphasis>
</taxonomicName>
the patches are just in sternite VIII. Also, the sternal pockets are in sternites VI and VII in
<taxonomicName id="FBF1F07C8593C766C5CA3681FE49CEEE" lsidName="C. colombicus" pageId="0" pageNumber="301" rank="species" species="colombicus">
<emphasis id="162BA436972FE837669A3856977507E4" italics="true" pageId="0" pageNumber="301">C. colombicus</emphasis>
</taxonomicName>
and in V-VIII in
<taxonomicName id="86F3F1F64E634F0E2ECAF7A34AC44F49" lsidName="C. florezi" pageId="0" pageNumber="301" rank="species" species="florezi">
<emphasis id="1D138A39E40AAE4430173B870782133E" italics="true" pageId="0" pageNumber="301">C. florezi</emphasis>
</taxonomicName>
. It differs from
<taxonomicName id="CED07D6258492A6D60317545E5774D68" lsidName="C. chamberlini" pageId="0" pageNumber="301" rank="species" species="chamberlini">
<emphasis id="5FC7110EE25C9568A50C3476E8678E7A" italics="true" pageId="0" pageNumber="301">C. chamberlini</emphasis>
</taxonomicName>
by the chelal hand being smooth rather than strongly granulated in
<taxonomicName id="3BB4A540F1D0F8BDFF241B88F5E1D346" lsidName="C. florezi" pageId="0" pageNumber="301" rank="species" species="florezi">
<emphasis id="ECBA3BB46D16FE8B8E1812235F5515CB" italics="true" pageId="0" pageNumber="301">C. florezi</emphasis>
</taxonomicName>
, and the position of the tactile setae on tarsus IV located close to midway of the tarsus, rather than distal in
<taxonomicName id="BC6E11BD4DC9029880A6093800CF8641" lsidName="C. florezi" pageId="0" pageNumber="301" rank="species" species="florezi">
<emphasis id="2A70FC53852765D0CCA17DCEE0BCAA78" italics="true" pageId="0" pageNumber="301">C. florezi</emphasis>
</taxonomicName>
; from
<taxonomicName id="C44925EE78F9C2BEE3980EAD472EB282" lsidName="C. ankeri" pageId="0" pageNumber="301" rank="species" species="ankeri">
<emphasis id="C31814314AF204141E9280910EDEA2FF" italics="true" pageId="0" pageNumber="301">C. ankeri</emphasis>
</taxonomicName>
by the length of the movable chelal finger which is longer in
<taxonomicName id="E1F44A0910D29BE793B921A0F5DB536D" lsidName="C. ankeri" pageId="0" pageNumber="301" rank="species" species="ankeri">
<emphasis id="093778B298BEFD80EB1CFDFA4963F844" italics="true" pageId="0" pageNumber="301">C. ankeri</emphasis>
</taxonomicName>
than in
<taxonomicName id="6F63DAA03B1E3B86F5ADC606A6759901" lsidName="C. florezi" pageId="0" pageNumber="301" rank="species" species="florezi">
<emphasis id="5544BD4CBE289D0F987D83A66DE86FB9" italics="true" pageId="0" pageNumber="301">C. florezi</emphasis>
</taxonomicName>
(0.70 mm vs 0.52 mm); and from
<taxonomicName id="DAAE3CDA5EB329941BFCEBEBB1336A4D" lsidName="C. smithersi" pageId="0" pageNumber="301" rank="species" species="smithersi">
<emphasis id="DC4A0286DC951D18EABCEDD849942288" italics="true" pageId="0" pageNumber="301">C. smithersi</emphasis>
</taxonomicName>
by the length of the chela with pedicel which is longer in
<taxonomicName id="07F797AC1D29055F2513C24ECFB064D1" lsidName="C. smithersi" pageId="0" pageNumber="301" rank="species" species="smithersi">
<emphasis id="7F8EE9EF64A67CDBEB7061173BA7E1BE" italics="true" pageId="0" pageNumber="301">C. smithersi</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C49B2170D85EF4EC249FEF490C1D8EE" lsidName="C. florezi" pageId="0" pageNumber="301" rank="species" species="florezi">
<emphasis id="152E0F1734A7E307DA7E18C5890FF771" italics="true" pageId="0" pageNumber="301">C. florezi</emphasis>
</taxonomicName>
(1.35 mm vs 1.02 mm).
</paragraph>
<caption id="817D765CB7F29FAE67DE1B4A1D46434B" ID-Table-UUID="817D765CB7F29FAE67DE1B4A1D46434B" httpUri="http://table.plazi.org/id/817D765CB7F29FAE67DE1B4A1D46434B" pageId="0" pageNumber="301" start="Table 1" startId="T1">
<paragraph id="A9C2858ED8010D48A7E08ABAE0F7C4F2" pageId="0" pageNumber="301">
<emphasis id="0CA062E42A3541EF3000811F6D641278" bold="true" pageId="0" pageNumber="301">Table 1.</emphasis>
Comparative characters between three species of
<taxonomicName id="3B02DBD967AE4CCC76605FE0925CBDEF" authorityName="Harvey" authorityYear="2004" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="DF661E5E83969F1C106E8B97F640B686" italics="true" pageId="0" pageNumber="301">Cystowithius</emphasis>
</taxonomicName>
.
<emphasis id="27647DE4168612C164D645EE0C2C6443" bold="true" pageId="0" pageNumber="301">Bold</emphasis>
font denotes ratios.
</paragraph>
</caption>
<paragraph id="A5D9B64666A3224248EF9BF11793E8EC" pageId="0" pageNumber="301">
<table id="6B65236E8BAD8FD2A760C1C8F29D40A1" pageId="0" pageNumber="301">
<tr id="3E059B5F37C1C724B0F859874FD8B84E" pageId="0" pageNumber="301">
<th id="DA552781ECE7F7422CB6C90FC483D32D" colspan="1" pageId="0" pageNumber="301" rowspan="1">Species/ Characters</th>
<th id="AEC37DAF1D6EF833EF9A7D4A750CCAF4" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<taxonomicName id="269B9A1ADA4CCE3F406420ED05DB6C61" lsidName="C. colombicus" pageId="0" pageNumber="301" rank="species" species="colombicus">
<emphasis id="6E48C59E47E13B7CED4AAD2D28356075" italics="true" pageId="0" pageNumber="301">C. colombicus</emphasis>
</taxonomicName>
Harvey, 2004
</th>
<th id="7943221BA43D0C2E017B392624E2E999" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<taxonomicName id="55A9DAB6EBEB1A7B723C452F7D0BCEAB" lsidName="C. ecuadoricus" pageId="0" pageNumber="301" rank="species" species="ecuadoricus">
<emphasis id="C6704D9B158B0D03FA739FACE0DBD415" italics="true" pageId="0" pageNumber="301">C. ecuadoricus</emphasis>
</taxonomicName>
Harvey, 2004
</th>
<th id="9D2447A330D545315D1C1EF8FABBC682" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<taxonomicName id="9B315581B8CF2F28F4F37D6A5704E702" lsidName="C. florezi" pageId="0" pageNumber="301" rank="species" species="florezi">
<emphasis id="731D6EF113A3C1233762EFBA2E22AC58" italics="true" pageId="0" pageNumber="301">C. florezi</emphasis>
</taxonomicName>
sp. nov.
</th>
</tr>
<tr id="1F9AE3755FE45417DA9D03CD2B784991" pageId="0" pageNumber="301">
<td id="9EB3E4BFCAACF42FFAB9B4142B8191F4" colspan="1" pageId="0" pageNumber="301" rowspan="1">Body length</td>
<td id="D68A92050E26C6BD8289CACA8B499F41" colspan="1" pageId="0" pageNumber="301" rowspan="1">2.00-2.11</td>
<td id="432D295A9947FE2727B01DD6FA69D655" colspan="1" pageId="0" pageNumber="301" rowspan="1">2.16-2.29</td>
<td id="59257F8844170274FDBD127A15D79D63" colspan="1" pageId="0" pageNumber="301" rowspan="1">1.94</td>
</tr>
<tr id="6D48C07DAAE7B09BCE04CE94C3174878" pageId="0" pageNumber="301">
<td id="6E1ABEDA42B2616FDCA915FCB6DB4018" colspan="1" pageId="0" pageNumber="301" rowspan="2">Carapace length/width</td>
<td id="17A7C7003F5527E82FE991F0D8EAC455" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.74/0.52</td>
<td id="C4F276FE089025D04CEA6196DF6AD209" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.75-0.80/0.62</td>
<td id="845B121FE703ED2AC389DBFBEBCFFB25" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.67/0.51</td>
</tr>
<tr id="9EAB241AC69BA3CFD0EC5D1A2C0C7639" pageId="0" pageNumber="301">
<td id="52910D14681C0614D6DDD0DC82D94498" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="20C33CF71FC07488642D7013021511D4" bold="true" pageId="0" pageNumber="301">1.43</emphasis>
</td>
<td id="A72A79CC4B18EFFC10A1CCD492A35A6F" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="B39D8A21FCA546877D2ADCD47D4A7AE9" bold="true" pageId="0" pageNumber="301">1.20-1.28</emphasis>
</td>
<td id="E93901A9F7D7BB205F71298CA871839D" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="0D4EBB95FC6821049DEBAF5446C31119" bold="true" pageId="0" pageNumber="301">1.31</emphasis>
</td>
</tr>
<tr id="D360C3618C40BB18280B3DBF6314F3CC" pageId="0" pageNumber="301">
<td id="C9D00A08CB5C2ED464395CDD2E132543" colspan="1" pageId="0" pageNumber="301" rowspan="2">Pedipalpal femur length/width</td>
<td id="DB134AE8D55F8188C3B1D434329A645E" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.87-0.97/0.17-0.20</td>
<td id="B197F0BBE8CD66094A6DF1919DB7D518" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.80-0.82/ 0.17-0.18</td>
<td id="2EBFA073140E17F89FB2970A97B72BE1" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.60/0.17</td>
</tr>
<tr id="0F8A4CE8A4C7A15089F3E8C443C63FA7" pageId="0" pageNumber="301">
<td id="CBD75B8F761BADE68F162C605BFC58F6" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="B75D6FC25EE4FFFAD05F0D2787E337E4" bold="true" pageId="0" pageNumber="301">4.78-5.25</emphasis>
</td>
<td id="6DBEE7476CFA30121F6E01497BCB95AB" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="9D56F00B4F19A1BDB835FDCDA779DA1A" bold="true" pageId="0" pageNumber="301">4.47-4.74</emphasis>
</td>
<td id="56A264664AF37DF301C60015BB11D84E" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="20714838618A1329365034FA9136A954" bold="true" pageId="0" pageNumber="301">3.60</emphasis>
</td>
</tr>
<tr id="332BD6C7CFDA3266D8C2AEF41DC826AB" pageId="0" pageNumber="301">
<td id="67E78E2FEE64FFD06CE5ADD0D8C919BB" colspan="1" pageId="0" pageNumber="301" rowspan="2">Pedipalp patella length/width</td>
<td id="9A1C8ABB8E443493FCD573E5BD36F0C7" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.77-0.88/0.19-0.21</td>
<td id="7520D19556B5FF1F953885CA5BEDACE6" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.67-0.69/0.20-0.21</td>
<td id="E8CEDF83524C860207A3FAFFB2F4C80A" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.61/0.18</td>
</tr>
<tr id="CAFCEDB6CF4C90F6F0ED370D71AB0524" pageId="0" pageNumber="301">
<td id="7D97E3DDF2CE02E33787FC0B12BD77E1" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="CB92DBF8E2A19EE3F3FB488EBCEA94AD" bold="true" pageId="0" pageNumber="301">3.88-4.46</emphasis>
</td>
<td id="910B71A7362F0990FC37DC732BF4C4CD" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="2D3FD3E972B7172418479109FB3FCDD8" bold="true" pageId="0" pageNumber="301">3.18-3.47</emphasis>
</td>
<td id="2F2FB593B3A93E3D3D94EA868C0056EB" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="282BC5E0440A76490BE469D06901F938" bold="true" pageId="0" pageNumber="301">3.30</emphasis>
</td>
</tr>
<tr id="11834FF4DA13DFDF2D982F0B7A5B94F3" pageId="0" pageNumber="301">
<td id="3D26FD2E5A7F549D2AF46E6993A316B0" colspan="1" pageId="0" pageNumber="301" rowspan="2">Chela with pedicel length/width</td>
<td id="5AF37B9985386347B6691231E77374A5" colspan="1" pageId="0" pageNumber="301" rowspan="1">1.12-1.26/0.28-0.33</td>
<td id="B6EFF6E8B1574C37762DA604B34E6E7F" colspan="1" pageId="0" pageNumber="301" rowspan="1">1.09-1.14/0.28-0.30</td>
<td id="76B6E7C0CAA8496ED59650C9074185AA" colspan="1" pageId="0" pageNumber="301" rowspan="1">1.02/0.28</td>
</tr>
<tr id="46DABA421726CCC0770633C8E4C4B73D" pageId="0" pageNumber="301">
<td id="9B003219E71765F9756BFEDE10B5BBF1" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="5DE33F7E36CFB1F3C281EAE6DC83F5CC" bold="true" pageId="0" pageNumber="301">3.80-4.04</emphasis>
</td>
<td id="B0A72DE775340AF295799497DAD2BE73" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="2317B2711685B1CC0BA16395EC2E7C3B" bold="true" pageId="0" pageNumber="301">3.65-4.01</emphasis>
</td>
<td id="A0A565255711CEACD335427350A2BF79" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="286F9D9D95CCF76CF67B6DCF733CACA4" bold="true" pageId="0" pageNumber="301">3.64</emphasis>
</td>
</tr>
<tr id="487D80CDFB02E0E6A3C209DAF4DF234B" pageId="0" pageNumber="301">
<td id="3F094AD7B3827FB11CF7FEED13E364F4" colspan="1" pageId="0" pageNumber="301" rowspan="2">Movable finger ratio (length/hand width)</td>
<td id="9688852FE50E241ECFDE56FDEE677A5A" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.49-0.61</td>
<td id="0BC0E68BA4EB0C82F629F712455338B1" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.50-0.56</td>
<td id="8F240F54A5D6801102F01910567393DA" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.52</td>
</tr>
<tr id="3D35329006BFC3D391535B711C29DDAD" pageId="0" pageNumber="301">
<td id="3BC40086A80A227F7E3C95D1C191ADD0" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="842BD58FB13ECF52558EE258622E70AF" bold="true" pageId="0" pageNumber="301">0.90-1.24</emphasis>
</td>
<td id="C4ED44D77D5EA311413E75693C630798" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="AF5893FCF84B91D93713EE24565B7711" bold="true" pageId="0" pageNumber="301">0.86-1.11</emphasis>
</td>
<td id="380DA1CF94023F7E3822C3A035225BDC" colspan="1" isEmpty="true" pageId="0" pageNumber="301" rowspan="1">-</td>
</tr>
<tr id="8B8DFFAC60814780132753F5AE15C252" pageId="0" pageNumber="301">
<td id="BA38BA237DA4A4E34167E3620AE206FC" colspan="1" pageId="0" pageNumber="301" rowspan="1">Hand length/width</td>
<td id="694A6AF21259257623B08B43B0BF9A99" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="28D1B38030EAFB4D1BA3B1983BA0DEF1" bold="true" pageId="0" pageNumber="301">0.50-0.60</emphasis>
</td>
<td id="68AF2C5C94484E80BB682344867FD531" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="690C2B72EE8128738C68545D6A0E01A0" bold="true" pageId="0" pageNumber="301">0.50-0.58</emphasis>
</td>
<td id="ECDC7FFC94EB3132BBBB9B211356BBF5" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="E75C6857B29D7EAE51567378B4281E9F" bold="true" pageId="0" pageNumber="301">0.49</emphasis>
</td>
</tr>
<tr id="D7A79349CE3224DF7541FA05F76BEAB2" pageId="0" pageNumber="301">
<td id="07C0F3675C0E4D1DAC44D73C13890171" colspan="1" pageId="0" pageNumber="301" rowspan="2">Femur/patella IV</td>
<td id="6EC088924A8B5A35E3E3C3FE8DAD877B" colspan="1" pageId="0" pageNumber="301" rowspan="2">0.67/0.17</td>
<td id="87F808D5F1CD05FF9B98AAABC0846137" colspan="1" pageId="0" pageNumber="301" rowspan="1">0.60/0.21</td>
<td id="0B387F07951D7492779F28B101AA7AF4" colspan="1" pageId="0" pageNumber="301" rowspan="2">0.57/0.18</td>
</tr>
<tr id="149A8B02B016F0410FC0DDFBD1063AD3" pageId="0" pageNumber="301">
<td id="F468EC549DF9610FDAF488E3D5B18D8C" colspan="1" pageId="0" pageNumber="301" rowspan="1">
<emphasis id="920F65BCC5FB9941AECC66658C363A7D" bold="true" pageId="0" pageNumber="301">2.93</emphasis>
</td>
</tr>
<tr id="073BD4D05B58403967EFAFD672388EB4" pageId="0" pageNumber="301">
<td id="DA718EA0EC887BC4F72C584D2BDDF9D1" colspan="1" pageId="0" pageNumber="301" rowspan="1">Glandular setae</td>
<td id="0AE5AF4270DD77843CDC5D7B9781EAFE" colspan="1" pageId="0" pageNumber="301" rowspan="1">
sternites VII-IX with patches of
<emphasis id="F14F0F8340619122D29C18BB048F174D" italics="true" pageId="0" pageNumber="301">gls</emphasis>
arranged ca. 20: 27: 13 respectively
</td>
<td id="D3F7013A65D23F6E21A0FE74CA8A563D" colspan="1" pageId="0" pageNumber="301" rowspan="1">
sternites VI-IX with patches of
<emphasis id="EE52C7F506EAC1A0EA22232E4D9B6C44" italics="true" pageId="0" pageNumber="301">gls</emphasis>
, arranged 6: 42: 10: 8 (lectotype) respectively
</td>
<td id="96D100F933794356AEACAEEF163B6E8C" colspan="1" pageId="0" pageNumber="301" rowspan="1">
sternite VIII with patches of
<emphasis id="1FC9D2D547ED69094C4020BB01BEE647" italics="true" pageId="0" pageNumber="301">gls</emphasis>
, arranged 40 in a small circle
</td>
</tr>
<tr id="0A8AC9F5344E280D66A2E552B9E07D51" pageId="0" pageNumber="301">
<td id="261BBCFF64CB646C36444D869198C8FC" colspan="1" pageId="0" pageNumber="301" rowspan="1">Sternal pockets on anterior margins of sternites</td>
<td id="3D29F0E9BD5D0E10B39661BD9DD7BB36" colspan="1" pageId="0" pageNumber="301" rowspan="1">VI-VII</td>
<td id="0916FA06162A5E1E11BB698A60E75D75" colspan="1" pageId="0" pageNumber="301" rowspan="1">V-VIII</td>
<td id="3A4C8482EA94647FF8EE3F5DA799BC0F" colspan="1" pageId="0" pageNumber="301" rowspan="1">VI-VIII</td>
</tr>
<tr id="8828440879314CE4F707B1BD478553EC" pageId="0" pageNumber="301">
<td id="E532C7936719A1CA2AF5449645F1E7B6" colspan="1" pageId="0" pageNumber="301" rowspan="1">Sense spots on chelal fingers</td>
<td id="2678D393153A5B56461F9E7C8F2E5CFD" colspan="1" pageId="0" pageNumber="301" rowspan="1">without sense spots on movable finger</td>
<td id="A72931EF3AA6B7049D01CEFECE8524E1" colspan="1" pageId="0" pageNumber="301" rowspan="1">
without sense spots on movable finger but with small sense spot slightly distal to
<emphasis id="B3E89C9D0313AC77266D64592700BF52" italics="true" pageId="0" pageNumber="301">st</emphasis>
that contains three small nubbins
</td>
<td id="D35D84C2E4EB52FCF74CF832859E1AE3" colspan="1" pageId="0" pageNumber="301" rowspan="1">
without sense spots on movable finger but with small spot slightly distal to
<emphasis id="53F9FD979EC0B27CFED1742E05CDEEC2" italics="true" pageId="0" pageNumber="301">st</emphasis>
that contains two small nubbins
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="E10CFF914FEF49FB8564D0766934AF65" pageId="0" pageNumber="301" type="description">
<paragraph id="438B8D876CC10CCF36CA7794F42569E3" pageId="0" pageNumber="301">Description.</paragraph>
<paragraph id="DD2502C26276B7404E48CC31183FA51A" pageId="0" pageNumber="301">
<emphasis id="DE92F6461BDA223BD1C4869F37286315" bold="true" italics="true" pageId="0" pageNumber="301">Color</emphasis>
: with sclerotized portions, generally yellow-brown; carapace and pedipalps darker; carapace metazone with paired pale spots; legs darker at the edges (Fig.
<figureCitation id="A69C07C5CC61EC59B58793F9D520F9F6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of the new species (males) A Paciwithius chimbilacus sp. nov. B Cystowithius florezi sp. nov. C Parawithius bromelicola sp. nov. D Oligowithius achagua sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.106698.figure1" httpUri="https://binary.pensoft.net/fig/940870" pageId="0" pageNumber="301">1B</figureCitation>
).
</paragraph>
<paragraph id="35F43E9E5F70892331C410EFB35316AE" pageId="0" pageNumber="301">
<emphasis id="4B4B716B060EA926ACC64C4C6CD90A5A" bold="true" italics="true" pageId="0" pageNumber="301">Dimensions</emphasis>
(mm): male: holotype: body length 1.94. Pedipalps: trochanter 0.36/0.18, femur 0.60/0.17, patella 0.61/0.18, chela (with pedicel) 1.02/0.28, chela (without pedicel) 0.96, hand (without pedicel) length 0.49, movable finger length 0.52. Chelicera 0.19, movable finger length 0.14. Carapace 0.67/0.51 (width at medial area); eye diameter 0.06. Leg I: femur 0.14/0.14, patella 0.29/0.14, tibia 0.20/0.10, tarsus 0.25/0.06. Leg IV: femur + patella 0.57/0.18, tibia 0.43/0.18, tarsus 0.34/0.07, TS 0.74.
</paragraph>
<paragraph id="F5A6CE88055667D276E1E5FED8C1D5A9" pageId="0" pageNumber="301">
<emphasis id="55064558CD5838E891C65E630D45D34E" bold="true" italics="true" pageId="0" pageNumber="301">Carapace</emphasis>
(Fig.
<figureCitation id="96BC22616581893D91EBCB76B498447F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5A</figureCitation>
): 1.31
<normalizedToken id="40E25D77A23DDCAF9092F8217E5C5F60" originalValue="×">x</normalizedToken>
longer than broad; lateral margins convex, not posteriorly widened; with two non-corneate eyes; with 52 setae, including four near anterior margin, seven near posterior margin, 13 in the medial zone, and 32 in the anterior region; with two distinct furrows; posterior furrow slightly closer to posterior carapace margin than to median furrow (Fig.
<figureCitation id="3ED6A42C8ABA70D2CAAFE09C2E0CF810" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5A</figureCitation>
); deeply granulated.
</paragraph>
<caption id="B14C2DF68695C3A1F3D4555A0B3CD8B8" doi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301" start="Figure 5" startId="F5">
<paragraph id="74474624EE23AA5797874FF4B544A26F" pageId="0" pageNumber="301">
<emphasis id="BDDC5D0F1DE3FA5EC85A0DF92A5D3FFA" bold="true" pageId="0" pageNumber="301">Figure 5.</emphasis>
<taxonomicName id="2836499F73F29F86997335583C10BCB8" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius florezi" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="CB9C7C3042CD11513A2254E98CCCA35D" italics="true" pageId="0" pageNumber="301">Cystowithius florezi</emphasis>
</taxonomicName>
sp. nov. (male holotype, ICN-Aps-077)
<emphasis id="DF4E32350B673C354E845EB64F22700B" bold="true" pageId="0" pageNumber="301">A</emphasis>
carapace
<emphasis id="90CC0E0F61C550A2382838692DCFD668" bold="true" pageId="0" pageNumber="301">B</emphasis>
chelicera
<emphasis id="25652C2AB7A8BB5609C095E243FE10E9" bold="true" pageId="0" pageNumber="301">C</emphasis>
left leg I
<emphasis id="911027DD91A048D21A7D0CD38843805A" bold="true" pageId="0" pageNumber="301">D</emphasis>
abdomen, sternites
<emphasis id="6ECA45F15F6BAADAE100A623BC1D3905" bold="true" pageId="0" pageNumber="301">E</emphasis>
right chela
<emphasis id="A35788952D26B0B8247C78045CDEA6C5" bold="true" pageId="0" pageNumber="301">F</emphasis>
left leg IV
<emphasis id="F37064F020C5F88BC8137B41A34D7A3C" bold="true" pageId="0" pageNumber="301">G</emphasis>
left pedipalp
<emphasis id="DE7931D35EC17EB4319D70D9AA134AE3" bold="true" pageId="0" pageNumber="301">H</emphasis>
right chelal fingers (arrows show sense-spots)
<emphasis id="51171C319D3747A0735F763F034764D4" bold="true" pageId="0" pageNumber="301">I</emphasis>
rallum. Abbreviations: Pedipalp movable finger:
<emphasis id="0D5F8B6FA0E5DF830E7C97E0FDA36C83" bold="true" italics="true" pageId="0" pageNumber="301">b</emphasis>
= basal;
<emphasis id="D77789B4E8BB1145598C373EBBE6EF00" bold="true" italics="true" pageId="0" pageNumber="301">sb</emphasis>
= sub-basal;
<emphasis id="308B112D00C148323D724567135EEF30" bold="true" italics="true" pageId="0" pageNumber="301">st</emphasis>
= sub-terminal;
<emphasis id="B10125011C7B720BB602C7AA358AAF43" bold="true" italics="true" pageId="0" pageNumber="301">t</emphasis>
= terminal; fixed finger:
<emphasis id="B77DACEBEE1FED3450E23193ADFE7D9B" bold="true" italics="true" pageId="0" pageNumber="301">eb</emphasis>
= exterior basal;
<emphasis id="74E0DFC2C74A888F9AD19E1FF0C23B93" bold="true" italics="true" pageId="0" pageNumber="301">esb</emphasis>
= exterior sub-basal;
<emphasis id="43B99F8F8DBA6100AA8FFFC778F0693F" bold="true" italics="true" pageId="0" pageNumber="301">est</emphasis>
= exterior sub-terminal;
<emphasis id="EC0B54458D0471AB7A6CA4571099DB1B" bold="true" italics="true" pageId="0" pageNumber="301">et</emphasis>
= exterior terminal;
<emphasis id="B48511FF6E58EBE3017CA9A71AF17EB0" bold="true" italics="true" pageId="0" pageNumber="301">ib</emphasis>
= interior basal;
<emphasis id="620F17D8DF44ABCF3D694019F9B5D331" bold="true" italics="true" pageId="0" pageNumber="301">isb</emphasis>
= interior sub-basal;
<emphasis id="86CAF45BB0785EC59DF0D7F5E6E2FB02" bold="true" italics="true" pageId="0" pageNumber="301">ist</emphasis>
= interior sub-terminal;
<emphasis id="C16ADC7E7598304F82FF9696DBA68B41" bold="true" italics="true" pageId="0" pageNumber="301">it</emphasis>
= interior terminal. Cheliceral setae:
<emphasis id="CDF8023AEF0612F6BE75B06A5D997E33" bold="true" italics="true" pageId="0" pageNumber="301">b</emphasis>
= basal seta;
<emphasis id="88F6F7AEC6B7F9AB4A8B84C44B3BEDB6" bold="true" italics="true" pageId="0" pageNumber="301">es</emphasis>
= exterior seta;
<emphasis id="45E86BE2690C96F9A5DB988D00B37A96" bold="true" italics="true" pageId="0" pageNumber="301">is</emphasis>
= interior seta;
<emphasis id="1A42CE0D02AD76DA8612CCB66DD485E6" bold="true" italics="true" pageId="0" pageNumber="301">ls</emphasis>
= laminal seta. Scale bars: 0.5 mm (
<emphasis id="FDBBA482EC67884BD23C6D91F667A13D" bold="true" pageId="0" pageNumber="301">G</emphasis>
); 0.1 mm (
<emphasis id="F9BFC8A81B61A5CBF4D70D71BA3635D4" bold="true" pageId="0" pageNumber="301">A, C-F, H</emphasis>
); 0.05 mm (
<emphasis id="E0FCA4230E116B9115C2F7F435B3DAF9" bold="true" pageId="0" pageNumber="301">B</emphasis>
); 0.0125 mm (
<emphasis id="1C430B66D3EBD05E9222EB3A908598B9" bold="true" pageId="0" pageNumber="301">I</emphasis>
).
</paragraph>
</caption>
<paragraph id="1BC0D56D01741A2FDCFCB130FE54CB62" pageId="0" pageNumber="301">
<emphasis id="36E951D4311DB2E3DA3AE92A7DF84118" bold="true" italics="true" pageId="0" pageNumber="301">Chelicera</emphasis>
(Fig.
<figureCitation id="5F9012C14B0B6AD3B6F48F0AB1E6C4BA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5B</figureCitation>
): with five setae on hand,
<emphasis id="3A537652D7C6C97783C92B7E3759E0D3" italics="true" pageId="0" pageNumber="301">sb</emphasis>
and
<emphasis id="2ECA396EAB9D3899B3D5C93EF3BC72C5" italics="true" pageId="0" pageNumber="301">b</emphasis>
denticulate, all others acuminate; movable finger with one subdistal seta; galea with one sub-terminal and three terminal rami; rallum of four blades, the most distal blade with several serrations on leading edge, other blades smooth (Fig.
<figureCitation id="CB3A6877965AAEF41D3044F65F9A900A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5I</figureCitation>
); lamina exterior present; two dorsal lyrifissures.
</paragraph>
<paragraph id="77B6647D1C69C2DCC24A93054E999B9F" pageId="0" pageNumber="301">
<emphasis id="DDC18A7A73D9BD4E5AA281CB36643B58" bold="true" italics="true" pageId="0" pageNumber="301">Pedipalp</emphasis>
(Fig.
<figureCitation id="39101BD245E0056B91AA3A4CB37171B7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5E, G, H</figureCitation>
): trochanter, femur, patella and chelal hand granulate, chelal fingers smooth; setae clavate and denticulate; trochanter 2.04
<normalizedToken id="E9566857C4A9C644E21CA1DC11FEE740" originalValue="×">x</normalizedToken>
, femur 3.81
<normalizedToken id="8CDE29C9F2DF508BDE495B1D8D632E64" originalValue="×">x</normalizedToken>
, patella 3.30
<normalizedToken id="9CC35D5E1268662EB68C34F97D307CB3" originalValue="×">x</normalizedToken>
, chela (with pedicel) 3.64
<normalizedToken id="ECC585B9F19CDFB65841F1CFDFDC86C0" originalValue="×">x</normalizedToken>
, chela (without pedicel) 3.42
<normalizedToken id="974CF02DCCD5734DFF2894C610CDF856" originalValue="×">x</normalizedToken>
, hand 1.74
<normalizedToken id="2B46FA0426C66410CD5CBD9DE5F767C1" originalValue="×">x</normalizedToken>
longer than broad, movable finger 1.07
<normalizedToken id="EEE3A0ED98D498168405B7CFC2227F7E" originalValue="×">x</normalizedToken>
longer than hand. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria (Fig.
<figureCitation id="330E8A7332F380EE748624364E78160F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5E, H</figureCitation>
):
<emphasis id="832902FC9DBE5505A6D64279EE410724" italics="true" pageId="0" pageNumber="301">eb</emphasis>
and
<emphasis id="236E092563ECFFDA6AAA91217D866C02" italics="true" pageId="0" pageNumber="301">esb</emphasis>
situated basally, as well as
<emphasis id="AEBBBD9227789E9229CC71CD7FCE7478" italics="true" pageId="0" pageNumber="301">ib</emphasis>
and
<emphasis id="3D52073FABDD86836B886F66B348FE6F" italics="true" pageId="0" pageNumber="301">ist</emphasis>
;
<emphasis id="C85DC5FB10D262D8E307E5D6E8732037" italics="true" pageId="0" pageNumber="301">isb</emphasis>
,
<emphasis id="094E5E3A2F400278B75267A6AD757626" italics="true" pageId="0" pageNumber="301">it</emphasis>
and
<emphasis id="C42CFBA78159F13A84305C9F932AB54E" italics="true" pageId="0" pageNumber="301">est</emphasis>
grouped together submedially;
<emphasis id="6E425D3F6B138E9F7969DDB6C243D00D" italics="true" pageId="0" pageNumber="301">et</emphasis>
near the distal end of the finger;
<emphasis id="2CC7DAE36C7193BA6C2843F54EFFECCC" italics="true" pageId="0" pageNumber="301">b</emphasis>
and
<emphasis id="396599A5C345E0EFA48F392A3A5E8A31" italics="true" pageId="0" pageNumber="301">sb</emphasis>
situated near one another;
<emphasis id="D4BB9142466DFBADAFA75A7A57CE6F8A" italics="true" pageId="0" pageNumber="301">st</emphasis>
slightly closer to
<emphasis id="11461A87D0ACDB5F8441A6375EAE7349" italics="true" pageId="0" pageNumber="301">sb</emphasis>
than to
<emphasis id="EB483B9A65A28421D787FF8D4742B4C6" italics="true" pageId="0" pageNumber="301">t</emphasis>
;
<emphasis id="549CCC655781750994320BE2B2D2A1AB" italics="true" pageId="0" pageNumber="301">t</emphasis>
parallel to
<emphasis id="158CDA9118871EFE05B78F07871F02B7" italics="true" pageId="0" pageNumber="301">isb</emphasis>
. Venom apparatus not visible, venom ducts not visible in fixed finger; nodus ramosus distal to
<emphasis id="A61069E00F5B707F6EE9348A156D56BD" italics="true" pageId="0" pageNumber="301">t</emphasis>
on movable finger, not visible on fixed finger. Retrolateral margin of fixed finger with three sense-spots (Fig.
<figureCitation id="6D1ACAC3B9D001CF71C765CD1DC3F613" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5H</figureCitation>
) situated linearly between
<emphasis id="25D64688A5B19458E48A65BEBF29D3CE" italics="true" pageId="0" pageNumber="301">esb</emphasis>
and
<emphasis id="84EBCD784B95674F6E236C34EDF83584" italics="true" pageId="0" pageNumber="301">est</emphasis>
; movable finger with small structure between
<emphasis id="9A3CA8BEA3E15AE98ABAE472598E656D" italics="true" pageId="0" pageNumber="301">t</emphasis>
and
<emphasis id="D5B6EEF36277FA9A66E8433AC302361B" italics="true" pageId="0" pageNumber="301">st</emphasis>
that contains two small nubbins. Chelal teeth rounded with an apical spot; fixed and movable finger with 32 teeth; accessory teeth absent.
</paragraph>
<paragraph id="08129FA821B84E3D56AC227DA6902A2E" pageId="0" pageNumber="301">
<emphasis id="32B2D74D69CFC11D8B667193C1F535AC" bold="true" italics="true" pageId="0" pageNumber="301">Coxal region</emphasis>
: coxal chaetotaxy: 8: 8: 8: 12, with multiple small lyrifissures; maxilla with two apical setae, one very small internal, sub-oral seta and 12 setae; median maxillary lyrifissure medial in position, posterior lyrifissure present.
</paragraph>
<paragraph id="27B152EC89D9983713432D87194DD029" pageId="0" pageNumber="301">
<emphasis id="058F84621291F6F4BDAF5ED0C5211266" bold="true" italics="true" pageId="0" pageNumber="301">Legs</emphasis>
(Fig.
<figureCitation id="259A0F1AE769994A2D5481709794EE7F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5C, F</figureCitation>
): junction between femora and patellae I and II parallel, junction in legs III and IV oblique; tarsal tactile seta of leg IV situated distally, 0.74 of tarsus length (Fig.
<figureCitation id="82A14AD9C0AD0907FD93C26F939240D4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5F</figureCitation>
); subterminal tarsal seta acute, distal to tactile seta; arolium same level as claws. Ratios: leg I: femur 1.06
<normalizedToken id="84885D542CACD364DE542D72CE127738" originalValue="×">x</normalizedToken>
, patella 2.12
<normalizedToken id="C99FA28D048E955FFE815F64C3356841" originalValue="×">x</normalizedToken>
, tibia 2.13
<normalizedToken id="5920261C93EB7D85D2CD9DAAB29CA4EB" originalValue="×">x</normalizedToken>
, tarsus 4.43
<normalizedToken id="B6013C8B6E632B7856CE7C0BFB5EFD78" originalValue="×">x</normalizedToken>
deeper than broad; Leg IV: femur + patella 3.09
<normalizedToken id="C3E12F89BF40459E1B89E85233ACC876" originalValue="×">x</normalizedToken>
, tibia 2.35
<normalizedToken id="DE9C6E50CFE5B0F08F15D8C8202A97C5" originalValue="×">x</normalizedToken>
, tarsus 4.78
<normalizedToken id="939EDE889FE90BAF12CBC2BBF568BEA5" originalValue="×">x</normalizedToken>
deeper than broad.
</paragraph>
<paragraph id="2FEDC8D00C447F9C76F4F86C72EEECBE" pageId="0" pageNumber="301">
<emphasis id="01F0AF9B405077A67E1C26678A9482C2" bold="true" italics="true" pageId="0" pageNumber="301">Abdomen</emphasis>
(Fig.
<figureCitation id="F9FE5E83F4D475DBF9F15B91A52F6E6C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5D</figureCitation>
): all tergites divided but the first two only with a faint medial suture; all sternites entire. Tergal chaetotaxy: 8: 7: 8: 14: 15: 13: 15: 15: 15: 13: 8 (including two tactile setae): 2; mostly uniseriate but some tergites with a few setae placed anteriorly; all setae foliate, except in the last tergite; tergites densely granulated. Sternal chaetotaxy: 10: (2) 9 (2): (2) 9 (2): 12: 12: 12: 2 + 40
<emphasis id="B14C40064AD69B412B622327DC1937D9" italics="true" pageId="0" pageNumber="301">gls</emphasis>
: 9: 8: 9 (including two tactile and three clavate setae): 2; only sternite VIII with a small circular patch of glandular setae (Fig.
<figureCitation id="0450FF7BD045E4B88CC3B950E42683A5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glandular setae patches of the new species A Paciwithius chimbilacus sp. nov. (holotype, ICN-Aps- 076) B Parawithius bromelicola sp. nov. (holotype, ICN-Aps- 082) C Paciwithius valduparensis sp. nov. (holotype, ICN-Aps- 597) D Cystowithius florezi sp. nov. (holotype, ICN-Aps- 07) E, F Oligowithius achagua sp. nov. E male paratype (ICN-Aps- 388) F male holotype (ICN-Aps- 388). Scale bars: 0.1 mm (A-C, E, F); 0.05 mm (D)." figureDoi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301">9D</figureCitation>
); with paired invaginations on anterior margins of sternites VI-VIII (Fig.
<figureCitation id="8BBEACE02457869B32D0E22FB9930BC9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cystowithius florezi sp. nov. (male holotype, ICN-Aps- 077) A carapace B chelicera C left leg I D abdomen, sternites E right chela F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, C-F, H); 0.05 mm (B); 0.0125 mm (I)." figureDoi="10.3897/zookeys.1184.106698.figure5" httpUri="https://binary.pensoft.net/fig/940874" pageId="0" pageNumber="301">5D</figureCitation>
).
</paragraph>
<paragraph id="85037893A90AA4C4F5D9B74682C66E5E" pageId="0" pageNumber="301">
<emphasis id="1673D8866E3B30EB59AD931549E3D95E" bold="true" italics="true" pageId="0" pageNumber="301">Genitalia</emphasis>
: see
<bibRefCitation id="A0937930BC7F13A7AD8091DDA4E8CF32" DOI="https://doi.org/10.1636/JoA-S-19-068" author="Romero-Ortiz, C" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="301" pagination="108 - 121" refId="B23" refString="Romero-Ortiz, C, Sarmiento, CE, 2021. A comparative study of the male genitalia of the Cacodemoniini (Pseudoscorpiones: Withiidae). The Journal of Arachnology 49 (1): 108 - 121, DOI: https://doi.org/10.1636/JoA-S-19-068" title="A comparative study of the male genitalia of the Cacodemoniini (Pseudoscorpiones: Withiidae)." url="https://doi.org/10.1636/JoA-S-19-068" volume="49" year="2021">Romero-Ortiz and Sarmiento 2021</bibRefCitation>
as &quot;
<taxonomicName id="51F07DC317499A9E2994BB9A5CBF039D" authorityName="Harvey" authorityYear="2004" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="87933AC3ED96C8DBED2C1681B40B47A4" italics="true" pageId="0" pageNumber="301">Cystowithius</emphasis>
</taxonomicName>
msp1&quot;.
</paragraph>
</subSubSection>
<subSubSection id="5440D2A109201023A28C1C8C0FC61375" pageId="0" pageNumber="301" type="etymology">
<paragraph id="7419F71F9A02FF0788314FA3F69D322D" pageId="0" pageNumber="301">Etymology.</paragraph>
<paragraph id="F8373E91E5A831700826A38FB7613843" pageId="0" pageNumber="301">This species is dedicated to Professor Eduardo Florez, considered the father of Arachnology in Colombia. He has also been the curator of the Arachnological collection of the Instituto de Ciencias Naturales, where all the material used in this study is lodged.</paragraph>
</subSubSection>
</treatment>
</document>
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