treatments-xml/data/6F/53/C6/6F53C675875CFF9EFF7BF8B71ECCB56D.xml

<document id="0D73444CD31ACEF297F422527D51D0DE" ID-CLB-Dataset="298743" ID-DOI="10.11646/zootaxa.5471.4.3" ID-GBIF-Dataset="7ffa6c79-8784-4c3f-b7b8-1e562dd9f607" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190343" ID-ZooBank="7CD97873-F527-4EE5-98C4-0297B17A26EC" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1718893888664" checkinUser="plazi" docAuthor="Hernández-Austria, Raquel, García-Castillo, Mirna G. &amp; Parra-Olea, Gabriela" docDate="2024" docId="6F53C675875CFF9EFF7BF8B71ECCB56D" docLanguage="en" docName="zootaxa.5471.4.3.pdf" docOrigin="Zootaxa 5471 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eleutherodactylus (Syrrhophus) coelum Hernández-Austria &amp; García-Castillo &amp; Parra-Olea 2024, sp. nov." docType="treatment" docVersion="7" lastPageNumber="447" masterDocId="936ABE0D875BFF90FFECFFC61D5BB42D" masterDocTitle="A new species of direct-developing frog of the genus Eleutherodactylus (Anura: Eleutherodactylidae) from Tamaulipas, Mexico" masterLastPageNumber="450" masterPageNumber="433" pageNumber="440" updateTime="1719009346240" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DA2874C540CC3FB3C7F265CD2A0693DA">A new species of direct-developing frog of the genus Eleutherodactylus (Anura: Eleutherodactylidae) from Tamaulipas, Mexico</mods:title>
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<mods:namePart id="181F3874901432C4BBC11C5713D6761A">Hernández-Austria, Raquel</mods:namePart>
<mods:affiliation id="ED46FB073716BFD98BCADF41BD6C9D4B">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, A. P. 70 - 153, 04510, Ciudad de Mexico, México. &amp; Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México, A. P. 70 - 153, 04510, Ciudad de Mexico, México.</mods:affiliation>
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<mods:namePart id="CEF21D4BA9EE2BEB6300546347087B6F">García-Castillo, Mirna G.</mods:namePart>
<mods:affiliation id="952B64020CB84D4A45E8912EFC98D3C2">Facultad de Ciencias, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, Ciudad de México, México.</mods:affiliation>
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<mods:namePart id="E66CBE0163C1347FB2ED3B240384BA02">Parra-Olea, Gabriela</mods:namePart>
<mods:affiliation id="106D10FB1C50024A7E88B5CDEE89FD30">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, A. P. 70 - 153, 04510, Ciudad de Mexico, México.</mods:affiliation>
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<subSubSection id="AFE024E8875CFF97FF7BF8B71FBBB3A1" box="[151,736,1905,1932]" pageId="7" pageNumber="440" type="nomenclature">
<paragraph id="E7457763875CFF97FF7BF8B71FBBB3A1" blockId="7.[151,736,1905,1968]" box="[151,736,1905,1932]" pageId="7" pageNumber="440">
<heading id="BC0DC00F875CFF97FF7BF8B71FBBB3A1" bold="true" box="[151,736,1905,1932]" fontSize="11" level="1" pageId="7" pageNumber="440" reason="1">
<emphasis id="D58EAB71875CFF97FF7BF8B71FBBB3A1" bold="true" box="[151,736,1905,1932]" pageId="7" pageNumber="440">
<taxonomicName id="20FA0CE0875CFF97FF7BF8B71F20B3A1" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[151,635,1905,1932]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="440" phylum="Chordata" rank="species" species="coelum" status="sp. nov." subGenus="Syrrhophus">
<emphasis id="D58EAB71875CFF97FF7BF8B71C28B3A1" bold="true" box="[151,371,1905,1932]" italics="true" pageId="7" pageNumber="440">Eleutherodactylus</emphasis>
(
<emphasis id="D58EAB71875CFF97FE68F8B71F49B3A1" bold="true" box="[388,530,1905,1932]" italics="true" pageId="7" pageNumber="440">Syrrhophus</emphasis>
) 
<emphasis id="D58EAB71875CFF97FDCFF8B71F20B3A1" bold="true" box="[547,635,1905,1932]" italics="true" pageId="7" pageNumber="440">coelum</emphasis>
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<taxonomicNameLabel id="CEBD160A875CFF97FD6FF8B41FBBB3A1" box="[643,736,1906,1932]" pageId="7" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="AFE024E8875CFF97FF7BF8501C17B39D" box="[151,332,1942,1968]" pageId="7" pageNumber="440" type="description">
<paragraph id="E7457763875CFF97FF7BF8501C17B39D" blockId="7.[151,736,1905,1968]" box="[151,332,1942,1968]" pageId="7" pageNumber="440">
(
<figureCitation id="7FC16BE6875CFF97FF73F8501DBEB39D" box="[159,229,1942,1968]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="7" pageNumber="440">Fig. 4</figureCitation>
; 
<tableCitation id="AA7842D8875CFF97FF1CF8501C18B39D" box="[240,323,1942,1968]" captionStart="TABLE 3" captionStartId="10.[152,244,594,620]" captionTargetPageId="10" captionText="TABLE 3. Comparative morphological characters of the Eleutherodactylus longipes species clade. Morphological information was obtained by examining type specimens (E. potosiensis) or photographs of type specimens of all species (except for E. marnockii)." httpUri="http://table.plazi.org/id/B38527EB8751FF9AFF74FD941969B6BD" pageId="7" pageNumber="440" tableUuid="B38527EB8751FF9AFF74FD941969B6BD">Table 3</tableCitation>
)
</paragraph>
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<subSubSection id="AFE024E8875CFF97FF7BF8181FA4B3D5" box="[151,767,2014,2040]" pageId="7" pageNumber="440" type="vernacular_names">
<paragraph id="E7457763875CFF97FF7BF8181FA4B3D5" blockId="7.[151,767,2014,2040]" box="[151,767,2014,2040]" pageId="7" pageNumber="440">El Cielo Chirping Frog / Rana Chirriadora de El Cielo:</paragraph>
</subSubSection>
<subSubSection id="AFE024E88753FF98FF2BFF51180CB64C" pageId="8" pageNumber="441" type="materials_examined">
<paragraph id="E74577638753FF98FF2BFF511E27B4D4" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="441">
<materialsCitation id="57927D3E8753FF98FF2BFF511E27B4D4" ID-GBIF-Occurrence="4898140302" collectingDate="2018-09-13" collectionCode="IBH, WGS" collectorName="C. Omar Becerra Soria &amp; M. Delia Basanta" country="Mexico" elevation="350" latitude="23.042309" location="El Cielo Biosphere Reserve" longLatPrecision="1" longitude="-99.15486" pageId="8" pageNumber="441" specimenCode="IBH 36351, WGS84" specimenCount="1" stateProvince="Tamaulipas" typeStatus="holotype">
<emphasis id="D58EAB718753FF98FF2BFF511C60B49C" bold="true" box="[199,315,151,177]" pageId="8" pageNumber="441">
<typeStatus id="3841C9C18753FF98FF2BFF511C6DB49C" box="[199,310,151,177]" pageId="8" pageNumber="441" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="B75CDF188753FF98FEAAFF511C90B49C" box="[326,459,150,177]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 36351</specimenCode>
, an adult male from Gómez Farías (
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, - 
<geoCoordinate id="82CE11A48753FF98FBC1FF501997B49C" box="[1069,1228,150,177]" degrees="99.154858" direction="west" orientation="longitude" pageId="8" pageNumber="441" precision="1" value="-99.15486">99.154858°W</geoCoordinate>
, 
<specimenCode id="B75CDF188753FF98FB37FF511862B49C" box="[1243,1337,151,177]" collectionCode="WGS" pageId="8" pageNumber="441">WGS84</specimenCode>
), 
<quantity id="2002DA868753FF98FABDFF5018C7B49C" box="[1361,1436,150,177]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="441" unit="m" value="350.0">
<elevation id="6CD790508753FF98FABDFF5018C7B49C" box="[1361,1436,150,177]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="441" unit="m" value="350.0">350 m</elevation>
</quantity>
elevation, in the 
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, municipality of Gómez Farías, 
<collectingRegion id="253EB9818753FF98FC11FF7D19DAB4F8" box="[1021,1153,187,213]" country="Mexico" name="Tamaulipas" pageId="8" pageNumber="441">Tamaulipas</collectingRegion>
, 
<collectingCountry id="9FED37F38753FF98FB61FF7D19BEB4F8" box="[1165,1253,187,213]" name="Mexico" pageId="8" pageNumber="441">Mexico</collectingCountry>
, collected by 
<collectorName id="4A0F12B58753FF98FA6EFF7D1C2CB4D4" pageId="8" pageNumber="441">C. Omar Becerra Soria</collectorName>
and 
<collectorName id="4A0F12B58753FF98FE42FF191F2AB4D4" box="[430,625,223,249]" pageId="8" pageNumber="441">M. Delia Basanta</collectorName>
on 
<date id="934451A38753FF98FD77FF191E23B4D4" box="[667,888,223,249]" pageId="8" pageNumber="441" value="2018-09-13">
<collectingDate id="8300A84B8753FF98FD77FF191E23B4D4" box="[667,888,223,249]" pageId="8" pageNumber="441" value="2018-09-13">13 September 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="E74577638753FF98FF2BFEC51F29B5A4" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="441">
<materialsCitation id="57927D3E8753FF98FF2BFEC51FD7B530" ID-GBIF-Occurrence="4898140310" box="[199,652,259,285]" collectingDate="2018-09-13" collectionCode="IBH" collectorName="C. Omar Becerra Soria &amp; M. Delia Basanta" country="Mexico" elevation="950" latitude="23.042309" location="El Cielo Biosphere Reserve" longLatPrecision="1" longitude="-99.15486" municipality="Gomez Farias" pageId="8" pageNumber="441" specimenCode="IBH 36349, IBH 36352, IBH 36350, IBH 36353, IBH 36354" specimenCount="5" stateProvince="Tamaulipas" typeStatus="paratype">
<emphasis id="D58EAB718753FF98FF2BFEC51C1DB530" bold="true" box="[199,326,259,285]" pageId="8" pageNumber="441">
<typeStatus id="3841C9C18753FF98FF2BFEC51C1AB530" box="[199,321,259,285]" pageId="8" pageNumber="441" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Five specimens. 
<specimenCode id="B75CDF188753FF98FDE1FEC51FD7B530" box="[525,652,259,285]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 36349</specimenCode>
</materialsCitation>
<materialsCitation id="57927D3E8753FF98FD78FEC51977B530" ID-GBIF-Occurrence="4898140315" box="[660,1068,258,285]" collectionCode="IBH" country="Mexico" elevation="950" latitude="23.042309" location="El Cielo Biosphere Reserve" longitude="-99.15486" municipality="Gomez Farias" pageId="8" pageNumber="441" specimenCode="IBH 36349, IBH 36352, IBH 36350, IBH 36353, IBH 36354" specimenCount="5" stateProvince="Tamaulipas" typeStatus="paratype">
and 
<specimenCode id="B75CDF188753FF98FD29FEC51E1CB530" box="[709,839,258,285]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 36352</specimenCode>
, both adult females,
</materialsCitation>
<materialsCitation id="57927D3E8753FF98FBD8FEC5184EB530" ID-GBIF-Occurrence="4898140307" box="[1076,1301,258,285]" collectionCode="IBH" country="Mexico" elevation="950" latitude="23.042309" location="El Cielo Biosphere Reserve" longitude="-99.15486" municipality="Gomez Farias" pageId="8" pageNumber="441" specimenCode="IBH 36349, IBH 36352, IBH 36350, IBH 36353, IBH 36354" specimenCount="5" stateProvince="Tamaulipas" typeStatus="paratype">
and 
<specimenCode id="B75CDF188753FF98FB89FEC519BFB530" box="[1125,1252,258,285]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 36350</specimenCode>
and
</materialsCitation>
<materialsCitation id="57927D3E8753FF98FAF1FEC51EE2B56C" ID-GBIF-Occurrence="4898140304" collectionCode="IBH" country="Mexico" elevation="950" latitude="23.042309" location="El Cielo Biosphere Reserve" longitude="-99.15486" municipality="Gomez Farias" pageId="8" pageNumber="441" specimenCode="IBH 36349, IBH 36352, IBH 36350, IBH 36353, IBH 36354" specimenCount="5" stateProvince="Tamaulipas" typeStatus="paratype">
<specimenCode id="B75CDF188753FF98FAF1FEC518C7B530" box="[1309,1436,258,285]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 36353</specimenCode>
adult males from the same locality and recollection data as the holotype.
</materialsCitation>
<materialsCitation id="57927D3E8753FF98FC53FEE11F35B5A4" ID-GBIF-Occurrence="4898140313" collectionCode="IBH" country="Mexico" elevation="950" latitude="23.042309" location="El Cielo Biosphere Reserve" longitude="-99.202869" municipality="Gomez Farias" pageId="8" pageNumber="441" specimenCode="IBH 36349, IBH 36352, IBH 36350, IBH 36353, IBH 36354" specimenCount="5" stateProvince="Tamaulipas" typeStatus="paratype">
<specimenCode id="B75CDF188753FF98FC53FEE11960B56C" box="[959,1083,294,321]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 36354</specimenCode>
from Alta Cima (
<geoCoordinate id="82CE11A48753FF98FB11FEE118C3B56C" box="[1277,1432,295,321]" degrees="23.064144" direction="north" orientation="latitude" pageId="8" pageNumber="441" precision="1" value="23.064144">23.064144°N</geoCoordinate>
, - 
<geoCoordinate id="82CE11A48753FF98FF73FE8D1C60B548" box="[159,315,331,357]" degrees="99.202869" direction="north" orientation="latitude" pageId="8" pageNumber="441" precision="1" value="99.202866">99.202869°N</geoCoordinate>
), 950 m elevation, in the 
<location id="E22521B88753FF98FDBBFE8D1ECBB548" LSID="urn:lsid:plazi:treatment:6F53C675875CFF9EFF7BF8B71ECCB56D:E22521B88753FF98FDBBFE8D1ECBB548" box="[599,912,331,357]" country="Mexico" latitude="99.202866" municipality="Gomez Farias" name="El Cielo Biosphere Reserve" pageId="8" pageNumber="441" stateProvince="Tamaulipas">El Cielo Biosphere Reserve</location>
, 
<collectingMunicipality id="0721ED198753FF98FC77FE8D1961B548" box="[923,1082,331,357]" pageId="8" pageNumber="441">Gómez Farías</collectingMunicipality>
, 
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, 
<collectingCountry id="9FED37F38753FF98FB38FE8D1877B548" box="[1236,1324,331,357]" name="Mexico" pageId="8" pageNumber="441">
<collectingRegion id="253EB9818753FF98FB38FE8D1877B548" box="[1236,1324,331,357]" country="Mexico" name="Mexico" pageId="8" pageNumber="441">Mexico</collectingRegion>
</collectingCountry>
, collected by the same individuals than the holotype
</materialsCitation>
.
</paragraph>
<paragraph id="E74577638753FF98FF2BFE54180CB64C" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="441">
<emphasis id="D58EAB718753FF98FF2BFE541CE1B581" bold="true" box="[199,442,402,428]" pageId="8" pageNumber="441">Referred specimens.</emphasis>
<materialsCitation id="57927D3E8753FF98FE2DFE5518C2B5D8" ID-GBIF-Occurrence="4898140309" collectingDate="2010-08-02" collectionCode="IBH, WGS" collectorName="Sean M. Rovito &amp; Gabriela Parra-Olea &amp; Thomas J. Devitt &amp; Susan Cameron &amp; Jorge Sanchez &amp; Eli Garcia-Padilla" county="El Cielo Biosphere Reserve" elevation="1382" latitude="23.06062" location="Alta Cima" longLatPrecision="1" longitude="-99.22171" municipality="San Jose" pageId="8" pageNumber="441" specimenCode="IBH 30477, WGS84" specimenCount="1">
<specimenCode id="B75CDF188753FF98FE2DFE551F65B580" box="[449,574,403,429]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 30477</specimenCode>
from on road from 
<location id="E22521B88753FF98FCF0FE551ECBB580" LSID="urn:lsid:plazi:treatment:6F53C675875CFF9EFF7BF8B71ECCB56D:E22521B88753FF98FCF0FE551ECBB580" box="[796,912,403,429]" county="El Cielo Biosphere Reserve" latitude="23.06062" longLatPrecision="1" longitude="-99.22171" municipality="San Jose" name="Alta Cima" pageId="8" pageNumber="441">Alta Cima</location>
to 
<location id="E22521B88753FF98FC58FE551943B580" LSID="urn:lsid:plazi:treatment:6F53C675875CFF9EFF7BF8B71ECCB56D:E22521B88753FF98FC58FE551943B580" box="[948,1048,403,429]" county="El Cielo Biosphere Reserve" latitude="23.06062" longLatPrecision="1" longitude="-99.22171" municipality="San Jose" name="San Jose" pageId="8" pageNumber="441">San José</location>
, at 
<locationDeviation id="F6F76E418753FF98FBD3FE5519E1B580" bearing="45.0" box="[1087,1210,403,429]" distance="1700.0" pageId="8" pageNumber="441">
<quantity id="2002DA868753FF98FBD3FE5519D5B580" box="[1087,1166,403,429]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="441" unit="km" value="1.7">1.7 km</quantity>
NE
</locationDeviation>
(by air) of 
<collectingMunicipality id="0721ED198753FF98FAD7FE5518C7B580" box="[1339,1436,403,429]" pageId="8" pageNumber="441">San José</collectingMunicipality>
(
<geoCoordinate id="82CE11A48753FF98FF73FE711C77B5FC" box="[159,300,439,465]" degrees="23.06062" direction="north" orientation="latitude" pageId="8" pageNumber="441" precision="1" value="23.06062">23.06062°N</geoCoordinate>
, - 
<geoCoordinate id="82CE11A48753FF98FEACFE711C8AB5FC" box="[320,465,439,465]" degrees="99.22171" direction="west" orientation="longitude" pageId="8" pageNumber="441" precision="1" value="-99.22171">99.22171°W</geoCoordinate>
, 
<specimenCode id="B75CDF188753FF98FE32FE711F67B5FC" box="[478,572,439,465]" collectionCode="WGS" pageId="8" pageNumber="441">WGS84</specimenCode>
), 
<quantity id="2002DA868753FF98FDBDFE711FF1B5FC" box="[593,682,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.382" pageId="8" pageNumber="441" unit="m" value="1382.0">
<elevation id="6CD790508753FF98FDBDFE711FF1B5FC" box="[593,682,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.382" pageId="8" pageNumber="441" unit="m" value="1382.0">1382 m</elevation>
</quantity>
, in the 
<collectingCounty id="0E240FEF8753FF98FCEEFE71191AB5FC" box="[770,1089,439,465]" pageId="8" pageNumber="441">El Cielo Biosphere Reserve</collectingCounty>
, collected by 
<collectorName id="4A0F12B58753FF98FB0DFE7118C3B5FC" box="[1249,1432,439,465]" pageId="8" pageNumber="441">Sean M. Rovito</collectorName>
, 
<collectorName id="4A0F12B58753FF98FF7BFE1D1C27B5D8" box="[151,380,475,501]" pageId="8" pageNumber="441">Gabriela Parra-Olea</collectorName>
, 
<collectorName id="4A0F12B58753FF98FE6BFE1D1F17B5D8" box="[391,588,475,501]" pageId="8" pageNumber="441">Thomas J. Devitt</collectorName>
, 
<collectorName id="4A0F12B58753FF98FDB4FE1D1E56B5D8" box="[600,781,475,501]" pageId="8" pageNumber="441">Susan Cameron</collectorName>
, 
<collectorName id="4A0F12B58753FF98FCF5FE1D1EE6B5D8" box="[793,957,475,501]" pageId="8" pageNumber="441">Jorge Sánchez</collectorName>
, and 
<collectorName id="4A0F12B58753FF98FC16FE1D199FB5D8" box="[1018,1220,475,501]" pageId="8" pageNumber="441">Elí Garcia-Padilla</collectorName>
on 
<date id="934451A38753FF98FB1CFE1D18C2B5D8" box="[1264,1433,475,501]" pageId="8" pageNumber="441" value="2010-08-02">
<collectingDate id="8300A84B8753FF98FB1CFE1D18C2B5D8" box="[1264,1433,475,501]" pageId="8" pageNumber="441" value="2010-08-02">2 August 2010</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="57927D3E8753FF98FF7BFE391FB2B610" ID-GBIF-Occurrence="4898140316" collectingDate="2010-08-02" collectionCode="IBH" collectorName="Sean M. Rovito &amp; Gabriela Parra-Olea &amp; Thomas J. Devitt &amp; Susan Cameron &amp; Jorge Sanchez &amp; Eli Garcia-Padilla" county="El Cielo Biosphere Reserve" elevation="360" latitude="23.06593" location="Centro Interpretativo Ecologico" longLatPrecision="5" longitude="-99.1687" municipality="El Cielo Biosphere Reserve" pageId="8" pageNumber="441" specimenCode="IBH 30480" specimenCount="1">
<specimenCode id="B75CDF188753FF98FF7BFE391C4CB634" box="[151,279,511,537]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 30480</specimenCode>
from 
<location id="E22521B88753FF98FE8DFE391F90B634" LSID="urn:lsid:plazi:treatment:6F53C675875CFF9EFF7BF8B71ECCB56D:E22521B88753FF98FE8DFE391F90B634" box="[353,715,511,537]" county="El Cielo Biosphere Reserve" latitude="23.06593" longLatPrecision="5" longitude="-99.1687" municipality="El Cielo Biosphere Reserve" name="Centro Interpretativo Ecologico" pageId="8" pageNumber="441">Centro Interpretativo Ecológico</location>
(
<geoCoordinate id="82CE11A48753FF98FD31FE381E32B634" box="[733,873,510,537]" degrees="23.06593" direction="north" orientation="latitude" pageId="8" pageNumber="441" precision="1" value="23.06593">23.06593°N</geoCoordinate>
, - 
<geoCoordinate id="82CE11A48753FF98FC93FE39195EB634" box="[895,1029,511,537]" degrees="99.1687" direction="west" orientation="longitude" pageId="8" pageNumber="441" precision="5" value="-99.1687">99.1687°W</geoCoordinate>
), 
<quantity id="2002DA868753FF98FBF7FE39193CB634" box="[1051,1127,511,537]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="441" unit="m" value="360.0">
<elevation id="6CD790508753FF98FBF7FE39193CB634" box="[1051,1127,511,537]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="441" unit="m" value="360.0">360 m</elevation>
</quantity>
, in the 
<collectingMunicipality id="0721ED198753FF98FB2DFE391DAFB610" pageId="8" pageNumber="441">El Cielo Biosphere Reserve</collectingMunicipality>
, with the same recollection data as the IBH
</materialsCitation>
30477. 
<materialsCitation id="57927D3E8753FF98FCA5FDE5180AB610" ID-GBIF-Occurrence="4898140317" box="[841,1361,547,573]" collectingDate="2010-08-02" collectionCode="IBH" collectorName="Sean M. Rovito &amp; Gabriela Parra-Olea &amp; Thomas J. Devitt &amp; Susan Cameron &amp; Jorge Sanchez &amp; Eli Garcia-Padilla" county="El Cielo Biosphere Reserve" elevation="360" latitude="23.06593" longLatPrecision="5" longitude="-99.1687" municipality="El Cielo Biosphere Reserve" pageId="8" pageNumber="441" specimenCode="IBH 32711" specimenCount="1" specimenCount-adult="1" specimenCount-male="1">
<specimenCode id="B75CDF188753FF98FCA5FDE51E91B610" box="[841,970,547,573]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 32711</specimenCode>
, an adult 
<specimenCount id="F1FCBCEA8753FF98FBD7FDE5192AB610" box="[1083,1137,547,573]" pageId="8" pageNumber="441" type="male">male</specimenCount>
with the same data
</materialsCitation>
of the 
<typeStatus id="3841C9C18753FF98FF7BFD811DA6B64C" box="[151,253,583,609]" pageId="8" pageNumber="441" type="holotype">holotype</typeStatus>
. 
<materialsCitation id="57927D3E8753FF98FEEBFD811E70B64C" ID-GBIF-Occurrence="4898140320" box="[263,811,583,609]" collectingDate="2010-08-02" collectionCode="IBH" collectorName="Sean M. Rovito &amp; Gabriela Parra-Olea &amp; Thomas J. Devitt &amp; Susan Cameron &amp; Jorge Sanchez &amp; Eli Garcia-Padilla" county="El Cielo Biosphere Reserve" elevation="360" latitude="23.06593" longLatPrecision="5" longitude="-99.1687" municipality="El Cielo Biosphere Reserve" pageId="8" pageNumber="441" specimenCode="IBH 32712" specimenCount="1" specimenCount-male="1">
<specimenCode id="B75CDF188753FF98FEEBFD811CDCB64C" box="[263,391,583,609]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 32712</specimenCode>
, an adult 
<specimenCount id="F1FCBCEA8753FF98FE1FFD811F72B64C" box="[499,553,583,609]" pageId="8" pageNumber="441" type="male">male</specimenCount>
from the same locality
</materialsCitation>
<materialsCitation id="57927D3E8753FF98FCDEFD811991B64C" ID-GBIF-Occurrence="4898140311" box="[818,1226,583,609]" collectingDate="2010-08-02" collectionCode="IBH" collectorName="Sean M. Rovito &amp; Gabriela Parra-Olea &amp; Thomas J. Devitt &amp; Susan Cameron &amp; Jorge Sanchez &amp; Eli Garcia-Padilla" county="El Cielo Biosphere Reserve" elevation="360" latitude="23.06593" longLatPrecision="5" longitude="-99.1687" municipality="El Cielo Biosphere Reserve" pageId="8" pageNumber="441" specimenCount="1" typeStatus="as the paratype">
and recollection data 
<typeStatus id="3841C9C18753FF98FBCFFD811991B64C" box="[1059,1226,583,609]" pageId="8" pageNumber="441">as the paratype</typeStatus>
</materialsCitation>
<materialsCitation id="57927D3E8753FF98FB3EFD811808B64C" ID-GBIF-Occurrence="4898140314" box="[1234,1363,582,609]" collectionCode="IBH" pageId="8" pageNumber="441" specimenCode="IBH 36354" specimenCount="1">
<specimenCode id="B75CDF188753FF98FB3EFD811808B64C" box="[1234,1363,582,609]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="8" pageNumber="441">IBH 36354</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AFE024E88753FF98FF2BFDAC1E70B075" pageId="8" pageNumber="441" type="diagnosis">
<paragraph id="E74577638753FF98FF2BFDAC1CF2B7C0" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="441">
<emphasis id="D58EAB718753FF98FF2BFDAC1C19B6A9" bold="true" box="[199,322,618,644]" pageId="8" pageNumber="441">Diagnosis.</emphasis>
A member of the genus 
<taxonomicName id="20FA0CE08753FF98FD64FDAD1E08B6A8" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[648,851,619,645]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="genus">
<emphasis id="D58EAB718753FF98FD64FDAD1E08B6A8" box="[648,851,619,645]" italics="true" pageId="8" pageNumber="441">Eleutherodactylus</emphasis>
</taxonomicName>
, subgenus 
<taxonomicName id="20FA0CE08753FF98FC0EFDAD1938B6A8" box="[994,1123,619,645]" pageId="8" pageNumber="441" rank="subGenus" subGenus="Syrrhophus">
<emphasis id="D58EAB718753FF98FC0EFDAD1938B6A8" box="[994,1123,619,645]" italics="true" pageId="8" pageNumber="441">Syrrhophus</emphasis>
</taxonomicName>
, in the 
<taxonomicName id="20FA0CE08753FF98FB3EFDAD1CCBB684" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="longipes" subGenus="Syrrhophus">
<emphasis id="D58EAB718753FF98FB3EFDAD18C6B6A8" box="[1234,1437,619,645]" italics="true" pageId="8" pageNumber="441">Eleutherodactylus</emphasis>
(
<emphasis id="D58EAB718753FF98FF4DFD491C79B684" box="[161,290,655,681]" italics="true" pageId="8" pageNumber="441">Syrrhophus</emphasis>
) 
<emphasis id="D58EAB718753FF98FEDDFD491CCBB684" box="[305,400,655,681]" italics="true" pageId="8" pageNumber="441">longipes</emphasis>
</taxonomicName>
clade as defined by 
<bibRefCitation id="836B0A928753FF98FD98FD491E80B684" author="Hernandez-Austria, R. &amp; Garcia-Vazquez, U. O. &amp; Grunwald, C. I. &amp; Parra-Olea, G." box="[628,987,655,681]" pageId="8" pageNumber="441" pagination="1 - 20" refId="ref9960" refString="Hernandez-Austria, R., Garcia-Vazquez, U. O., Grunwald, C. I. &amp; Parra-Olea, G. (2022) Molecular phylogeny of the subgenus Syrrhophus (Amphibia: Anura: Eleutherodactylidae), with the description of a new species from Eastern Mexico. Systematics and Biodiversity, 20, 1 - 20. https: // doi. org / 10.1080 / 14772000.2021.2014597" type="journal article" year="2022">
Hernández-Austria 
<emphasis id="D58EAB718753FF98FCA3FD491EDBB684" box="[847,896,655,681]" italics="true" pageId="8" pageNumber="441">et al</emphasis>
. (2022)
</bibRefCitation>
, morphologically most like 
<taxonomicName id="20FA0CE08753FF98FAFFFD49182DB685" box="[1299,1398,655,681]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="campi">
<emphasis id="D58EAB718753FF98FAFFFD49187FB685" box="[1299,1316,655,680]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FADEFD49182DB685" box="[1330,1398,655,680]" italics="true" pageId="8" pageNumber="441">campi</emphasis>
</taxonomicName>
, 
<taxonomicName id="20FA0CE08753FF98FA68FD491C1EB6E0" class="Amphibia" family="Dendrobatidae" genus="Phyllobates" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="cystignathoides">
<emphasis id="D58EAB718753FF98FA68FD4918CEB685" box="[1412,1429,655,680]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FF7BFD751C1EB6E0" box="[151,325,691,717]" italics="true" pageId="8" pageNumber="441">cystignathoides</emphasis>
</taxonomicName>
, and 
<taxonomicName id="20FA0CE08753FF98FE6CFD751F45B6E1" box="[384,542,691,717]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="potosiensis">
<emphasis id="D58EAB718753FF98FE6CFD751CCAB6E1" box="[384,401,691,716]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FE72FD751F45B6E1" box="[414,542,691,716]" italics="true" pageId="8" pageNumber="441">potosiensis</emphasis>
</taxonomicName>
<emphasis id="D58EAB718753FF98FDF2FD751E12B6E0" box="[542,841,691,717]" italics="true" pageId="8" pageNumber="441">
. 
<taxonomicName id="20FA0CE08753FF98FDCBFD751E12B6E0" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[551,841,691,717]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">Eleutherodactylus coelum</taxonomicName>
</emphasis>
<emphasis id="D58EAB718753FF98FCA3FD751EFCB6E0" bold="true" box="[847,935,691,717]" pageId="8" pageNumber="441">
<taxonomicNameLabel id="CEBD160A8753FF98FCA3FD751EFCB6E0" box="[847,935,691,717]" pageId="8" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other species of the subgenus by the following combination of characters: small-size frog with a mean SVL 
<quantity id="2002DA868753FF98FBC0FD1119F9B6DC" box="[1068,1186,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.7269999999999999" pageId="8" pageNumber="441" unit="mm" value="17.27">17.27 mm</quantity>
± 
<quantity id="2002DA868753FF98FB2AFD10184CB6DC" box="[1222,1303,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.9116" pageId="8" pageNumber="441" unit="in" value="1.54">1.54 in</quantity>
<specimenCount id="F1FCBCEA8753FF98FACEFD1118C7B6DC" box="[1314,1436,727,753]" count="8" pageId="8" pageNumber="441" type="adult">eight adult</specimenCount>
males (range 
<quantity id="2002DA868753FF98FEDEFD3C1CAFB738" box="[306,500,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="8" pageNumber="441" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
) and mean SVL 
<quantity id="2002DA868753FF98FD55FD3D1E76B738" box="[697,813,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" pageId="8" pageNumber="441" unit="mm" value="16.64">16.64 mm</quantity>
± 
<quantity id="2002DA868753FF98FCA1FD3D1EC6B738" box="[845,925,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.7272" pageId="8" pageNumber="441" unit="in" value="0.68">0.68 in</quantity>
<specimenCount id="F1FCBCEA8753FF98FC4AFD3D194AB738" box="[934,1041,763,789]" count="2" pageId="8" pageNumber="441" type="adult">two adult</specimenCount>
females (range 
<quantity id="2002DA868753FF98FB27FD3D18D4B738" box="[1227,1423,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="8" pageNumber="441" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
); interorbital bar present (
<figureCitation id="7FC16BE68753FF98FE4AFCD91F5AB714" box="[422,513,799,825]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="8" pageNumber="441">Fig. 4A</figureCitation>
); snout truncate in lateral view, and rounded in dorsal view; tympanum membrane round and annulus distinct, interrupted above (
<figureCitation id="7FC16BE68753FF98FD40FC851E5DB770" box="[684,774,835,861]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="8" pageNumber="441">Fig. 4B</figureCitation>
); Finger I shorter than Finger II; fingertips truncated with pads having a terminal groove; Finger III and IV having dilated pads 1.4–2.3 times width of the narrowest part of these fingers (
<figureCitation id="7FC16BE68753FF98FED5FC4D1CC8B788" box="[313,403,907,933]" captionStart="FIGURE 3" captionStartId="7.[151,249,1451,1475]" captionTargetBox="[172,1418,186,1419]" captionTargetId="figure-19@7.[161,1426,181,1428]" captionTargetPageId="7" captionText="FIGURE 3. Non-Metric Multidimensional Scaling (nMDS) ordination of 24 morphometrical variables of Eleutherodactylus coelum sp. nov. (magenta), E. cystignathoides (green), E. dennisi (blue), E. potosiensis (orange), and E. sp. Ahuacatlán (black)." figureDoi="http://doi.org/10.5281/zenodo.12190351" httpUri="https://zenodo.org/record/12190351/files/figure.png" pageId="8" pageNumber="441">Fig. 3C</figureCitation>
); palmar subarticular tubercles large and conical; outer metatarsal tubercle round, conical, 1.3–2.2 times larger than the ovoid inner metatarsal tubercle; skin of the dorsum granular; diffuse dark-colored transverse bands in legs.
</paragraph>
<paragraph id="E74577638753FF98FF2BFC301E70B075" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="441">
<emphasis id="D58EAB718753FF98FF2BFC301C36B03D" bold="true" box="[199,365,1014,1040]" pageId="8" pageNumber="441">Comparisons.</emphasis>
We limited our comparison to members belonging to the 
<taxonomicName id="20FA0CE08753FF98FBB1FC3119BDB03C" box="[1117,1254,1015,1041]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="longipes">
<emphasis id="D58EAB718753FF98FBB1FC311935B03D" box="[1117,1134,1015,1040]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FB6BFC3119BDB03C" box="[1159,1254,1015,1041]" italics="true" pageId="8" pageNumber="441">longipes</emphasis>
</taxonomicName>
species clade. 
<taxonomicName id="20FA0CE08753FF98FF7BFBDC1CE5B019" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[151,446,1050,1076]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718753FF98FF7BFBDC1CE5B019" box="[151,446,1050,1076]" italics="true" pageId="8" pageNumber="441">Eleutherodactylus coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718753FF98FE24FBDC1F7DB019" bold="true" box="[456,550,1050,1076]" pageId="8" pageNumber="441">
<taxonomicNameLabel id="CEBD160A8753FF98FE24FBDC1F7DB019" box="[456,550,1050,1076]" pageId="8" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other species of the 
<taxonomicName id="20FA0CE08753FF98FC58FBDD196DB019" box="[948,1078,1050,1077]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="longipes">
<emphasis id="D58EAB718753FF98FC58FBDD1E9EB019" box="[948,965,1051,1076]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FC3BFBDC196DB019" box="[983,1078,1050,1076]" italics="true" pageId="8" pageNumber="441">longipes</emphasis>
</taxonomicName>
species clade in the following characters (condition of 
<taxonomicName id="20FA0CE08753FF98FE44FBF91F43B075" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[424,536,1086,1113]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718753FF98FE44FBF91CE2B075" box="[424,441,1087,1112]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FE2BFBF81F43B075" box="[455,536,1086,1112]" italics="true" pageId="8" pageNumber="441">coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718753FF98FDF3FBF81F22B075" bold="true" box="[543,633,1086,1112]" pageId="8" pageNumber="441">
<taxonomicNameLabel id="CEBD160A8753FF98FDF3FBF81F22B075" box="[543,633,1086,1112]" pageId="8" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in parenthesis).
</paragraph>
</subSubSection>
<subSubSection id="AFE024E88753FF9DFF2BFBA41EE8B6A8" lastPageId="13" lastPageNumber="446" pageId="8" pageNumber="441" type="description">
<paragraph id="E74577638753FF98FF2BFBA4194EB17F" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="441">
<taxonomicName id="20FA0CE08753FF98FF2BFBA41CE3B051" box="[199,440,1122,1148]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="D58EAB718753FF98FF2BFBA41CC9B051" box="[199,402,1122,1148]" italics="true" pageId="8" pageNumber="441">Eleutherodactylus</emphasis>
sp.
</taxonomicName>
Ahuacatlán differs from 
<taxonomicName id="20FA0CE08753FF98FD3CFBA51E64B051" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[720,831,1122,1148]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718753FF98FD3CFBA51FBAB051" box="[720,737,1123,1148]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FD02FBA41E64B051" box="[750,831,1122,1148]" italics="true" pageId="8" pageNumber="441">coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718753FF98FCAAFBA41EC4B051" bold="true" box="[838,927,1122,1148]" pageId="8" pageNumber="441">
<taxonomicNameLabel id="CEBD160A8753FF98FCAAFBA41EC4B051" box="[838,927,1122,1148]" pageId="8" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a larger size, 
<specimenCount id="F1FCBCEA8753FF98FB42FBA41872B051" box="[1198,1321,1122,1148]" count="1" pageId="8" pageNumber="441" type="female">one female</specimenCount>
with SVL 
<quantity id="2002DA868753FF98FF7BFB401C56B08D" box="[151,269,1158,1184]" metricMagnitude="-2" metricUnit="m" metricValue="2.307" pageId="8" pageNumber="441" unit="mm" value="23.07">23.07 mm</quantity>
(females SVL 
<quantity id="2002DA868753FF98FE2CFB431FDDB08D" box="[448,646,1157,1184]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="8" pageNumber="441" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), and a smaller tympanum, TD/EW=0.16 (TD/EW 0.91); 
<taxonomicName id="20FA0CE08753FF98FAC5FB4018C7B0B2" box="[1321,1436,1157,1184]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="dennisi">
<emphasis id="D58EAB718753FF98FAC5FB401861B0B2" box="[1321,1338,1158,1183]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FAA7FB4318C7B0B2" box="[1355,1436,1157,1183]" italics="true" pageId="8" pageNumber="441">dennisi</emphasis>
</taxonomicName>
has a larger size, 
<specimenCount id="F1FCBCEA8753FF98FE8DFB6C1C92B0EE" box="[353,457,1193,1220]" count="1" pageId="8" pageNumber="441" type="male">one male</specimenCount>
with SVL 
<quantity id="2002DA868753FF98FDA6FB6F1FE4B0E9" box="[586,703,1193,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.464" pageId="8" pageNumber="441" unit="mm" value="24.64">24.64 mm</quantity>
and males SVL 
<quantity id="2002DA868753FF98FC6DFB6F1919B0E9" box="[897,1090,1193,1220]" metricMagnitude="-2" metricUnit="m" metricValue="3.3095" metricValueMax="3.4829999999999997" metricValueMin="3.136" pageId="8" pageNumber="441" unit="mm" value="33.095" valueMax="34.83" valueMin="31.36">31.36–34.83 mm</quantity>
(males SVL 
<quantity id="2002DA868753FF98FB37FB6F18C6B0E9" box="[1243,1437,1193,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="8" pageNumber="441" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
and females SVL 
<quantity id="2002DA868753FF98FE88FB0B1F72B0CA" box="[356,553,1229,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="8" pageNumber="441" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), digital tips of the Finger III and Finger IV expanded 3.1–3.7 times width of the narrowest part of these same fingers (digital tips of the Finger III and Finger IV expanded 1.4–2.3 times width of the narrowest part of these same fingers), and by having a dorsal coloration light brown to tan with brown vermiculations (dorsum with brown markings without forming vermiculations).
</paragraph>
<paragraph id="E74577638753FF98FF2BFA9A1FCFB29A" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="441">
<taxonomicName id="20FA0CE08753FF98FF2BFA9A1F16B15B" box="[199,589,1372,1398]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="cystignathoides">
<emphasis id="D58EAB718753FF98FF2BFA9A1F16B15B" box="[199,589,1372,1398]" italics="true" pageId="8" pageNumber="441">Eleutherodactylus cystignathoides</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="20FA0CE08753FF98FD19FA9B1E37B15B" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[757,876,1372,1399]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718753FF98FD19FA9B1E5DB15B" box="[757,774,1373,1398]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FCF7FA9A1E37B15B" box="[795,876,1372,1398]" italics="true" pageId="8" pageNumber="441">coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718753FF98FC95FA9A1E82B15B" bold="true" box="[889,985,1372,1398]" pageId="8" pageNumber="441">
<taxonomicNameLabel id="CEBD160A8753FF98FC95FA9A1E82B15B" box="[889,985,1372,1398]" pageId="8" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a smaller tympanum, TD/ EW= 
<quantity id="2002DA868753FF98FF38FA461C78B1B7" box="[212,291,1408,1434]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" pageId="8" pageNumber="441" unit="in" value="1.03">1.03 in</quantity>
males and TD/EW= 
<quantity id="2002DA868753FF98FDFEFA461F38B1B7" box="[530,611,1408,1434]" metricMagnitude="-2" metricUnit="m" metricValue="1.8796" pageId="8" pageNumber="441" unit="in" value="0.74">0.74 in</quantity>
females (TD/EW= 
<quantity id="2002DA868753FF98FCACFA461ECBB1B7" box="[832,912,1408,1434]" metricMagnitude="-2" metricUnit="m" metricValue="2.8956" pageId="8" pageNumber="441" unit="in" value="1.14">1.14 in</quantity>
males and TD/EW= 
<quantity id="2002DA868753FF98FB6CFA461994B1B7" box="[1152,1231,1408,1434]" metricMagnitude="-2" metricUnit="m" metricValue="2.3114" pageId="8" pageNumber="441" unit="in" value="0.91">0.91 in</quantity>
females), smaller tibia, TL/SVL= 
<quantity id="2002DA868753FF98FEA8FA651CCAB190" box="[324,401,1443,1469]" metricMagnitude="-2" metricUnit="m" metricValue="1.1684" pageId="8" pageNumber="441" unit="in" value="0.46">0.46 in</quantity>
males and TL/SVL= 
<quantity id="2002DA868753FF98FD6EFA621F94B190" box="[642,719,1443,1470]" metricMagnitude="-2" metricUnit="m" metricValue="1.0922" pageId="8" pageNumber="441" unit="in" value="0.43">0.43 in</quantity>
females (TL/SVL= 
<quantity id="2002DA868753FF98FC45FA651EACB190" box="[937,1015,1443,1470]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="441" unit="in" value="0.5">0.50 in</quantity>
males and TL/SVL= 
<quantity id="2002DA868753FF98FB0EFA621874B190" box="[1250,1327,1443,1470]" metricMagnitude="-2" metricUnit="m" metricValue="1.1938" pageId="8" pageNumber="441" unit="in" value="0.47">0.47 in</quantity>
females), and rounded subarticular tubercles (conical subarticular tubercles); 
<taxonomicName id="20FA0CE08753FF98FC7FFA0E1EACB1CC" box="[915,1015,1480,1506]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="campi">
<emphasis id="D58EAB718753FF98FC7FFA0E1EFFB1CC" box="[915,932,1480,1505]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FC5FFA0E1EACB1CC" box="[947,1015,1480,1505]" italics="true" pageId="8" pageNumber="441">campi</emphasis>
</taxonomicName>
has a coarsely granular dorsum (less granular dorsum), generally no interorbital bar (interorbital bar present), and rounded subarticular tubercles (conical subarticular tubercles); 
<taxonomicName id="20FA0CE08753FF98FE4CF9C91F60B205" box="[416,571,1551,1577]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="potosiensis">
<emphasis id="D58EAB718753FF98FE4CF9C91CEAB205" box="[416,433,1551,1576]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FE53F9C91F60B205" box="[447,571,1551,1576]" italics="true" pageId="8" pageNumber="441">potosiensis</emphasis>
</taxonomicName>
has a larger size, with males SVL 
<quantity id="2002DA868753FF98FC2DF9C919DBB204" box="[961,1152,1551,1577]" metricMagnitude="-2" metricUnit="m" metricValue="2.2195" metricValueMax="2.316" metricValueMin="2.123" pageId="8" pageNumber="441" unit="mm" value="22.195" valueMax="23.16" valueMin="21.23">21.23–23.16 mm</quantity>
and females SVL 
<quantity id="2002DA868753FF98FAA3F9C91C51B260" metricMagnitude="-2" metricUnit="m" metricValue="2.503" metricValueMax="2.7239999999999998" metricValueMin="2.282" pageId="8" pageNumber="441" unit="mm" value="25.03" valueMax="27.24" valueMin="22.82">22.82– 27.24 mm</quantity>
(males SVL 
<quantity id="2002DA868753FF98FE4DF9F41F3AB260" box="[417,609,1586,1613]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="8" pageNumber="441" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
and females SVL 
<quantity id="2002DA868753FF98FCD9F9F41EA2B261" box="[821,1017,1586,1612]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="8" pageNumber="441" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), digital tips of Finger III and Finger IV expanded 1.52–1.72 times width of the narrowest part of these same fingers (digital tips of Finger III and Finger IV expanded 1.4–2.3 times width of the narrowest part of these same fingers), and belly unpigmented, without markings (belly with a few brown markings).
</paragraph>
<paragraph id="E74577638753FF99FF2BF9071900B5FC" blockId="8.[151,1437,150,2043]" lastBlockId="9.[151,1437,150,1221]" lastPageId="9" lastPageNumber="442" pageId="8" pageNumber="441">
<taxonomicName id="20FA0CE08753FF98FF2BF9071CA3B2F6" box="[199,504,1729,1755]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="longipes">
<emphasis id="D58EAB718753FF98FF2BF9071CA3B2F6" box="[199,504,1729,1755]" italics="true" pageId="8" pageNumber="441">Eleutherodactylus longipes</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="20FA0CE08753FF98FD60F9041FA6B2F6" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[652,765,1729,1756]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718753FF98FD60F9041FC6B2F6" box="[652,669,1730,1755]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FD40F9071FA6B2F6" box="[684,765,1729,1755]" italics="true" pageId="8" pageNumber="441">coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718753FF98FCE8F9071E06B2F6" bold="true" box="[772,861,1729,1755]" pageId="8" pageNumber="441">
<taxonomicNameLabel id="CEBD160A8753FF98FCE8F9071E06B2F6" box="[772,861,1729,1755]" pageId="8" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a larger size, with males SVL 
<quantity id="2002DA868753FF98FAC0F9071D98B32D" metricMagnitude="-2" metricUnit="m" metricValue="2.765" metricValueMax="3.32" metricValueMin="2.21" pageId="8" pageNumber="441" unit="mm" value="27.65" valueMax="33.2" valueMin="22.1">22.1–33.2 mm</quantity>
and females SVL 26.8–39. 
<quantity id="2002DA868753FF98FE1BF9231F6CB32D" box="[503,567,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="441" unit="mm" value="6.0">6 mm</quantity>
(males SVL 
<quantity id="2002DA868753FF98FD24F9231EDDB32D" box="[712,902,1765,1792]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="8" pageNumber="441" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
and females SVL 
<quantity id="2002DA868753FF98FBBEF923184FB2D2" box="[1106,1300,1765,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="8" pageNumber="441" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), digital tips of Finger III and Finger IV greatly expanded, more than twice the width of the narrowest part of these same fingers (digital tips of Finger III and Finger IV expanded 1.4–2.3 times width of the narrowest part of these same fingers), and dorsum with irregular brown blotches, spots, or reticulations on a light background (dorsum with scattered olive-brown markings without forming vermiculations on light background); 
<taxonomicName id="20FA0CE08753FF98FC14F8B31907B3A3" box="[1016,1116,1908,1935]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="leprus">
<emphasis id="D58EAB718753FF98FC14F8B31952B3A3" box="[1016,1033,1909,1934]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FBFAF8B21907B3A3" box="[1046,1116,1908,1934]" italics="true" pageId="8" pageNumber="441">leprus</emphasis>
</taxonomicName>
has a larger size, with males SVL 
<quantity id="2002DA868753FF98FF3FF85E1C2DB39E" box="[211,374,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.65" metricValueMin="2.05" pageId="8" pageNumber="441" unit="mm" value="23.5" valueMax="26.5" valueMin="20.5">20.5–26.5 mm</quantity>
and females SVL 22.0– 
<quantity id="2002DA868753FF98FD6BF85E1FB0B39E" box="[647,747,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="8" pageNumber="441" unit="mm" value="29.3">29.3 mm</quantity>
(males SVL 
<quantity id="2002DA868753FF98FC6CF85E1964B39E" box="[896,1087,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="8" pageNumber="441" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
and females SVL 
<quantity id="2002DA868753FF98FAFCF85E1D9CB3FB" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="8" pageNumber="441" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), dorsum with brown, yellow or green blotches or spots forming reticulations on a dark background (dorsum with scattered olive-brown markings without forming vermiculations on a light background); 
<taxonomicName id="20FA0CE08753FF98FB2AF8271802B3D7" box="[1222,1369,2016,2043]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="441" phylum="Chordata" rank="species" species="marnockii">
<emphasis id="D58EAB718753FF98FB2AF827198CB3D7" box="[1222,1239,2017,2042]" italics="true" pageId="8" pageNumber="441">E</emphasis>
. 
<emphasis id="D58EAB718753FF98FB0BF8261802B3D7" box="[1255,1369,2016,2042]" italics="true" pageId="8" pageNumber="441">marnockii</emphasis>
</taxonomicName>
has a larger size, with males SVL 
<quantity id="2002DA868752FF99FE3AFF511F21B49C" box="[470,634,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.3649999999999998" metricValueMax="2.8899999999999997" metricValueMin="1.8399999999999999" pageId="9" pageNumber="442" unit="mm" value="23.65" valueMax="28.9" valueMin="18.4">18.4–28.9 mm</quantity>
and females SVL 20.4–35.4 (males SVL 
<quantity id="2002DA868752FF99FBA3FF501855B49C" box="[1103,1294,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="9" pageNumber="442" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
and females SVL 
<quantity id="2002DA868752FF99FF3AFF7D1CC6B4F8" box="[214,413,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="9" pageNumber="442" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), and dorsum tan to light brown with rusty-brown marks (dorsum with scattered olive-brown markings on light background), and interorbital bar absent (interorbital bar present); 
<taxonomicName id="20FA0CE08752FF99FB63FF191843B4D4" box="[1167,1304,223,249]" class="Amphibia" family="Eleutherodactylidae" genus="Malachylodes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="442" phylum="Chordata" rank="species" species="guttilatus">
<emphasis id="D58EAB718752FF99FB63FF1919FBB4D5" box="[1167,1184,223,248]" italics="true" pageId="9" pageNumber="442">E</emphasis>
. 
<emphasis id="D58EAB718752FF99FB42FF191843B4D4" box="[1198,1304,223,249]" italics="true" pageId="9" pageNumber="442">guttilatus</emphasis>
</taxonomicName>
has a larger size, with males SVL 20.6–29.0 mm and females SVL 25.7–31.0 mm (males SVL 
<quantity id="2002DA868752FF99FBA7FEC41857B530" box="[1099,1292,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="9" pageNumber="442" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
and females SVL 
<quantity id="2002DA868752FF99FF3EFEE11CCFB56C" box="[210,404,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="9" pageNumber="442" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), lumbo-inguinal gland present (lumbo-inguinal gland absent), areolate ventral skin (smooth ventral skin); 
<taxonomicName id="20FA0CE08752FF99FEDCFE8D1C9FB549" box="[304,452,331,357]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="442" phylum="Chordata" rank="species" species="verrucipes">
<emphasis id="D58EAB718752FF99FEDCFE8D1C1AB549" box="[304,321,331,356]" italics="true" pageId="9" pageNumber="442">E</emphasis>
. 
<emphasis id="D58EAB718752FF99FEA2FE8D1C9FB549" box="[334,452,331,356]" italics="true" pageId="9" pageNumber="442">verrucipes</emphasis>
</taxonomicName>
has a larger size, with males SVL 
<quantity id="2002DA868752FF99FCD3FE8C1EBBB548" box="[831,992,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" metricValueMax="2.61" metricValueMin="1.75" pageId="9" pageNumber="442" unit="mm" value="21.8" valueMax="26.1" valueMin="17.5">17.5–26.1 mm</quantity>
and females SVL 28.0– 
<quantity id="2002DA868752FF99FB06FE8D1817B548" box="[1258,1356,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.17" pageId="9" pageNumber="442" unit="mm" value="31.7">31.7 mm</quantity>
(males SVL 
<quantity id="2002DA868752FF99FF3AFEA81CC2B5A4" box="[214,409,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.6695" metricValueMax="1.988" metricValueMin="1.351" pageId="9" pageNumber="442" unit="mm" value="16.695" valueMax="19.88" valueMin="13.51">13.51–19.88 mm</quantity>
and females SVL 
<quantity id="2002DA868752FF99FD94FEA91E64B5A4" box="[632,831,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" metricValueMax="1.712" metricValueMin="1.616" pageId="9" pageNumber="442" unit="mm" value="16.64" valueMax="17.12" valueMin="16.16">16.16–17.12 mm</quantity>
), and lumbo-inguinal gland present (lumbo-inguinal gland absent), areolate ventral skin (smooth ventral skin), and interorbital bar absent or present (interorbital bar present). General characteristics for the 
<taxonomicName id="20FA0CE08752FF99FDBBFE711F8EB5FC" box="[599,725,439,465]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="442" phylum="Chordata" rank="species" species="longipes">
<emphasis id="D58EAB718752FF99FDBBFE711F8EB5FC" box="[599,725,439,465]" italics="true" pageId="9" pageNumber="442">E. longipes</emphasis>
</taxonomicName>
species clade are given in 
<tableCitation id="AA7842D88752FF99FBEDFE71190EB5FC" box="[1025,1109,439,465]" captionStart="TABLE 3" captionStartId="10.[152,244,594,620]" captionTargetPageId="10" captionText="TABLE 3. Comparative morphological characters of the Eleutherodactylus longipes species clade. Morphological information was obtained by examining type specimens (E. potosiensis) or photographs of type specimens of all species (except for E. marnockii)." httpUri="http://table.plazi.org/id/B38527EB8751FF9AFF74FD941969B6BD" pageId="9" pageNumber="442" tableUuid="B38527EB8751FF9AFF74FD941969B6BD">Table 3</tableCitation>
.
</paragraph>
<paragraph id="E74577638752FF99FF2BFE1C1C35B7C0" blockId="9.[151,1437,150,1221]" pageId="9" pageNumber="442">
<emphasis id="D58EAB718752FF99FF2BFE1C1F42B5D8" bold="true" box="[199,537,474,501]" pageId="9" pageNumber="442">
Description of the 
<typeStatus id="3841C9C18752FF99FE47FE1D1F4FB5D8" box="[427,532,475,501]" pageId="9" pageNumber="442" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult of small size (
<quantity id="2002DA868752FF99FCFDFE1D1E26B5D8" box="[785,893,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="9" pageNumber="442" unit="mm" value="18.7">18.7 mm</quantity>
); head longer than wide (HL 
<quantity id="2002DA868752FF99FB38FE1D1860B5D8" box="[1236,1339,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="442" unit="mm" value="7.62">7.62 mm</quantity>
vs. HW 
<quantity id="2002DA868752FF99FF7BFE391C5BB634" box="[151,256,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="7.16" pageId="9" pageNumber="442" unit="mm" value="7.16">7.16 mm</quantity>
); interorbital bar diffuse (
<figureCitation id="7FC16BE68752FF99FDC8FE391FDBB634" box="[548,640,511,537]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="9" pageNumber="442">Fig. 4A</figureCitation>
); snout truncate in lateral view (
<figureCitation id="7FC16BE68752FF99FC1CFE391912B634" box="[1008,1097,511,537]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="9" pageNumber="442">Fig. 4B</figureCitation>
), and rounded in dorsal view; nostrils small, directed laterally and much nearer the tip of snout than the eye; eyes large, with a length similar to the END (EL 
<quantity id="2002DA868752FF99FED8FD811CD1B64C" box="[308,394,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="442" unit="mm" value="2.3">2.3 mm</quantity>
vs END 
<quantity id="2002DA868752FF99FE1EFD811F01B64C" box="[498,602,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.43" pageId="9" pageNumber="442" unit="mm" value="2.43">2.43 mm</quantity>
), prominent, directed laterally; lower lip with a few pale spots; tympanum distinct, circular, its annulus not interrupted above and membrane visible; tympanum large, spans 80% of the eye; poorly developed supratympanic fold; a pair of low, and rounded postrictal tubercles; vocal slits present; subgular vocal sac not evident. Forelimbs relatively short and slim; fingers lack interdigital webbing; Finger I shorter than Finger II; fingertips truncated and expanded; digital pad on Finger III expanded 1.7 times the width of the narrowest part of that same finger; palmar subarticular tubercles large and conical (
<figureCitation id="7FC16BE68752FF99FC0DFD3D1966B738" box="[993,1085,763,789]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="9" pageNumber="442">Fig. 4C</figureCitation>
); three palmar tubercles: inner palmar tubercle slightly projection in elevation, oval middle palmar tubercle, and outer palmar tubercle almost imperceptible; supernumerary tubercles are not numerous (
<figureCitation id="7FC16BE68752FF99FCCBFC851E24B770" box="[807,895,835,861]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="9" pageNumber="442">Fig. 4C</figureCitation>
). Inguinal gland absent; skin on dorsum granular and venter smooth, and areolate in flanks. Hindlimbs are long; when the leg is extended forward along the body, the heels reach the anterior corner of the eye; the inner metatarsal tubercle is large and elongated; the outer metatarsal tubercle is small, 1.5 times the size of the inner, slightly prominent; tips of the toes truncated; toes lack interdigital webbing (
<figureCitation id="7FC16BE68752FF99FEEBFC151C39B7C0" box="[263,354,979,1005]" captionStart="FIGURE 4" captionStartId="9.[151,250,1931,1955]" captionTargetBox="[262,1326,1252,1908]" captionTargetId="figure-508@9.[248,1340,1245,1908]" captionTargetPageId="9" captionText="FIGURE 4. A) Photo in life of an individual of Eleutherodactylus coelum sp. nov. from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of E. coelum sp. nov. (IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D)." figureDoi="http://doi.org/10.5281/zenodo.12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="9" pageNumber="442">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="E74577638752FF99FF2BFC301C65B0E8" blockId="9.[151,1437,150,1221]" pageId="9" pageNumber="442">
<emphasis id="D58EAB718752FF99FF2BFC301FE2B03D" bold="true" box="[199,697,1014,1041]" pageId="9" pageNumber="442">
Coloration of the 
<typeStatus id="3841C9C18752FF99FE7AFC311CA6B03C" box="[406,509,1015,1041]" pageId="9" pageNumber="442" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Lateral aspect of the canthus rostralis with a stripe Sepia (279) that fades inferiorly to Olive-Brown (278); the stripe extends posterior to the eye across the tympanum; interorbital bar Olive-Brown (278); dorsal background of head, body and limbs Light Buff (2); many Olive-Brown (278) markings on the back with two ill-defined longitudinal bands Light Buff; legs with irregular transverse dark bands Olive-Brown (278); venter surface Pale Buff (1); chin, ventral surface of thighs and legs sprinkled with Vandyke Brown pigment (281).
</paragraph>
<caption id="B38527EB8752FF99FF7BF84D19BFB3C6" ID-DOI="http://doi.org/10.5281/zenodo.12190353" ID-Zenodo-Dep="12190353" httpUri="https://zenodo.org/record/12190353/files/figure.png" pageId="9" pageNumber="442" startId="9.[151,250,1931,1955]" targetBox="[262,1326,1252,1908]" targetPageId="9" targetType="figure">
<paragraph id="E74577638752FF99FF7BF84D19BFB3C6" blockId="9.[151,1437,1931,2027]" pageId="9" pageNumber="442">
<emphasis id="D58EAB718752FF99FF7BF84D1C4AB38E" bold="true" box="[151,273,1931,1955]" pageId="9" pageNumber="442">FIGURE 4.</emphasis>
A) Photo in life of an individual of 
<taxonomicName id="20FA0CE08752FF99FD83F84D1E2FB38E" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[623,884,1931,1955]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="442" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718752FF99FD83F84D1E2FB38E" box="[623,884,1931,1955]" italics="true" pageId="9" pageNumber="442">Eleutherodactylus coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718752FF99FC95F84D1E93B38E" bold="true" box="[889,968,1931,1955]" pageId="9" pageNumber="442">
<taxonomicNameLabel id="CEBD160A8752FF99FC95F84D1E93B38E" box="[889,968,1931,1955]" pageId="9" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from El Cielo Biosphere Reserve, Tamaulipas, Mexico. B) Head in lateral view of the holotype of 
<taxonomicName id="20FA0CE08752FF99FD77F8761E5BB3EA" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[667,768,1967,1991]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="442" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718752FF99FD77F8761FF1B3EA" box="[667,682,1968,1991]" italics="true" pageId="9" pageNumber="442">E</emphasis>
. 
<emphasis id="D58EAB718752FF99FD5BF8691E5BB3EA" box="[695,768,1967,1991]" italics="true" pageId="9" pageNumber="442">coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718752FF99FCEAF8691E0DB3EA" bold="true" box="[774,854,1967,1991]" pageId="9" pageNumber="442">
<taxonomicNameLabel id="CEBD160A8752FF99FCEAF8691E0DB3EA" box="[774,854,1967,1991]" pageId="9" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(IBH 36351) in preservative; C) hand in ventral view; D) foot in ventral view. Scale bar is 1 mm. Photos: M. Delia Basanta (4A) and Susana Guzmán Gómez (4B–D).
</paragraph>
</caption>
<caption id="B38527EB8751FF9AFF74FD941969B6BD" ID-Table-UUID="B38527EB8751FF9AFF74FD941969B6BD" httpUri="http://table.plazi.org/id/B38527EB8751FF9AFF74FD941969B6BD" pageId="10" pageNumber="443" startId="10.[152,244,594,620]" targetBox="[162,2026,671,1595]" targetIsTable="true" targetPageId="10" targetType="table">
<paragraph id="E74577638751FF9AFF74FD941969B6BD" blockId="10.[151,2037,594,656]" pageId="10" pageNumber="443">
<emphasis id="D58EAB718751FF9AFF74FD941C52B641" bold="true" box="[152,265,594,620]" pageId="10" pageNumber="443">TABLE 3</emphasis>
. Comparative morphological characters of the 
<taxonomicName id="20FA0CE08751FF9AFCEBFD941972B641" box="[775,1065,594,620]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="longipes">
<emphasis id="D58EAB718751FF9AFCEBFD941972B641" box="[775,1065,594,620]" italics="true" pageId="10" pageNumber="443">Eleutherodactylus longipes</emphasis>
</taxonomicName>
species clade. Morphological information was obtained by examining type specimens (
<taxonomicName id="20FA0CE08751FF9AF832FD941C55B6BD" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="potosiensis">
<emphasis id="D58EAB718751FF9AF832FD941AB5B641" box="[2014,2030,594,620]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFF7BFDB01C55B6BD" box="[151,270,630,656]" italics="true" pageId="10" pageNumber="443">potosiensis</emphasis>
</taxonomicName>
) or photographs of type specimens of all species (except for 
<taxonomicName id="20FA0CE08751FF9AFC74FDB01979B6BD" box="[920,1058,630,656]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="marnockii">
<emphasis id="D58EAB718751FF9AFC74FDB01EF3B6BD" box="[920,936,630,656]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFC5AFDB01979B6BD" box="[950,1058,630,656]" italics="true" pageId="10" pageNumber="443">marnockii</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="E74577638751FF9AFF4EFD591D81B216" pageId="10" pageNumber="443">
<table id="95FA85C38751006FFF4EFD591AB1B216" box="[162,2026,671,1595]" gridcols="11" gridrows="25" pageId="10" pageNumber="443">
<tr id="59CA75218751006FFF4EFD591AB1B695" box="[162,2026,671,696]" gridrow="0" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFD591C65B695" box="[162,318,671,696]" gridcol="0" gridrow="0" pageId="10" pageNumber="443">Species</th>
<th id="1A1B1C5D8751006FFEB1FD591C99B695" box="[349,450,671,696]" gridcol="1" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AFEB1FD591CECB695" box="[349,439,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="campi">
<emphasis id="D58EAB718751FF9AFEB1FD591C37B695" box="[349,364,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFE95FD591CECB695" box="[377,439,671,696]" italics="true" pageId="10" pageNumber="443">campi</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FFE17FD591F22B695" box="[507,633,671,696]" gridcol="2" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AFE17FD591F3BB695" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[507,608,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="coelum">
<emphasis id="D58EAB718751FF9AFE17FD591F51B695" box="[507,522,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFDFBFD591F3BB695" box="[535,608,671,696]" italics="true" pageId="10" pageNumber="443">coelum</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FFD74FD591E08B695" box="[664,851,671,696]" gridcol="3" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AFD74FD591E08B695" box="[664,851,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="cystignathoides">
<emphasis id="D58EAB718751FF9AFD74FD591FFCB695" box="[664,679,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFD59FD591E08B695" box="[693,851,671,696]" italics="true" pageId="10" pageNumber="443">cystignathoides</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FFC91FD591955B695" box="[893,1038,671,696]" gridcol="4" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AFC91FD591EB9B695" box="[893,994,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="dennisi">
<emphasis id="D58EAB718751FF9AFC91FD591ED7B695" box="[893,908,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFC75FD591EB9B695" box="[921,994,671,696]" italics="true" pageId="10" pageNumber="443">dennisi</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FFBDCFD5919F7B695" box="[1072,1196,671,696]" gridcol="5" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AFBDCFD5919F7B695" box="[1072,1196,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Malachylodes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="guttilatus">
<emphasis id="D58EAB718751FF9AFBDCFD591964B695" box="[1072,1087,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFBA0FD5919F7B695" box="[1100,1196,671,696]" italics="true" pageId="10" pageNumber="443">guttilatus</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FFB32FD59181DB695" box="[1246,1350,671,696]" gridcol="6" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AFB32FD591862B695" box="[1246,1337,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="leprus">
<emphasis id="D58EAB718751FF9AFB32FD5919B6B695" box="[1246,1261,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFB16FD591862B695" box="[1274,1337,671,696]" italics="true" pageId="10" pageNumber="443">leprus</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FFA93FD5918AAB695" box="[1407,1521,671,696]" gridcol="7" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AFA93FD5918AAB695" box="[1407,1521,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="longipes">
<emphasis id="D58EAB718751FF9AFA93FD5918D5B695" box="[1407,1422,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AFA77FD5918AAB695" box="[1435,1521,671,696]" italics="true" pageId="10" pageNumber="443">longipes</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FF9FBFD591BFBB695" box="[1559,1696,671,696]" gridcol="8" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AF9FBFD591BC1B695" box="[1559,1690,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="marnockii">
<emphasis id="D58EAB718751FF9AF9FBFD591B7DB695" box="[1559,1574,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AF9DFFD591BC1B695" box="[1587,1690,671,696]" italics="true" pageId="10" pageNumber="443">marnockii</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FF95BFD591A1FB695" box="[1719,1860,671,696]" gridcol="9" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AF95BFD591A1FB695" box="[1719,1860,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="potosiensis">
<emphasis id="D58EAB718751FF9AF95BFD591A1FB695" box="[1719,1860,671,696]" italics="true" pageId="10" pageNumber="443">E. potosiensis</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8751006FF88FFD591AB1B695" box="[1891,2026,671,696]" gridcol="10" gridrow="0" pageId="10" pageNumber="443">
<taxonomicName id="20FA0CE08751FF9AF88FFD591AB1B695" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[1891,2026,671,696]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="443" phylum="Chordata" rank="species" species="verrucipes" status="sp. nov.">
<emphasis id="D58EAB718751FF9AF88FFD591A29B695" box="[1891,1906,671,696]" italics="true" pageId="10" pageNumber="443">E</emphasis>
. 
<emphasis id="D58EAB718751FF9AF893FD591AB1B695" box="[1919,2026,671,696]" italics="true" pageId="10" pageNumber="443">verrucipes</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="59CA75218751006FFF4EFD051AB1B6F1" box="[162,2026,707,732]" gridrow="1" pageId="10" pageNumber="443" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td id="1A1B1C5D8751006FFE17FD051F22B6F1" box="[507,633,707,732]" gridcol="2" gridrow="1" pageId="10" pageNumber="443">
<emphasis id="D58EAB718751FF9AFE17FD051F10B6F1" bold="true" box="[507,587,707,732]" pageId="10" pageNumber="443">
<taxonomicNameLabel id="CEBD160A8751FF9AFE17FD051F10B6F1" box="[507,587,707,732]" pageId="10" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="59CA75218751006FFF4EFD2D1AB1B729" box="[162,2026,747,772]" gridrow="2" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFD2D1C65B729" box="[162,318,747,772]" gridcol="0" gridrow="2" pageId="10" pageNumber="443">Size</th>
<td id="1A1B1C5D8751006FFEB1FD2D1C99B729" box="[349,450,747,772]" gridcol="1" gridrow="2" pageId="10" pageNumber="443">Small</td>
<td id="1A1B1C5D8751006FFE17FD2D1F22B729" box="[507,633,747,772]" gridcol="2" gridrow="2" pageId="10" pageNumber="443">Small</td>
<td id="1A1B1C5D8751006FFD74FD2D1E08B729" box="[664,851,747,772]" gridcol="3" gridrow="2" pageId="10" pageNumber="443">Small</td>
<td id="1A1B1C5D8751006FFC91FD2D1955B729" box="[893,1038,747,772]" gridcol="4" gridrow="2" pageId="10" pageNumber="443">Medium</td>
<td id="1A1B1C5D8751006FFBDCFD2D19F7B729" box="[1072,1196,747,772]" gridcol="5" gridrow="2" pageId="10" pageNumber="443">Medium</td>
<td id="1A1B1C5D8751006FFB32FD2D181DB729" box="[1246,1350,747,772]" gridcol="6" gridrow="2" pageId="10" pageNumber="443">Medium</td>
<td id="1A1B1C5D8751006FFA93FD2D18AAB729" box="[1407,1521,747,772]" gridcol="7" gridrow="2" pageId="10" pageNumber="443">Large</td>
<td id="1A1B1C5D8751006FF9FBFD2D1BFBB729" box="[1559,1696,747,772]" gridcol="8" gridrow="2" pageId="10" pageNumber="443">Medium</td>
<td id="1A1B1C5D8751006FF95BFD2D1A1FB729" box="[1719,1860,747,772]" gridcol="9" gridrow="2" pageId="10" pageNumber="443">Small</td>
<td id="1A1B1C5D8751006FF88FFD2D1AB1B729" box="[1891,2026,747,772]" gridcol="10" gridrow="2" pageId="10" pageNumber="443">Medium</td>
</tr>
<tr id="59CA75218751006FFF4EFCD31AB1B703" box="[162,2026,789,814]" gridrow="3" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFCD31C65B703" box="[162,318,789,814]" gridcol="0" gridrow="3" pageId="10" pageNumber="443">SVL adult</th>
<td id="1A1B1C5D8751006FFEB1FCD31C99B703" box="[349,450,789,814]" gridcol="1" gridrow="3" pageId="10" pageNumber="443">16.3–23.5</td>
<td id="1A1B1C5D8751006FFE17FCD31F22B703" box="[507,633,789,814]" gridcol="2" gridrow="3" pageId="10" pageNumber="443">13.51–19.88</td>
<td id="1A1B1C5D8751006FFD74FCD31E08B703" box="[664,851,789,814]" gridcol="3" gridrow="3" pageId="10" pageNumber="443">15.97–21.18</td>
<td id="1A1B1C5D8751006FFC91FCD31955B703" box="[893,1038,789,814]" gridcol="4" gridrow="3" pageId="10" pageNumber="443">22.8–28.4</td>
<td id="1A1B1C5D8751006FFBDCFCD319F7B703" box="[1072,1196,789,814]" gridcol="5" gridrow="3" pageId="10" pageNumber="443">20.6–29.0</td>
<td id="1A1B1C5D8751006FFB32FCD3181DB703" box="[1246,1350,789,814]" gridcol="6" gridrow="3" pageId="10" pageNumber="443">20.0–26.5</td>
<td id="1A1B1C5D8751006FFA93FCD318AAB703" box="[1407,1521,789,814]" gridcol="7" gridrow="3" pageId="10" pageNumber="443">22.1–33.2</td>
<td id="1A1B1C5D8751006FF9FBFCD31BFBB703" box="[1559,1696,789,814]" gridcol="8" gridrow="3" pageId="10" pageNumber="443">18.4–28.9</td>
<td id="1A1B1C5D8751006FF95BFCD31A1FB703" box="[1719,1860,789,814]" gridcol="9" gridrow="3" pageId="10" pageNumber="443">21.23–23.16</td>
<td id="1A1B1C5D8751006FF88FFCD31AB1B703" box="[1891,2026,789,814]" gridcol="10" gridrow="3" pageId="10" pageNumber="443">17.5–26.1</td>
</tr>
<tr id="59CA75218751006FFF4EFCFF1AB1B77F" box="[162,2026,825,850]" gridrow="4" pageId="10" pageNumber="443" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFCFF1C65B77F" box="[162,318,825,850]" gridcol="0" gridrow="4" pageId="10" pageNumber="443">males (range)</th>
</tr>
<tr id="59CA75218751006FFF4EFC9B1AB1B75B" box="[162,2026,861,886]" gridrow="5" pageId="10" pageNumber="443" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFC9B1C65B75B" box="[162,318,861,886]" gridcol="0" gridrow="5" pageId="10" pageNumber="443">in mm</th>
</tr>
<tr id="59CA75218751006FFF4EFC431AB1B7B3" box="[162,2026,901,926]" gridrow="6" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFC431C65B7B3" box="[162,318,901,926]" gridcol="0" gridrow="6" pageId="10" pageNumber="443">Condition of</th>
<td id="1A1B1C5D8751006FFEB1FC431C99B7B3" box="[349,450,901,926]" gridcol="1" gridrow="6" pageId="10" pageNumber="443">Usually</td>
<td id="1A1B1C5D8751006FFE17FC431F22B7B3" box="[507,633,901,926]" gridcol="2" gridrow="6" pageId="10" pageNumber="443">Present</td>
<td id="1A1B1C5D8751006FFD74FC431E08B7B3" box="[664,851,901,926]" gridcol="3" gridrow="6" pageId="10" pageNumber="443">Present</td>
<td id="1A1B1C5D8751006FFC91FC431955B7B3" box="[893,1038,901,926]" gridcol="4" gridrow="6" pageId="10" pageNumber="443">Present</td>
<td id="1A1B1C5D8751006FFBDCFC4319F7B7B3" box="[1072,1196,901,926]" gridcol="5" gridrow="6" pageId="10" pageNumber="443">Present</td>
<td id="1A1B1C5D8751006FFB32FC43181DB7B3" box="[1246,1350,901,926]" gridcol="6" gridrow="6" pageId="10" pageNumber="443">Present</td>
<td id="1A1B1C5D8751006FFA93FC4318AAB7B3" box="[1407,1521,901,926]" gridcol="7" gridrow="6" pageId="10" pageNumber="443">Present or</td>
<td id="1A1B1C5D8751006FF9FBFC431BFBB7B3" box="[1559,1696,901,926]" gridcol="8" gridrow="6" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FF95BFC431A1FB7B3" box="[1719,1860,901,926]" gridcol="9" gridrow="6" pageId="10" pageNumber="443">Present</td>
<td id="1A1B1C5D8751006FF88FFC431AB1B7B3" box="[1891,2026,901,926]" gridcol="10" gridrow="6" pageId="10" pageNumber="443">Present or</td>
</tr>
<tr id="59CA75218751006FFF4EFC6F1AB1B7EF" box="[162,2026,937,962]" gridrow="7" pageId="10" pageNumber="443" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFC6F1C65B7EF" box="[162,318,937,962]" gridcol="0" gridrow="7" pageId="10" pageNumber="443">interorbital bar</th>
<td id="1A1B1C5D8751006FFEB1FC6F1C99B7EF" box="[349,450,937,962]" gridcol="1" gridrow="7" pageId="10" pageNumber="443">absent</td>
<td id="1A1B1C5D8751006FFA93FC6F18AAB7EF" box="[1407,1521,937,962]" gridcol="7" gridrow="7" pageId="10" pageNumber="443">triangle</td>
<td id="1A1B1C5D8751006FF88FFC6F1AB1B7EF" box="[1891,2026,937,962]" gridcol="10" gridrow="7" pageId="10" pageNumber="443">not</td>
</tr>
<tr id="59CA75218751006FFF4EFC0B1AB1B7CB" box="[162,2026,973,998]" gridrow="8" pageId="10" pageNumber="443" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<td id="1A1B1C5D8751006FFA93FC0B18AAB7CB" box="[1407,1521,973,998]" gridcol="7" gridrow="8" pageId="10" pageNumber="443">present</td>
</tr>
<tr id="59CA75218751006FFF4EFC331AB1B023" box="[162,2026,1013,1038]" gridrow="9" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFC331C65B023" box="[162,318,1013,1038]" gridcol="0" gridrow="9" pageId="10" pageNumber="443">Tips of Finger</th>
<td id="1A1B1C5D8751006FFEB1FC331C99B023" box="[349,450,1013,1038]" gridcol="1" gridrow="9" pageId="10" pageNumber="443">Expanded</td>
<td id="1A1B1C5D8751006FFE17FC331F22B023" box="[507,633,1013,1038]" gridcol="2" gridrow="9" pageId="10" pageNumber="443">Expanded</td>
<td id="1A1B1C5D8751006FFD74FC331E08B023" box="[664,851,1013,1038]" gridcol="3" gridrow="9" pageId="10" pageNumber="443">Expanded and</td>
<td id="1A1B1C5D8751006FFC91FC331955B023" box="[893,1038,1013,1038]" gridcol="4" gridrow="9" pageId="10" pageNumber="443">Greatly</td>
<td id="1A1B1C5D8751006FFBDCFC3319F7B023" box="[1072,1196,1013,1038]" gridcol="5" gridrow="9" pageId="10" pageNumber="443">Slightly</td>
<td id="1A1B1C5D8751006FFB32FC33181DB023" box="[1246,1350,1013,1038]" gridcol="6" gridrow="9" pageId="10" pageNumber="443">Slightly</td>
<td id="1A1B1C5D8751006FFA93FC3318AAB023" box="[1407,1521,1013,1038]" gridcol="7" gridrow="9" pageId="10" pageNumber="443">Greatly</td>
<td id="1A1B1C5D8751006FF9FBFC331BFBB023" box="[1559,1696,1013,1038]" gridcol="8" gridrow="9" pageId="10" pageNumber="443">Expanded</td>
<td id="1A1B1C5D8751006FF95BFC331A1FB023" box="[1719,1860,1013,1038]" gridcol="9" gridrow="9" pageId="10" pageNumber="443">Expanded</td>
<td id="1A1B1C5D8751006FF88FFC331AB1B023" box="[1891,2026,1013,1038]" gridcol="10" gridrow="9" pageId="10" pageNumber="443">Slightly</td>
</tr>
<tr id="59CA75218751006FFF4EFBDF1AB1B01F" box="[162,2026,1049,1074]" gridrow="10" pageId="10" pageNumber="443" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFBDF1C65B01F" box="[162,318,1049,1074]" gridcol="0" gridrow="10" pageId="10" pageNumber="443">III and IV</th>
<td id="1A1B1C5D8751006FFEB1FBDF1C99B01F" box="[349,450,1049,1074]" gridcol="1" gridrow="10" pageId="10" pageNumber="443">and</td>
<td id="1A1B1C5D8751006FFE17FBDF1F22B01F" box="[507,633,1049,1074]" gridcol="2" gridrow="10" pageId="10" pageNumber="443">and</td>
<td id="1A1B1C5D8751006FFD74FBDF1E08B01F" box="[664,851,1049,1074]" gridcol="3" gridrow="10" pageId="10" pageNumber="443">truncated</td>
<td id="1A1B1C5D8751006FFC91FBDF1955B01F" box="[893,1038,1049,1074]" gridcol="4" gridrow="10" pageId="10" pageNumber="443">expanded and</td>
<td id="1A1B1C5D8751006FFBDCFBDF19F7B01F" box="[1072,1196,1049,1074]" gridcol="5" gridrow="10" pageId="10" pageNumber="443">expanded</td>
<td id="1A1B1C5D8751006FFB32FBDF181DB01F" box="[1246,1350,1049,1074]" gridcol="6" gridrow="10" pageId="10" pageNumber="443">expanded,</td>
<td id="1A1B1C5D8751006FFA93FBDF18AAB01F" box="[1407,1521,1049,1074]" gridcol="7" gridrow="10" pageId="10" pageNumber="443">expanded</td>
<td id="1A1B1C5D8751006FF9FBFBDF1BFBB01F" box="[1559,1696,1049,1074]" gridcol="8" gridrow="10" pageId="10" pageNumber="443">and truncated</td>
<td id="1A1B1C5D8751006FF88FFBDF1AB1B01F" box="[1891,2026,1049,1074]" gridcol="10" gridrow="10" pageId="10" pageNumber="443">expanded</td>
</tr>
<tr id="59CA75218751006FFF4EFBFB1AB1B07B" box="[162,2026,1085,1110]" gridrow="11" pageId="10" pageNumber="443" rowspan-0="1" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<td id="1A1B1C5D8751006FFEB1FBFB1C99B07B" box="[349,450,1085,1110]" gridcol="1" gridrow="11" pageId="10" pageNumber="443">truncated</td>
<td id="1A1B1C5D8751006FFE17FBFB1F22B07B" box="[507,633,1085,1110]" gridcol="2" gridrow="11" pageId="10" pageNumber="443">truncated</td>
<td id="1A1B1C5D8751006FFC91FBFB1955B07B" box="[893,1038,1085,1110]" gridcol="4" gridrow="11" pageId="10" pageNumber="443">truncated</td>
<td id="1A1B1C5D8751006FFB32FBFB181DB07B" box="[1246,1350,1085,1110]" gridcol="6" gridrow="11" pageId="10" pageNumber="443">truncated</td>
<td id="1A1B1C5D8751006FFA93FBFB18AAB07B" box="[1407,1521,1085,1110]" gridcol="7" gridrow="11" pageId="10" pageNumber="443">and</td>
</tr>
<tr id="59CA75218751006FFF4EFBA71AB1B057" box="[162,2026,1121,1146]" gridrow="12" pageId="10" pageNumber="443" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<td id="1A1B1C5D8751006FFA93FBA718AAB057" box="[1407,1521,1121,1146]" gridcol="7" gridrow="12" pageId="10" pageNumber="443">truncated</td>
</tr>
<tr id="59CA75218751006FFF4EFB4C1AB1B08E" box="[162,2026,1162,1187]" gridrow="13" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFB4C1C65B08E" box="[162,318,1162,1187]" gridcol="0" gridrow="13" pageId="10" pageNumber="443">Subarticular</th>
<td id="1A1B1C5D8751006FFEB1FB4C1C99B08E" box="[349,450,1162,1187]" gridcol="1" gridrow="13" pageId="10" pageNumber="443">Rounded</td>
<td id="1A1B1C5D8751006FFE17FB4C1F22B08E" box="[507,633,1162,1187]" gridcol="2" gridrow="13" pageId="10" pageNumber="443">Conical</td>
<td id="1A1B1C5D8751006FFD74FB4C1E08B08E" box="[664,851,1162,1187]" gridcol="3" gridrow="13" pageId="10" pageNumber="443">Rounded</td>
<td id="1A1B1C5D8751006FFC91FB4C1955B08E" box="[893,1038,1162,1187]" gridcol="4" gridrow="13" pageId="10" pageNumber="443">Conical and</td>
<td id="1A1B1C5D8751006FFBDCFB4C19F7B08E" box="[1072,1196,1162,1187]" gridcol="5" gridrow="13" pageId="10" pageNumber="443">Conical</td>
<td id="1A1B1C5D8751006FFB32FB4C181DB08E" box="[1246,1350,1162,1187]" gridcol="6" gridrow="13" pageId="10" pageNumber="443">Conical</td>
<td id="1A1B1C5D8751006FFA93FB4C18AAB08E" box="[1407,1521,1162,1187]" gridcol="7" gridrow="13" pageId="10" pageNumber="443">Rounded</td>
<td id="1A1B1C5D8751006FF9FBFB4C1BFBB08E" box="[1559,1696,1162,1187]" gridcol="8" gridrow="13" pageId="10" pageNumber="443">Conical</td>
<td id="1A1B1C5D8751006FF95BFB4C1A1FB08E" box="[1719,1860,1162,1187]" gridcol="9" gridrow="13" pageId="10" pageNumber="443">Slightly</td>
<td id="1A1B1C5D8751006FF88FFB4C1AB1B08E" box="[1891,2026,1162,1187]" gridcol="10" gridrow="13" pageId="10" pageNumber="443">Conical</td>
</tr>
<tr id="59CA75218751006FFF4EFB681AB1B0EA" box="[162,2026,1198,1223]" gridrow="14" pageId="10" pageNumber="443" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="1A1B1C5D8751006FFF4EFB681C65B0EA" box="[162,318,1198,1223]" gridcol="0" gridrow="14" pageId="10" pageNumber="443">tubercles</th>
<td id="1A1B1C5D8751006FFC91FB681955B0EA" box="[893,1038,1198,1223]" gridcol="4" gridrow="14" pageId="10" pageNumber="443">rounded</td>
<td id="1A1B1C5D8751006FF95BFB681A1FB0EA" box="[1719,1860,1198,1223]" gridcol="9" gridrow="14" pageId="10" pageNumber="443">conical</td>
</tr>
<tr id="59CA75218751006FFF4EFB101AB1B0C2" box="[162,2026,1238,1263]" gridrow="15" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFB101C65B0C2" box="[162,318,1238,1263]" gridcol="0" gridrow="15" pageId="10" pageNumber="443">Dorsal skin</th>
<td id="1A1B1C5D8751006FFEB1FB101C99B0C2" box="[349,450,1238,1263]" gridcol="1" gridrow="15" pageId="10" pageNumber="443">Coarsely</td>
<td id="1A1B1C5D8751006FFE17FB101F22B0C2" box="[507,633,1238,1263]" gridcol="2" gridrow="15" pageId="10" pageNumber="443">Granular</td>
<td id="1A1B1C5D8751006FFD74FB101E08B0C2" box="[664,851,1238,1263]" gridcol="3" gridrow="15" pageId="10" pageNumber="443">Weakly granular</td>
<td id="1A1B1C5D8751006FFC91FB101955B0C2" box="[893,1038,1238,1263]" gridcol="4" gridrow="15" pageId="10" pageNumber="443">Shagreened to</td>
<td id="1A1B1C5D8751006FFBDCFB1019F7B0C2" box="[1072,1196,1238,1263]" gridcol="5" gridrow="15" pageId="10" pageNumber="443">Smooth to</td>
<td id="1A1B1C5D8751006FFB32FB10181DB0C2" box="[1246,1350,1238,1263]" gridcol="6" gridrow="15" pageId="10" pageNumber="443">Pustular</td>
<td id="1A1B1C5D8751006FFA93FB1018AAB0C2" box="[1407,1521,1238,1263]" gridcol="7" gridrow="15" pageId="10" pageNumber="443">Pustular</td>
<td id="1A1B1C5D8751006FF9FBFB101BFBB0C2" box="[1559,1696,1238,1263]" gridcol="8" gridrow="15" pageId="10" pageNumber="443">Smooth</td>
<td id="1A1B1C5D8751006FF95BFB101A1FB0C2" box="[1719,1860,1238,1263]" gridcol="9" gridrow="15" pageId="10" pageNumber="443">Granular</td>
<td id="1A1B1C5D8751006FF88FFB101AB1B0C2" box="[1891,2026,1238,1263]" gridcol="10" gridrow="15" pageId="10" pageNumber="443">Pustular</td>
</tr>
<tr id="59CA75218751006FFF4EFB3C1AB1B13E" box="[162,2026,1274,1299]" gridrow="16" pageId="10" pageNumber="443" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFB3C1C65B13E" box="[162,318,1274,1299]" gridcol="0" gridrow="16" pageId="10" pageNumber="443">texture</th>
<td id="1A1B1C5D8751006FFEB1FB3C1C99B13E" box="[349,450,1274,1299]" gridcol="1" gridrow="16" pageId="10" pageNumber="443">granular</td>
<td id="1A1B1C5D8751006FFC91FB3C1955B13E" box="[893,1038,1274,1299]" gridcol="4" gridrow="16" pageId="10" pageNumber="443">granular</td>
<td id="1A1B1C5D8751006FFBDCFB3C19F7B13E" box="[1072,1196,1274,1299]" gridcol="5" gridrow="16" pageId="10" pageNumber="443">moderately</td>
<td id="1A1B1C5D8751006FF9FBFB3C1BFBB13E" box="[1559,1696,1274,1299]" gridcol="8" gridrow="16" pageId="10" pageNumber="443">to weakly</td>
</tr>
<tr id="59CA75218751006FFF4EFAD81AB1B11A" box="[162,2026,1310,1335]" gridrow="17" pageId="10" pageNumber="443" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<td id="1A1B1C5D8751006FFBDCFAD819F7B11A" box="[1072,1196,1310,1335]" gridcol="5" gridrow="17" pageId="10" pageNumber="443">pustular</td>
<td id="1A1B1C5D8751006FF9FBFAD81BFBB11A" box="[1559,1696,1310,1335]" gridcol="8" gridrow="17" pageId="10" pageNumber="443">pustular</td>
</tr>
<tr id="59CA75218751006FFF4EFA801AB1B172" box="[162,2026,1350,1375]" gridrow="18" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFA801C65B172" box="[162,318,1350,1375]" gridcol="0" gridrow="18" pageId="10" pageNumber="443">Ventral skin</th>
<td id="1A1B1C5D8751006FFEB1FA801C99B172" box="[349,450,1350,1375]" gridcol="1" gridrow="18" pageId="10" pageNumber="443">Smooth</td>
<td id="1A1B1C5D8751006FFE17FA801F22B172" box="[507,633,1350,1375]" gridcol="2" gridrow="18" pageId="10" pageNumber="443">Smooth</td>
<td id="1A1B1C5D8751006FFD74FA801E08B172" box="[664,851,1350,1375]" gridcol="3" gridrow="18" pageId="10" pageNumber="443">Areolate</td>
<td id="1A1B1C5D8751006FFC91FA801955B172" box="[893,1038,1350,1375]" gridcol="4" gridrow="18" pageId="10" pageNumber="443">Weakly to</td>
<td id="1A1B1C5D8751006FFBDCFA8019F7B172" box="[1072,1196,1350,1375]" gridcol="5" gridrow="18" pageId="10" pageNumber="443">Areolate</td>
<td id="1A1B1C5D8751006FFB32FA80181DB172" box="[1246,1350,1350,1375]" gridcol="6" gridrow="18" pageId="10" pageNumber="443">Smooth</td>
<td id="1A1B1C5D8751006FFA93FA8018AAB172" box="[1407,1521,1350,1375]" gridcol="7" gridrow="18" pageId="10" pageNumber="443">Smooth</td>
<td id="1A1B1C5D8751006FF9FBFA801BFBB172" box="[1559,1696,1350,1375]" gridcol="8" gridrow="18" pageId="10" pageNumber="443">Smooth</td>
<td id="1A1B1C5D8751006FF95BFA801A1FB172" box="[1719,1860,1350,1375]" gridcol="9" gridrow="18" pageId="10" pageNumber="443">Smooth</td>
<td id="1A1B1C5D8751006FF88FFA801AB1B172" box="[1891,2026,1350,1375]" gridcol="10" gridrow="18" pageId="10" pageNumber="443">Areolate</td>
</tr>
<tr id="59CA75218751006FFF4EFAAC1AB1B1AE" box="[162,2026,1386,1411]" gridrow="19" pageId="10" pageNumber="443" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFAAC1C65B1AE" box="[162,318,1386,1411]" gridcol="0" gridrow="19" pageId="10" pageNumber="443">texture</th>
<td id="1A1B1C5D8751006FFC91FAAC1955B1AE" box="[893,1038,1386,1411]" gridcol="4" gridrow="19" pageId="10" pageNumber="443">moderately</td>
</tr>
<tr id="59CA75218751006FFF4EFA481AB1B18A" box="[162,2026,1422,1447]" gridrow="20" pageId="10" pageNumber="443" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td id="1A1B1C5D8751006FFC91FA481955B18A" box="[893,1038,1422,1447]" gridcol="4" gridrow="20" pageId="10" pageNumber="443">areolate</td>
</tr>
<tr id="59CA75218751006FFF4EFA701AB1B1E2" box="[162,2026,1462,1487]" gridrow="21" pageId="10" pageNumber="443">
<th id="1A1B1C5D8751006FFF4EFA701C65B1E2" box="[162,318,1462,1487]" gridcol="0" gridrow="21" pageId="10" pageNumber="443">Presence or</th>
<td id="1A1B1C5D8751006FFEB1FA701C99B1E2" box="[349,450,1462,1487]" gridcol="1" gridrow="21" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FFE17FA701F22B1E2" box="[507,633,1462,1487]" gridcol="2" gridrow="21" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FFD74FA701E08B1E2" box="[664,851,1462,1487]" gridcol="3" gridrow="21" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FFC91FA701955B1E2" box="[893,1038,1462,1487]" gridcol="4" gridrow="21" pageId="10" pageNumber="443">Present or not</td>
<td id="1A1B1C5D8751006FFBDCFA7019F7B1E2" box="[1072,1196,1462,1487]" gridcol="5" gridrow="21" pageId="10" pageNumber="443">Present</td>
<td id="1A1B1C5D8751006FFB32FA70181DB1E2" box="[1246,1350,1462,1487]" gridcol="6" gridrow="21" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FFA93FA7018AAB1E2" box="[1407,1521,1462,1487]" gridcol="7" gridrow="21" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FF9FBFA701BFBB1E2" box="[1559,1696,1462,1487]" gridcol="8" gridrow="21" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FF95BFA701A1FB1E2" box="[1719,1860,1462,1487]" gridcol="9" gridrow="21" pageId="10" pageNumber="443">Absent</td>
<td id="1A1B1C5D8751006FF88FFA701AB1B1E2" box="[1891,2026,1462,1487]" gridcol="10" gridrow="21" pageId="10" pageNumber="443">Present</td>
</tr>
<tr id="59CA75218751006FFF4EFA1C1AB1B1DE" box="[162,2026,1498,1523]" gridrow="22" pageId="10" pageNumber="443" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFA1C1C65B1DE" box="[162,318,1498,1523]" gridcol="0" gridrow="22" pageId="10" pageNumber="443">absence of</th>
</tr>
<tr id="59CA75218751006FFF4EFA381AB1B23A" box="[162,2026,1534,1559]" gridrow="23" pageId="10" pageNumber="443" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EFA381C65B23A" box="[162,318,1534,1559]" gridcol="0" gridrow="23" pageId="10" pageNumber="443">lumbo-inguinal</th>
</tr>
<tr id="59CA75218751006FFF4EF9E41AB1B216" box="[162,2026,1570,1595]" gridrow="24" pageId="10" pageNumber="443" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1A1B1C5D8751006FFF4EF9E41C65B216" box="[162,318,1570,1595]" gridcol="0" gridrow="24" pageId="10" pageNumber="443">gland</th>
</tr>
</table>
</paragraph>
<caption id="B38527EB8750FF9BFF74FE081B16B621" ID-Table-UUID="B38527EB8750FF9BFF74FE081B16B621" httpUri="http://table.plazi.org/id/B38527EB8750FF9BFF74FE081B16B621" pageId="11" pageNumber="444" startId="11.[152,244,462,488]" subCaptionStartIDs="11.[592,743,498,524]" subCaptionStarts="Abbr" targetBox="[162,2022,539,1680]" targetIsTable="true" targetPageId="11" targetType="table">
<paragraph id="E74577638750FF9BFF74FE081B16B621" pageId="11" pageNumber="444">
<emphasis id="D58EAB718750FF9BFF74FE081C54B5C5" bold="true" box="[152,271,462,488]" pageId="11" pageNumber="444">TABLE 4.</emphasis>
Morphometric data (in mm) of the type series of 
<taxonomicName id="20FA0CE08750FF9BFCC9FE081961B5C5" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[805,1082,462,488]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="444" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718750FF9BFCC9FE081961B5C5" box="[805,1082,462,488]" italics="true" pageId="11" pageNumber="444">Eleutherodactylus coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718750FF9BFBADFE0819CCB5C5" bold="true" box="[1089,1175,462,488]" pageId="11" pageNumber="444">
<taxonomicNameLabel id="CEBD160A8750FF9BFBADFE0819CCB5C5" box="[1089,1175,462,488]" pageId="11" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and closely related species. 
<emphasis id="D58EAB718750FF9BFA25FE081880B5C5" box="[1481,1499,462,488]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= sample size. For each character, the mean ± SD and range (in parentheses) are presented. Abbreviations for morphological characters are explained in the Materials and methods section.
</paragraph>
</caption>
<paragraph id="E74577638750FF9BFE58FDDD1AFFB2BD" pageId="11" pageNumber="444">
<table id="95FA85C38750006FFF4EFDDD1ABDB2BD" box="[162,2022,539,1680]" colsContinueIn="12.[151,2011,981,1252]" gridcols="8" gridrows="22" pageId="11" pageNumber="444">
<tr id="59CA75218750006FFF4EFDDD1ABDB619" box="[162,2022,539,564]" gridrow="0" pageId="11" pageNumber="444" rowspan-0="1">
<th id="1A1B1C5D8750006FFEE0FDDD1E4DB619" box="[268,790,539,564]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="11" pageNumber="444">
<taxonomicName id="20FA0CE08750FF9BFE58FDDD1F42B619" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[436,537,539,564]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="444" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718750FF9BFE58FDDD1F42B619" box="[436,537,539,564]" italics="true" pageId="11" pageNumber="444">E. coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718750FF9BFDF3FDDD1F2BB619" bold="true" box="[543,624,539,564]" pageId="11" pageNumber="444">
<taxonomicNameLabel id="CEBD160A8750FF9BFDF3FDDD1F2BB619" box="[543,624,539,564]" pageId="11" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="1A1B1C5D8750006FFCD6FDDD181CB619" box="[826,1351,539,564]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="11" pageNumber="444">
<taxonomicName id="20FA0CE08750FF9BFC0EFDDD19C6B619" box="[994,1181,539,564]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="444" phylum="Chordata" rank="species" species="cystignathoides">
<emphasis id="D58EAB718750FF9BFC0EFDDD19C6B619" box="[994,1181,539,564]" italics="true" pageId="11" pageNumber="444">E. cystignathoides</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8750006FFA89FDDD1A41B619" box="[1381,1818,539,564]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="11" pageNumber="444">
<taxonomicName id="20FA0CE08750FF9BFA21FDDD1B69B619" box="[1485,1586,539,564]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="444" phylum="Chordata" rank="species" species="dennisi">
<emphasis id="D58EAB718750FF9BFA21FDDD1B69B619" box="[1485,1586,539,564]" italics="true" pageId="11" pageNumber="444">E. dennisi</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8750006FF8D9FDDD1ABDB619" box="[1845,2022,539,564]" gridcol="7" gridrow="0" pageId="11" pageNumber="444">
<emphasis id="D58EAB718750FF9BF8D9FDDD1A1FB619" box="[1845,1860,539,564]" italics="true" pageId="11" pageNumber="444">E</emphasis>
. sp. Ahuacatlán
</th>
</tr>
<tr id="59CA75218750006FFF4EFD851ABDB671" box="[162,2022,579,604]" gridrow="1" pageId="11" pageNumber="444" rowspan-0="1">
<td id="1A1B1C5D8750006FFEE0FD851F58B671" box="[268,515,579,604]" gridcol="1" gridrow="1" pageId="11" pageNumber="444">
Males (
<emphasis id="D58EAB718750FF9BFE61FD851CC5B671" box="[397,414,579,604]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= 8)
</td>
<td id="1A1B1C5D8750006FFDCAFD851E4DB671" box="[550,790,579,604]" gridcol="2" gridrow="1" pageId="11" pageNumber="444">
Females (
<emphasis id="D58EAB718750FF9BFD43FD851F9BB671" box="[687,704,579,604]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= 2)
</td>
<td id="1A1B1C5D8750006FFCD6FD851971B671" box="[826,1066,579,604]" gridcol="3" gridrow="1" pageId="11" pageNumber="444">
Males (
<emphasis id="D58EAB718750FF9BFC5EFD851E98B671" box="[946,963,579,604]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= 10)
</td>
<td id="1A1B1C5D8750006FFBA7FD85181CB671" box="[1099,1351,579,604]" gridcol="4" gridrow="1" pageId="11" pageNumber="444">
Females (
<emphasis id="D58EAB718750FF9BFB36FD8519B0B671" box="[1242,1259,579,604]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= 2)
</td>
<td id="1A1B1C5D8750006FFA89FD8518B3B671" box="[1381,1512,579,604]" gridcol="5" gridrow="1" pageId="11" pageNumber="444">
Male (
<emphasis id="D58EAB718750FF9BFA4BFD8518E3B671" box="[1447,1464,579,604]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= 1)
</td>
<td id="1A1B1C5D8750006FF9E8FD851A41B671" box="[1540,1818,579,604]" gridcol="6" gridrow="1" pageId="11" pageNumber="444">
Females (
<emphasis id="D58EAB718750FF9BF94CFD851BEAB671" box="[1696,1713,579,604]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= 2)
</td>
<td id="1A1B1C5D8750006FF8D9FD851ABDB671" box="[1845,2022,579,604]" gridcol="7" gridrow="1" pageId="11" pageNumber="444">
Female (
<emphasis id="D58EAB718750FF9BF876FD851AF0B671" box="[1946,1963,579,604]" italics="true" pageId="11" pageNumber="444">N</emphasis>
= 1)
</td>
</tr>
<tr id="59CA75218750006FFF4EFDAD1ABDB685" box="[162,2022,619,680]" gridrow="2" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFDAD1DBDB685" box="[162,230,619,680]" gridcol="0" gridrow="2" pageId="11" pageNumber="444">SVL</th>
<td id="1A1B1C5D8750006FFEE0FDAD1F58B685" box="[268,515,619,680]" gridcol="1" gridrow="2" pageId="11" pageNumber="444">16.80 ± 1.95 (13.51– 19.88)</td>
<td id="1A1B1C5D8750006FFDCAFDAD1E4DB685" box="[550,790,619,680]" gridcol="2" gridrow="2" pageId="11" pageNumber="444">16.64 ± 0.68 (16.16– 17.12)</td>
<td id="1A1B1C5D8750006FFCD6FDAD1971B685" box="[826,1066,619,680]" gridcol="3" gridrow="2" pageId="11" pageNumber="444">18.60 ± 1.33 (15.97– 21.18)</td>
<td id="1A1B1C5D8750006FFBA7FDAD181CB685" box="[1099,1351,619,680]" gridcol="4" gridrow="2" pageId="11" pageNumber="444">17.92 ± 8.06 (12.22– 23.62)</td>
<td id="1A1B1C5D8750006FFA89FDAD18B3B685" box="[1381,1512,619,680]" gridcol="5" gridrow="2" pageId="11" pageNumber="444">24.64</td>
<td id="1A1B1C5D8750006FF9E8FDAD1A41B685" box="[1540,1818,619,680]" gridcol="6" gridrow="2" pageId="11" pageNumber="444">33.23 ± 2.26 (31.36–34.83)</td>
<td id="1A1B1C5D8750006FF8D9FDAD1ABDB685" box="[1845,2022,619,680]" gridcol="7" gridrow="2" pageId="11" pageNumber="444">23.07</td>
</tr>
<tr id="59CA75218750006FFF4EFD781ABDB6FA" box="[162,2022,702,727]" gridrow="3" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFD781DBDB6FA" box="[162,230,702,727]" gridcol="0" gridrow="3" pageId="11" pageNumber="444">HL</th>
<td id="1A1B1C5D8750006FFEE0FD781F58B6FA" box="[268,515,702,727]" gridcol="1" gridrow="3" pageId="11" pageNumber="444">6.96 ± 0.68 (5.91–7.94)</td>
<td id="1A1B1C5D8750006FFDCAFD781E4DB6FA" box="[550,790,702,727]" gridcol="2" gridrow="3" pageId="11" pageNumber="444">6.26 ± 0.27 (6.07–6.45)</td>
<td id="1A1B1C5D8750006FFCD6FD781971B6FA" box="[826,1066,702,727]" gridcol="3" gridrow="3" pageId="11" pageNumber="444">6.85 ± 0.45 (6.25–7.61)</td>
<td id="1A1B1C5D8750006FFBA7FD78181CB6FA" box="[1099,1351,702,727]" gridcol="4" gridrow="3" pageId="11" pageNumber="444">6.77 ± 2.56 (4.96–8.58)</td>
<td id="1A1B1C5D8750006FFA89FD7818B3B6FA" box="[1381,1512,702,727]" gridcol="5" gridrow="3" pageId="11" pageNumber="444">9.82</td>
<td id="1A1B1C5D8750006FF9E8FD781A41B6FA" box="[1540,1818,702,727]" gridcol="6" gridrow="3" pageId="11" pageNumber="444">13.53 ± 0.21 (13.38–13.67)</td>
<td id="1A1B1C5D8750006FF8D9FD781ABDB6FA" box="[1845,2022,702,727]" gridcol="7" gridrow="3" pageId="11" pageNumber="444">8.9</td>
</tr>
<tr id="59CA75218750006FFF4EFD351ABDB721" box="[162,2022,755,780]" gridrow="4" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFD351DBDB721" box="[162,230,755,780]" gridcol="0" gridrow="4" pageId="11" pageNumber="444">HW</th>
<td id="1A1B1C5D8750006FFEE0FD351F58B721" box="[268,515,755,780]" gridcol="1" gridrow="4" pageId="11" pageNumber="444">6.48 ± 0.73 (5.54–7.6)</td>
<td id="1A1B1C5D8750006FFDCAFD351E4DB721" box="[550,790,755,780]" gridcol="2" gridrow="4" pageId="11" pageNumber="444">5.93 ± 0.04 (5.9–5.95)</td>
<td id="1A1B1C5D8750006FFCD6FD351971B721" box="[826,1066,755,780]" gridcol="3" gridrow="4" pageId="11" pageNumber="444">6.60 ± 0.37 (5.83–6.98)</td>
<td id="1A1B1C5D8750006FFBA7FD35181CB721" box="[1099,1351,755,780]" gridcol="4" gridrow="4" pageId="11" pageNumber="444">6.19 ± 2.72 (4.27–8.11)</td>
<td id="1A1B1C5D8750006FFA89FD3518B3B721" box="[1381,1512,755,780]" gridcol="5" gridrow="4" pageId="11" pageNumber="444">9.08</td>
<td id="1A1B1C5D8750006FF9E8FD351A41B721" box="[1540,1818,755,780]" gridcol="6" gridrow="4" pageId="11" pageNumber="444">12.00 ± 0.19 (11.86–12.13)</td>
<td id="1A1B1C5D8750006FF8D9FD351ABDB721" box="[1845,2022,755,780]" gridcol="7" gridrow="4" pageId="11" pageNumber="444">8.73</td>
</tr>
<tr id="59CA75218750006FFF4EFCEE1ABDB76C" box="[162,2022,808,833]" gridrow="5" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFCEE1DBDB76C" box="[162,230,808,833]" gridcol="0" gridrow="5" pageId="11" pageNumber="444">EW</th>
<td id="1A1B1C5D8750006FFEE0FCEE1F58B76C" box="[268,515,808,833]" gridcol="1" gridrow="5" pageId="11" pageNumber="444">1.19 ± 0.18 (0.91–1.45)</td>
<td id="1A1B1C5D8750006FFDCAFCEE1E4DB76C" box="[550,790,808,833]" gridcol="2" gridrow="5" pageId="11" pageNumber="444">1.21 ± 0.14 (1.11–1.31)</td>
<td id="1A1B1C5D8750006FFCD6FCEE1971B76C" box="[826,1066,808,833]" gridcol="3" gridrow="5" pageId="11" pageNumber="444">1.35 ± 0.18 (0.98–1.61)</td>
<td id="1A1B1C5D8750006FFBA7FCEE181CB76C" box="[1099,1351,808,833]" gridcol="4" gridrow="5" pageId="11" pageNumber="444">1.36 ± 0.61 (0.93–1.79)</td>
<td id="1A1B1C5D8750006FFA89FCEE18B3B76C" box="[1381,1512,808,833]" gridcol="5" gridrow="5" pageId="11" pageNumber="444">1.76</td>
<td id="1A1B1C5D8750006FF9E8FCEE1A41B76C" box="[1540,1818,808,833]" gridcol="6" gridrow="5" pageId="11" pageNumber="444">2.56 ± 0.04 (2.53–2.58)</td>
<td id="1A1B1C5D8750006FF8D9FCEE1ABDB76C" box="[1845,2022,808,833]" gridcol="7" gridrow="5" pageId="11" pageNumber="444">9.28</td>
</tr>
<tr id="59CA75218750006FFF4EFC9B1ABDB75B" box="[162,2022,861,886]" gridrow="6" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFC9B1DBDB75B" box="[162,230,861,886]" gridcol="0" gridrow="6" pageId="11" pageNumber="444">END</th>
<td id="1A1B1C5D8750006FFEE0FC9B1F58B75B" box="[268,515,861,886]" gridcol="1" gridrow="6" pageId="11" pageNumber="444">1.99 ± 0.25 (1.57–2.13)</td>
<td id="1A1B1C5D8750006FFDCAFC9B1E4DB75B" box="[550,790,861,886]" gridcol="2" gridrow="6" pageId="11" pageNumber="444">1.92 ± 0.18 (1.79–2.05)</td>
<td id="1A1B1C5D8750006FFCD6FC9B1971B75B" box="[826,1066,861,886]" gridcol="3" gridrow="6" pageId="11" pageNumber="444">2.13 ± 0.19 (1.79–2.46)</td>
<td id="1A1B1C5D8750006FFBA7FC9B181CB75B" box="[1099,1351,861,886]" gridcol="4" gridrow="6" pageId="11" pageNumber="444">2.03 ± 0.8 (1.46–2.59)</td>
<td id="1A1B1C5D8750006FFA89FC9B18B3B75B" box="[1381,1512,861,886]" gridcol="5" gridrow="6" pageId="11" pageNumber="444">2.94</td>
<td id="1A1B1C5D8750006FF9E8FC9B1A41B75B" box="[1540,1818,861,886]" gridcol="6" gridrow="6" pageId="11" pageNumber="444">4.34 ± 0.43 (4.03–4.64)</td>
<td id="1A1B1C5D8750006FF8D9FC9B1ABDB75B" box="[1845,2022,861,886]" gridcol="7" gridrow="6" pageId="11" pageNumber="444">2.78</td>
</tr>
<tr id="59CA75218750006FFF4EFC541ABDB786" box="[162,2022,914,939]" gridrow="7" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFC541DBDB786" box="[162,230,914,939]" gridcol="0" gridrow="7" pageId="11" pageNumber="444">IOD</th>
<td id="1A1B1C5D8750006FFEE0FC541F58B786" box="[268,515,914,939]" gridcol="1" gridrow="7" pageId="11" pageNumber="444">2.46 ± 0.31 (2.06–2.91)</td>
<td id="1A1B1C5D8750006FFDCAFC541E4DB786" box="[550,790,914,939]" gridcol="2" gridrow="7" pageId="11" pageNumber="444">2.10 ± 0.06 (2.06–2.14)</td>
<td id="1A1B1C5D8750006FFCD6FC541971B786" box="[826,1066,914,939]" gridcol="3" gridrow="7" pageId="11" pageNumber="444">3.79 ± 0.34 (3.1–4.36)</td>
<td id="1A1B1C5D8750006FFBA7FC54181CB786" box="[1099,1351,914,939]" gridcol="4" gridrow="7" pageId="11" pageNumber="444">3.71 ± 1.4 (2.72–4.7)</td>
<td id="1A1B1C5D8750006FFA89FC5418B3B786" box="[1381,1512,914,939]" gridcol="5" gridrow="7" pageId="11" pageNumber="444">2.99</td>
<td id="1A1B1C5D8750006FF9E8FC541A41B786" box="[1540,1818,914,939]" gridcol="6" gridrow="7" pageId="11" pageNumber="444">4.63 ± 0.27 (4.44–4.82)</td>
<td id="1A1B1C5D8750006FF8D9FC541ABDB786" box="[1845,2022,914,939]" gridcol="7" gridrow="7" pageId="11" pageNumber="444">3.09</td>
</tr>
<tr id="59CA75218750006FFF4EFC011ABDB7CD" box="[162,2022,967,992]" gridrow="8" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFC011DBDB7CD" box="[162,230,967,992]" gridcol="0" gridrow="8" pageId="11" pageNumber="444">TD</th>
<td id="1A1B1C5D8750006FFEE0FC011F58B7CD" box="[268,515,967,992]" gridcol="1" gridrow="8" pageId="11" pageNumber="444">1.36 ± 0.32 (1.04–1.64)</td>
<td id="1A1B1C5D8750006FFDCAFC011E4DB7CD" box="[550,790,967,992]" gridcol="2" gridrow="8" pageId="11" pageNumber="444">1.10 ± 0.21 (0.95–1.24)</td>
<td id="1A1B1C5D8750006FFCD6FC011971B7CD" box="[826,1066,967,992]" gridcol="3" gridrow="8" pageId="11" pageNumber="444">1.39 ± 0.17 (1.07–1.62)</td>
<td id="1A1B1C5D8750006FFBA7FC01181CB7CD" box="[1099,1351,967,992]" gridcol="4" gridrow="8" pageId="11" pageNumber="444">1.01 ± 0.48 (0.67–1.35)</td>
<td id="1A1B1C5D8750006FFA89FC0118B3B7CD" box="[1381,1512,967,992]" gridcol="5" gridrow="8" pageId="11" pageNumber="444">1.56</td>
<td id="1A1B1C5D8750006FF9E8FC011A41B7CD" box="[1540,1818,967,992]" gridcol="6" gridrow="8" pageId="11" pageNumber="444">2.43 ± 0.23 (2.26–2.59)</td>
<td id="1A1B1C5D8750006FF8D9FC011ABDB7CD" box="[1845,2022,967,992]" gridcol="7" gridrow="8" pageId="11" pageNumber="444">1.51</td>
</tr>
<tr id="59CA75218750006FFF4EFC3A1ABDB038" box="[162,2022,1020,1045]" gridrow="9" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFC3A1DBDB038" box="[162,230,1020,1045]" gridcol="0" gridrow="9" pageId="11" pageNumber="444">EL</th>
<td id="1A1B1C5D8750006FFEE0FC3A1F58B038" box="[268,515,1020,1045]" gridcol="1" gridrow="9" pageId="11" pageNumber="444">2.24 ± 0.21 (2.02–2.5)</td>
<td id="1A1B1C5D8750006FFDCAFC3A1E4DB038" box="[550,790,1020,1045]" gridcol="2" gridrow="9" pageId="11" pageNumber="444">2.22 ± 0.24 (2.05–2.39)</td>
<td id="1A1B1C5D8750006FFCD6FC3A1971B038" box="[826,1066,1020,1045]" gridcol="3" gridrow="9" pageId="11" pageNumber="444">2.41 ± 0.15 (2.22–2.64)</td>
<td id="1A1B1C5D8750006FFBA7FC3A181CB038" box="[1099,1351,1020,1045]" gridcol="4" gridrow="9" pageId="11" pageNumber="444">2.35 ± 0.8 (1.78 2.91)</td>
<td id="1A1B1C5D8750006FFA89FC3A18B3B038" box="[1381,1512,1020,1045]" gridcol="5" gridrow="9" pageId="11" pageNumber="444">3.16</td>
<td id="1A1B1C5D8750006FF9E8FC3A1A41B038" box="[1540,1818,1020,1045]" gridcol="6" gridrow="9" pageId="11" pageNumber="444">4.27 ± 0.06 (4.23–4.31)</td>
<td id="1A1B1C5D8750006FF8D9FC3A1ABDB038" box="[1845,2022,1020,1045]" gridcol="7" gridrow="9" pageId="11" pageNumber="444">2.72</td>
</tr>
<tr id="59CA75218750006FFF4EFBF71ABDB067" box="[162,2022,1073,1098]" gridrow="10" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFBF71DBDB067" box="[162,230,1073,1098]" gridcol="0" gridrow="10" pageId="11" pageNumber="444">IND</th>
<td id="1A1B1C5D8750006FFEE0FBF71F58B067" box="[268,515,1073,1098]" gridcol="1" gridrow="10" pageId="11" pageNumber="444">1.83 ± 0.18 (1.63–2.08)</td>
<td id="1A1B1C5D8750006FFDCAFBF71E4DB067" box="[550,790,1073,1098]" gridcol="2" gridrow="10" pageId="11" pageNumber="444">1.69 ± 0.13 (1.6–1.78)</td>
<td id="1A1B1C5D8750006FFCD6FBF71971B067" box="[826,1066,1073,1098]" gridcol="3" gridrow="10" pageId="11" pageNumber="444">1.84 ± 0.07 (1.71–1.97)</td>
<td id="1A1B1C5D8750006FFBA7FBF7181CB067" box="[1099,1351,1073,1098]" gridcol="4" gridrow="10" pageId="11" pageNumber="444">1.68 ± 0.45 (1.36–1.99)</td>
<td id="1A1B1C5D8750006FFA89FBF718B3B067" box="[1381,1512,1073,1098]" gridcol="5" gridrow="10" pageId="11" pageNumber="444">2.39</td>
<td id="1A1B1C5D8750006FF9E8FBF71A41B067" box="[1540,1818,1073,1098]" gridcol="6" gridrow="10" pageId="11" pageNumber="444">3.10 ± 0.04 (3.07–3.13)</td>
<td id="1A1B1C5D8750006FF8D9FBF71ABDB067" box="[1845,2022,1073,1098]" gridcol="7" gridrow="10" pageId="11" pageNumber="444">2.57</td>
</tr>
<tr id="59CA75218750006FFF4EFBA01ABDB052" box="[162,2022,1126,1151]" gridrow="11" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFBA01DBDB052" box="[162,230,1126,1151]" gridcol="0" gridrow="11" pageId="11" pageNumber="444">FoL</th>
<td id="1A1B1C5D8750006FFEE0FBA01F58B052" box="[268,515,1126,1151]" gridcol="1" gridrow="11" pageId="11" pageNumber="444">4.60 ± 0.55 (3.75–5.24)</td>
<td id="1A1B1C5D8750006FFDCAFBA01E4DB052" box="[550,790,1126,1151]" gridcol="2" gridrow="11" pageId="11" pageNumber="444">4.12 ± 0.05 (4.08–4.15)</td>
<td id="1A1B1C5D8750006FFCD6FBA01971B052" box="[826,1066,1126,1151]" gridcol="3" gridrow="11" pageId="11" pageNumber="444">4.30 ± 0.46 (3.59–5.21)</td>
<td id="1A1B1C5D8750006FFBA7FBA0181CB052" box="[1099,1351,1126,1151]" gridcol="4" gridrow="11" pageId="11" pageNumber="444">4.35 ± 2.29 (2.73–5.97)</td>
<td id="1A1B1C5D8750006FFA89FBA018B3B052" box="[1381,1512,1126,1151]" gridcol="5" gridrow="11" pageId="11" pageNumber="444">6.43</td>
<td id="1A1B1C5D8750006FF9E8FBA01A41B052" box="[1540,1818,1126,1151]" gridcol="6" gridrow="11" pageId="11" pageNumber="444">8.72 ± 0.24 (8.55–8.89)</td>
<td id="1A1B1C5D8750006FF8D9FBA01ABDB052" box="[1845,2022,1126,1151]" gridcol="7" gridrow="11" pageId="11" pageNumber="444">6.16</td>
</tr>
<tr id="59CA75218750006FFF4EFB5D1ABDB099" box="[162,2022,1179,1204]" gridrow="12" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFB5D1DBDB099" box="[162,230,1179,1204]" gridcol="0" gridrow="12" pageId="11" pageNumber="444">HaL</th>
<td id="1A1B1C5D8750006FFEE0FB5D1F58B099" box="[268,515,1179,1204]" gridcol="1" gridrow="12" pageId="11" pageNumber="444">4.05 ± 0.49 (3.37–4.8)</td>
<td id="1A1B1C5D8750006FFDCAFB5D1E4DB099" box="[550,790,1179,1204]" gridcol="2" gridrow="12" pageId="11" pageNumber="444">3.59 ± 0.28 (3.39–3.79)</td>
<td id="1A1B1C5D8750006FFCD6FB5D1971B099" box="[826,1066,1179,1204]" gridcol="3" gridrow="12" pageId="11" pageNumber="444">4.6 ± 0.5 (3.64–5.39)</td>
<td id="1A1B1C5D8750006FFBA7FB5D181CB099" box="[1099,1351,1179,1204]" gridcol="4" gridrow="12" pageId="11" pageNumber="444">4.13 ± 2.26 (2.53–5.72)</td>
<td id="1A1B1C5D8750006FFA89FB5D18B3B099" box="[1381,1512,1179,1204]" gridcol="5" gridrow="12" pageId="11" pageNumber="444">5.65</td>
<td id="1A1B1C5D8750006FF9E8FB5D1A41B099" box="[1540,1818,1179,1204]" gridcol="6" gridrow="12" pageId="11" pageNumber="444">8.23 ± 0.03 (8.21–8.25)</td>
<td id="1A1B1C5D8750006FF8D9FB5D1ABDB099" box="[1845,2022,1179,1204]" gridcol="7" gridrow="12" pageId="11" pageNumber="444">5.29</td>
</tr>
<tr id="59CA75218750006FFF4EFB091ABDB0C5" box="[162,2022,1231,1256]" gridrow="13" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFB091DBDB0C5" box="[162,230,1231,1256]" gridcol="0" gridrow="13" pageId="11" pageNumber="444">IPTL</th>
<td id="1A1B1C5D8750006FFEE0FB091F58B0C5" box="[268,515,1231,1256]" gridcol="1" gridrow="13" pageId="11" pageNumber="444">0.54 ± 0.11 (0.44–0.72)</td>
<td id="1A1B1C5D8750006FFDCAFB091E4DB0C5" box="[550,790,1231,1256]" gridcol="2" gridrow="13" pageId="11" pageNumber="444">0.37 ± 0.01 (0.36–0.37)</td>
<td id="1A1B1C5D8750006FFCD6FB091971B0C5" box="[826,1066,1231,1256]" gridcol="3" gridrow="13" pageId="11" pageNumber="444">0.56 ± 0.07 (0.46–0.71)</td>
<td id="1A1B1C5D8750006FFBA7FB09181CB0C5" box="[1099,1351,1231,1256]" gridcol="4" gridrow="13" pageId="11" pageNumber="444">0.43 ± 0.04 (0.4–0.46)</td>
<td id="1A1B1C5D8750006FFA89FB0918B3B0C5" box="[1381,1512,1231,1256]" gridcol="5" gridrow="13" pageId="11" pageNumber="444">0.85</td>
<td id="1A1B1C5D8750006FF9E8FB091A41B0C5" box="[1540,1818,1231,1256]" gridcol="6" gridrow="13" pageId="11" pageNumber="444">1.14 ± 0.04 (1.11–1.17)</td>
<td id="1A1B1C5D8750006FF8D9FB091ABDB0C5" box="[1845,2022,1231,1256]" gridcol="7" gridrow="13" pageId="11" pageNumber="444">0.73</td>
</tr>
<tr id="59CA75218750006FFF4EFAC21ABDB130" box="[162,2022,1284,1309]" gridrow="14" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFAC21DBDB130" box="[162,230,1284,1309]" gridcol="0" gridrow="14" pageId="11" pageNumber="444">MPTL</th>
<td id="1A1B1C5D8750006FFEE0FAC21F58B130" box="[268,515,1284,1309]" gridcol="1" gridrow="14" pageId="11" pageNumber="444">0.70 ± 0.08 (0.83 –0.57)</td>
<td id="1A1B1C5D8750006FFDCAFAC21E4DB130" box="[550,790,1284,1309]" gridcol="2" gridrow="14" pageId="11" pageNumber="444">0.61 ± 0.01 (0.6–0.61)</td>
<td id="1A1B1C5D8750006FFCD6FAC21971B130" box="[826,1066,1284,1309]" gridcol="3" gridrow="14" pageId="11" pageNumber="444">0.74 ± 0.11 (0.57–0.93)</td>
<td id="1A1B1C5D8750006FFBA7FAC2181CB130" box="[1099,1351,1284,1309]" gridcol="4" gridrow="14" pageId="11" pageNumber="444">0.66 ± 0.17 (0.54–0.78)</td>
<td id="1A1B1C5D8750006FFA89FAC218B3B130" box="[1381,1512,1284,1309]" gridcol="5" gridrow="14" pageId="11" pageNumber="444">1.14</td>
<td id="1A1B1C5D8750006FF9E8FAC21A41B130" box="[1540,1818,1284,1309]" gridcol="6" gridrow="14" pageId="11" pageNumber="444">1.55 ± 0.01 (1.54–1.56)</td>
<td id="1A1B1C5D8750006FF8D9FAC21ABDB130" box="[1845,2022,1284,1309]" gridcol="7" gridrow="14" pageId="11" pageNumber="444">0.92</td>
</tr>
<tr id="59CA75218750006FFF4EFAFF1ABDB17F" box="[162,2022,1337,1362]" gridrow="15" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFAFF1DBDB17F" box="[162,230,1337,1362]" gridcol="0" gridrow="15" pageId="11" pageNumber="444">OPTL</th>
<td id="1A1B1C5D8750006FFEE0FAFF1F58B17F" box="[268,515,1337,1362]" gridcol="1" gridrow="15" pageId="11" pageNumber="444">0.21 ± 0.09 (0–0.29)</td>
<td id="1A1B1C5D8750006FFDCAFAFF1E4DB17F" box="[550,790,1337,1362]" gridcol="2" gridrow="15" pageId="11" pageNumber="444">0.16 ± 0.01 (0.15–0.16)</td>
<td id="1A1B1C5D8750006FFCD6FAFF1971B17F" box="[826,1066,1337,1362]" gridcol="3" gridrow="15" pageId="11" pageNumber="444">0.29 ± 0.04 (0.24–0.35)</td>
<td id="1A1B1C5D8750006FFBA7FAFF181CB17F" box="[1099,1351,1337,1362]" gridcol="4" gridrow="15" pageId="11" pageNumber="444">0.16 ± 0.13 (0.07–0.25)</td>
<td id="1A1B1C5D8750006FFA89FAFF18B3B17F" box="[1381,1512,1337,1362]" gridcol="5" gridrow="15" pageId="11" pageNumber="444">0.4</td>
<td id="1A1B1C5D8750006FF9E8FAFF1A41B17F" box="[1540,1818,1337,1362]" gridcol="6" gridrow="15" pageId="11" pageNumber="444">0.64 ± 0.13 (0.55–0.73)</td>
<td id="1A1B1C5D8750006FF8D9FAFF1ABDB17F" box="[1845,2022,1337,1362]" gridcol="7" gridrow="15" pageId="11" pageNumber="444">0.34</td>
</tr>
<tr id="59CA75218750006FFF4EFAA81ABDB1AA" box="[162,2022,1390,1415]" gridrow="16" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFAA81DBDB1AA" box="[162,230,1390,1415]" gridcol="0" gridrow="16" pageId="11" pageNumber="444">F3L</th>
<td id="1A1B1C5D8750006FFEE0FAA81F58B1AA" box="[268,515,1390,1415]" gridcol="1" gridrow="16" pageId="11" pageNumber="444">3.42 ± 0.47 (2.79–4.02)</td>
<td id="1A1B1C5D8750006FFDCAFAA81E4DB1AA" box="[550,790,1390,1415]" gridcol="2" gridrow="16" pageId="11" pageNumber="444">2.93 ± 0.23 (2.76–3.09)</td>
<td id="1A1B1C5D8750006FFCD6FAA81971B1AA" box="[826,1066,1390,1415]" gridcol="3" gridrow="16" pageId="11" pageNumber="444">2.79 ± 0.3 (2.31–3.35)</td>
<td id="1A1B1C5D8750006FFBA7FAA8181CB1AA" box="[1099,1351,1390,1415]" gridcol="4" gridrow="16" pageId="11" pageNumber="444">2.62 ± 1.49 (1.56–3.67)</td>
<td id="1A1B1C5D8750006FFA89FAA818B3B1AA" box="[1381,1512,1390,1415]" gridcol="5" gridrow="16" pageId="11" pageNumber="444">4.51</td>
<td id="1A1B1C5D8750006FF9E8FAA81A41B1AA" box="[1540,1818,1390,1415]" gridcol="6" gridrow="16" pageId="11" pageNumber="444">6.70 ± 0.17 (6.58–6.82)</td>
<td id="1A1B1C5D8750006FF8D9FAA81ABDB1AA" box="[1845,2022,1390,1415]" gridcol="7" gridrow="16" pageId="11" pageNumber="444">4.55</td>
</tr>
<tr id="59CA75218750006FFF4EFA651ABDB191" box="[162,2022,1443,1468]" gridrow="17" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFA651DBDB191" box="[162,230,1443,1468]" gridcol="0" gridrow="17" pageId="11" pageNumber="444">F3PW</th>
<td id="1A1B1C5D8750006FFEE0FA651F58B191" box="[268,515,1443,1468]" gridcol="1" gridrow="17" pageId="11" pageNumber="444">0.59 ± 0.08 (0.44–0.68)</td>
<td id="1A1B1C5D8750006FFDCAFA651E4DB191" box="[550,790,1443,1468]" gridcol="2" gridrow="17" pageId="11" pageNumber="444">0.51 ± 0.04 (0.48–0.53)</td>
<td id="1A1B1C5D8750006FFCD6FA651971B191" box="[826,1066,1443,1468]" gridcol="3" gridrow="17" pageId="11" pageNumber="444">0.65 ± 0.09 (0.51–0.81)</td>
<td id="1A1B1C5D8750006FFBA7FA65181CB191" box="[1099,1351,1443,1468]" gridcol="4" gridrow="17" pageId="11" pageNumber="444">0.62 ± 0.36 (0.36–0.87)</td>
<td id="1A1B1C5D8750006FFA89FA6518B3B191" box="[1381,1512,1443,1468]" gridcol="5" gridrow="17" pageId="11" pageNumber="444">1.28</td>
<td id="1A1B1C5D8750006FF9E8FA651A41B191" box="[1540,1818,1443,1468]" gridcol="6" gridrow="17" pageId="11" pageNumber="444">1.77 ± 0.07 (1.36–1.72)</td>
<td id="1A1B1C5D8750006FF8D9FA651ABDB191" box="[1845,2022,1443,1468]" gridcol="7" gridrow="17" pageId="11" pageNumber="444">0.92</td>
</tr>
<tr id="59CA75218750006FFF4EFA1E1ABDB1DC" box="[162,2022,1496,1521]" gridrow="18" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EFA1E1DBDB1DC" box="[162,230,1496,1521]" gridcol="0" gridrow="18" pageId="11" pageNumber="444">F4PW</th>
<td id="1A1B1C5D8750006FFEE0FA1E1F58B1DC" box="[268,515,1496,1521]" gridcol="1" gridrow="18" pageId="11" pageNumber="444">0.56 ± 0.10 (0.4–0.67)</td>
<td id="1A1B1C5D8750006FFDCAFA1E1E4DB1DC" box="[550,790,1496,1521]" gridcol="2" gridrow="18" pageId="11" pageNumber="444">0.50 ± 0.08 (0.44–0.55)</td>
<td id="1A1B1C5D8750006FFCD6FA1E1971B1DC" box="[826,1066,1496,1521]" gridcol="3" gridrow="18" pageId="11" pageNumber="444">0.61 ± 0.11 (0.42–0.8)</td>
<td id="1A1B1C5D8750006FFBA7FA1E181CB1DC" box="[1099,1351,1496,1521]" gridcol="4" gridrow="18" pageId="11" pageNumber="444">0.58 ± 0.37 (0.32–0.84)</td>
<td id="1A1B1C5D8750006FFA89FA1E18B3B1DC" box="[1381,1512,1496,1521]" gridcol="5" gridrow="18" pageId="11" pageNumber="444">1.27</td>
<td id="1A1B1C5D8750006FF9E8FA1E1A41B1DC" box="[1540,1818,1496,1521]" gridcol="6" gridrow="18" pageId="11" pageNumber="444">1.79 ± 0.01 (1.8–1.78)</td>
<td id="1A1B1C5D8750006FF8D9FA1E1ABDB1DC" box="[1845,2022,1496,1521]" gridcol="7" gridrow="18" pageId="11" pageNumber="444">0.81</td>
</tr>
<tr id="59CA75218750006FFF4EF9CB1ABDB20B" box="[162,2022,1549,1574]" gridrow="19" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EF9CB1DBDB20B" box="[162,230,1549,1574]" gridcol="0" gridrow="19" pageId="11" pageNumber="444">FeL</th>
<td id="1A1B1C5D8750006FFEE0F9CB1F58B20B" box="[268,515,1549,1574]" gridcol="1" gridrow="19" pageId="11" pageNumber="444">7.70 ± 0.79 (6.2–8.51)</td>
<td id="1A1B1C5D8750006FFDCAF9CB1E4DB20B" box="[550,790,1549,1574]" gridcol="2" gridrow="19" pageId="11" pageNumber="444">7.01 ± 0.06 (6.97–7.05)</td>
<td id="1A1B1C5D8750006FFCD6F9CB1971B20B" box="[826,1066,1549,1574]" gridcol="3" gridrow="19" pageId="11" pageNumber="444">7.68 ± 0.77 (6.55–9.15)</td>
<td id="1A1B1C5D8750006FFBA7F9CB181CB20B" box="[1099,1351,1549,1574]" gridcol="4" gridrow="19" pageId="11" pageNumber="444">7.63 ± 3.83 (4.92–10.34)</td>
<td id="1A1B1C5D8750006FFA89F9CB18B3B20B" box="[1381,1512,1549,1574]" gridcol="5" gridrow="19" pageId="11" pageNumber="444">11.16</td>
<td id="1A1B1C5D8750006FF9E8F9CB1A41B20B" box="[1540,1818,1549,1574]" gridcol="6" gridrow="19" pageId="11" pageNumber="444">14.19 ± 0.20 (14.05–14.33)</td>
<td id="1A1B1C5D8750006FF8D9F9CB1ABDB20B" box="[1845,2022,1549,1574]" gridcol="7" gridrow="19" pageId="11" pageNumber="444">10.62</td>
</tr>
<tr id="59CA75218750006FFF4EF9841ABDB276" box="[162,2022,1602,1627]" gridrow="20" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EF9841DBDB276" box="[162,230,1602,1627]" gridcol="0" gridrow="20" pageId="11" pageNumber="444">TL</th>
<td id="1A1B1C5D8750006FFEE0F9841F58B276" box="[268,515,1602,1627]" gridcol="1" gridrow="20" pageId="11" pageNumber="444">8.45 ± 0.83 (7.01–9.29)</td>
<td id="1A1B1C5D8750006FFDCAF9841E4DB276" box="[550,790,1602,1627]" gridcol="2" gridrow="20" pageId="11" pageNumber="444">7.83 ± 0.31 (7.61–8.05)</td>
<td id="1A1B1C5D8750006FFCD6F9841971B276" box="[826,1066,1602,1627]" gridcol="3" gridrow="20" pageId="11" pageNumber="444">8.6 ± 0.77 (7.55–9.84)</td>
<td id="1A1B1C5D8750006FFBA7F984181CB276" box="[1099,1351,1602,1627]" gridcol="4" gridrow="20" pageId="11" pageNumber="444">7.72 ± 3.5 (5.24–10.19)</td>
<td id="1A1B1C5D8750006FFA89F98418B3B276" box="[1381,1512,1602,1627]" gridcol="5" gridrow="20" pageId="11" pageNumber="444">11.3</td>
<td id="1A1B1C5D8750006FF9E8F9841A41B276" box="[1540,1818,1602,1627]" gridcol="6" gridrow="20" pageId="11" pageNumber="444">15.01 ± 0.20 (14.87–15.15)</td>
<td id="1A1B1C5D8750006FF8D9F9841ABDB276" box="[1845,2022,1602,1627]" gridcol="7" gridrow="20" pageId="11" pageNumber="444">10.77</td>
</tr>
<tr id="59CA75218750006FFF4EF9B11ABDB2BD" box="[162,2022,1655,1680]" gridrow="21" pageId="11" pageNumber="444">
<th id="1A1B1C5D8750006FFF4EF9B11DBDB2BD" box="[162,230,1655,1680]" gridcol="0" gridrow="21" pageId="11" pageNumber="444">IMTL</th>
<td id="1A1B1C5D8750006FFEE0F9B11F58B2BD" box="[268,515,1655,1680]" gridcol="1" gridrow="21" pageId="11" pageNumber="444">0.67 ± 0.13 (0.51–0.84)</td>
<td id="1A1B1C5D8750006FFDCAF9B11E4DB2BD" box="[550,790,1655,1680]" gridcol="2" gridrow="21" pageId="11" pageNumber="444">0.60 ± 0.04 (0.57–0.62)</td>
<td id="1A1B1C5D8750006FFCD6F9B11971B2BD" box="[826,1066,1655,1680]" gridcol="3" gridrow="21" pageId="11" pageNumber="444">0.64 ± 0.06 (0.58–0.76)</td>
<td id="1A1B1C5D8750006FFBA7F9B1181CB2BD" box="[1099,1351,1655,1680]" gridcol="4" gridrow="21" pageId="11" pageNumber="444">0.63 ± 0.18 (0.5–0.76)</td>
<td id="1A1B1C5D8750006FFA89F9B118B3B2BD" box="[1381,1512,1655,1680]" gridcol="5" gridrow="21" pageId="11" pageNumber="444">0.83</td>
<td id="1A1B1C5D8750006FF9E8F9B11A41B2BD" box="[1540,1818,1655,1680]" gridcol="6" gridrow="21" pageId="11" pageNumber="444">1.13 ± 0.06 (1.09–1.17)</td>
<td id="1A1B1C5D8750006FF8D9F9B11ABDB2BD" box="[1845,2022,1655,1680]" gridcol="7" gridrow="21" pageId="11" pageNumber="444">0.82</td>
</tr>
</table>
</paragraph>
<paragraph id="E74577638750FF9BF95FF9601AB1B292" blockId="11.[1715,2026,1702,1727]" box="[1715,2026,1702,1727]" pageId="11" pageNumber="444">
<tableNote id="1A1C76ED8750FF9BF95FF9601AB1B292" box="[1715,2026,1702,1727]" pageId="11" pageNumber="444" targetBox="[162,2022,539,1680]" targetPageId="11">.....Continued on the next page</tableNote>
</paragraph>
<caption id="B38527EB8757FF9CFF74FC6A1CCDB7EB" box="[152,406,940,966]" isContinuationCaption="true" pageId="12" pageNumber="445" startId="12.[152,244,940,966]" targetType="table">
<paragraph id="E74577638757FF9CFF74FC6A1CCDB7EB" blockId="12.[152,406,940,966]" box="[152,406,940,966]" pageId="12" pageNumber="445">
<emphasis id="D58EAB718757FF9CFF74FC6A1C55B7EB" bold="true" box="[152,270,940,966]" pageId="12" pageNumber="445">TABLE 4.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="E74577638757FF9CFE44FC131AC3B0C9" pageId="12" pageNumber="445">
<table id="95FA85C38757006FFF7BFC131A80B0C9" box="[151,2011,981,1252]" colsContinueFrom="11.[162,2022,539,1680]" gridcols="8" gridrows="6" pageId="12" pageNumber="445">
<tr id="59CA75218757006FFF7BFC131A80B7C3" box="[151,2011,981,1006]" gridrow="0" pageId="12" pageNumber="445" rowspan-0="1">
<th id="1A1B1C5D8757006FFEEDFC131E50B7C3" box="[257,779,981,1006]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="12" pageNumber="445">
<taxonomicName id="20FA0CE08757FF9CFE44FC131F55B7C3" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[424,526,981,1006]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="445" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718757FF9CFE44FC131F55B7C3" box="[424,526,981,1006]" italics="true" pageId="12" pageNumber="445">E. coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718757FF9CFDF8FC131F3EB7C3" bold="true" box="[532,613,981,1006]" pageId="12" pageNumber="445">
<taxonomicNameLabel id="CEBD160A8757FF9CFDF8FC131F3EB7C3" box="[532,613,981,1006]" pageId="12" pageNumber="445" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="1A1B1C5D8757006FFCC3FC131874B7C3" box="[815,1327,981,1006]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="12" pageNumber="445">
<taxonomicName id="20FA0CE08757FF9CFC3BFC1319CAB7C3" box="[983,1169,981,1006]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="445" phylum="Chordata" rank="species" species="cystignathoides">
<emphasis id="D58EAB718757FF9CFC3BFC1319CAB7C3" box="[983,1169,981,1006]" italics="true" pageId="12" pageNumber="445">E. cystignathoides</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8757006FFAB5FC131A55B7C3" box="[1369,1806,981,1006]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="12" pageNumber="445">
<taxonomicName id="20FA0CE08757FF9CFA2EFC131B7CB7C3" box="[1474,1575,981,1006]" class="Amphibia" family="Eleutherodactylidae" genus="Syrrhophus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="445" phylum="Chordata" rank="species" species="dennisi">
<emphasis id="D58EAB718757FF9CFA2EFC131B7CB7C3" box="[1474,1575,981,1006]" italics="true" pageId="12" pageNumber="445">E. dennisi</emphasis>
</taxonomicName>
</th>
<th id="1A1B1C5D8757006FF8C6FC131A80B7C3" box="[1834,2011,981,1006]" gridcol="7" gridrow="0" pageId="12" pageNumber="445">
<emphasis id="D58EAB718757FF9CF8C6FC131A62B7C3" box="[1834,1849,981,1006]" italics="true" pageId="12" pageNumber="445">E</emphasis>
. sp. Ahuacatlán
</th>
</tr>
<tr id="59CA75218757006FFF7BFC3B1A80B03B" box="[151,2011,1021,1046]" gridrow="1" pageId="12" pageNumber="445" rowspan-0="1">
<td id="1A1B1C5D8757006FFEEDFC3B1CACB03B" box="[257,503,1021,1046]" gridcol="1" gridrow="1" pageId="12" pageNumber="445">
Males (
<emphasis id="D58EAB718757FF9CFE6EFC3B1CC8B03B" box="[386,403,1021,1046]" italics="true" pageId="12" pageNumber="445">N</emphasis>
= 8)
</td>
<td id="1A1B1C5D8757006FFDF7FC3B1E50B03B" box="[539,779,1021,1046]" gridcol="2" gridrow="1" pageId="12" pageNumber="445">
Females (
<emphasis id="D58EAB718757FF9CFD48FC3B1FEEB03B" box="[676,693,1021,1046]" italics="true" pageId="12" pageNumber="445">N</emphasis>
= 2)
</td>
<td id="1A1B1C5D8757006FFCC3FC3B1944B03B" box="[815,1055,1021,1046]" gridcol="3" gridrow="1" pageId="12" pageNumber="445">
Males (
<emphasis id="D58EAB718757FF9CFC4AFC3B1EECB03B" box="[934,951,1021,1046]" italics="true" pageId="12" pageNumber="445">N</emphasis>
= 10)
</td>
<td id="1A1B1C5D8757006FFBA0FC3B1874B03B" box="[1100,1327,1021,1046]" gridcol="4" gridrow="1" pageId="12" pageNumber="445">
Females (
<emphasis id="D58EAB718757FF9CFB23FC3B19BBB03B" box="[1231,1248,1021,1046]" italics="true" pageId="12" pageNumber="445">N</emphasis>
= 2)
</td>
<td id="1A1B1C5D8757006FFAB5FC3B1886B03B" box="[1369,1501,1021,1046]" gridcol="5" gridrow="1" pageId="12" pageNumber="445">
Male (
<emphasis id="D58EAB718757FF9CFA70FC3B18F6B03B" box="[1436,1453,1021,1046]" italics="true" pageId="12" pageNumber="445">N</emphasis>
= 1)
</td>
<td id="1A1B1C5D8757006FFA14FC3B1A55B03B" box="[1528,1806,1021,1046]" gridcol="6" gridrow="1" pageId="12" pageNumber="445">
Females (
<emphasis id="D58EAB718757FF9CF978FC3B1BFEB03B" box="[1684,1701,1021,1046]" italics="true" pageId="12" pageNumber="445">N</emphasis>
= 2)
</td>
<td id="1A1B1C5D8757006FF8C6FC3B1A80B03B" box="[1834,2011,1021,1046]" gridcol="7" gridrow="1" pageId="12" pageNumber="445">
Female (
<emphasis id="D58EAB718757FF9CF863FC3B1AFBB03B" box="[1935,1952,1021,1046]" italics="true" pageId="12" pageNumber="445">N</emphasis>
= 1)
</td>
</tr>
<tr id="59CA75218757006FFF7BFBEB1A80B06B" box="[151,2011,1069,1094]" gridrow="2" pageId="12" pageNumber="445">
<th id="1A1B1C5D8757006FFF7BFBEB1D84B06B" box="[151,223,1069,1094]" gridcol="0" gridrow="2" pageId="12" pageNumber="445">OMTL</th>
<td id="1A1B1C5D8757006FFEEDFBEB1CACB06B" box="[257,503,1069,1094]" gridcol="1" gridrow="2" pageId="12" pageNumber="445">0.43 ± 0.08 (0.3–0.56)</td>
<td id="1A1B1C5D8757006FFDF7FBEB1E50B06B" box="[539,779,1069,1094]" gridcol="2" gridrow="2" pageId="12" pageNumber="445">0.32 ± 0.08 (0.26–0.37)</td>
<td id="1A1B1C5D8757006FFCC3FBEB1944B06B" box="[815,1055,1069,1094]" gridcol="3" gridrow="2" pageId="12" pageNumber="445">0.42 ± 0.06 (0.31–0.49)</td>
<td id="1A1B1C5D8757006FFBA0FBEB1874B06B" box="[1100,1327,1069,1094]" gridcol="4" gridrow="2" pageId="12" pageNumber="445">0.39 ± 0.2 (0.25–0.53)</td>
<td id="1A1B1C5D8757006FFAB5FBEB1886B06B" box="[1369,1501,1069,1094]" gridcol="5" gridrow="2" pageId="12" pageNumber="445">0.7</td>
<td id="1A1B1C5D8757006FFA14FBEB1A55B06B" box="[1528,1806,1069,1094]" gridcol="6" gridrow="2" pageId="12" pageNumber="445">0.83 ± 0.01 (0.82–0.83)</td>
<td id="1A1B1C5D8757006FF8C6FBEB1A80B06B" box="[1834,2011,1069,1094]" gridcol="7" gridrow="2" pageId="12" pageNumber="445">0.58</td>
</tr>
<tr id="59CA75218757006FFF7BFBA71A80B057" box="[151,2011,1121,1146]" gridrow="3" pageId="12" pageNumber="445">
<th id="1A1B1C5D8757006FFF7BFBA71D84B057" box="[151,223,1121,1146]" gridcol="0" gridrow="3" pageId="12" pageNumber="445">FL</th>
<td id="1A1B1C5D8757006FFEEDFBA71CACB057" box="[257,503,1121,1146]" gridcol="1" gridrow="3" pageId="12" pageNumber="445">7.69 ± 0.93 (8.79 –6.33)</td>
<td id="1A1B1C5D8757006FFDF7FBA71E50B057" box="[539,779,1121,1146]" gridcol="2" gridrow="3" pageId="12" pageNumber="445">6.88 ± 0.24 (6.71–7.05)</td>
<td id="1A1B1C5D8757006FFCC3FBA71944B057" box="[815,1055,1121,1146]" gridcol="3" gridrow="3" pageId="12" pageNumber="445">8.35 ± 0.82 (7.09–9.77)</td>
<td id="1A1B1C5D8757006FFBA0FBA71874B057" box="[1100,1327,1121,1146]" gridcol="4" gridrow="3" pageId="12" pageNumber="445">7.43 ± 3.49 (4.96–9.9)</td>
<td id="1A1B1C5D8757006FFAB5FBA71886B057" box="[1369,1501,1121,1146]" gridcol="5" gridrow="3" pageId="12" pageNumber="445">10.26</td>
<td id="1A1B1C5D8757006FFA14FBA71A55B057" box="[1528,1806,1121,1146]" gridcol="6" gridrow="3" pageId="12" pageNumber="445">13.50 ± 0.42 (13.2–13.8)</td>
<td id="1A1B1C5D8757006FF8C6FBA71A80B057" box="[1834,2011,1121,1146]" gridcol="7" gridrow="3" pageId="12" pageNumber="445">10.29</td>
</tr>
<tr id="59CA75218757006FFF7BFB501A80B082" box="[151,2011,1174,1199]" gridrow="4" pageId="12" pageNumber="445">
<th id="1A1B1C5D8757006FFF7BFB501D84B082" box="[151,223,1174,1199]" gridcol="0" gridrow="4" pageId="12" pageNumber="445">T4L</th>
<td id="1A1B1C5D8757006FFEEDFB501CACB082" box="[257,503,1174,1199]" gridcol="1" gridrow="4" pageId="12" pageNumber="445">7.0 ± 0.82 (5.91–8.18)</td>
<td id="1A1B1C5D8757006FFDF7FB501E50B082" box="[539,779,1174,1199]" gridcol="2" gridrow="4" pageId="12" pageNumber="445">6.29 ± 0.31 (6.07–6.51)</td>
<td id="1A1B1C5D8757006FFCC3FB501944B082" box="[815,1055,1174,1199]" gridcol="3" gridrow="4" pageId="12" pageNumber="445">5.09 ± 0.61 (4.2–6.24)</td>
<td id="1A1B1C5D8757006FFBA0FB501874B082" box="[1100,1327,1174,1199]" gridcol="4" gridrow="4" pageId="12" pageNumber="445">4.55 ± 2.18 (3.0–6.09)</td>
<td id="1A1B1C5D8757006FFAB5FB501886B082" box="[1369,1501,1174,1199]" gridcol="5" gridrow="4" pageId="12" pageNumber="445">9.33</td>
<td id="1A1B1C5D8757006FFA14FB501A55B082" box="[1528,1806,1174,1199]" gridcol="6" gridrow="4" pageId="12" pageNumber="445">12.74 ± 0.02 (12.72–12.75)</td>
<td id="1A1B1C5D8757006FF8C6FB501A80B082" box="[1834,2011,1174,1199]" gridcol="7" gridrow="4" pageId="12" pageNumber="445">9.59</td>
</tr>
<tr id="59CA75218757006FFF7BFB0D1A80B0C9" box="[151,2011,1227,1252]" gridrow="5" pageId="12" pageNumber="445">
<th id="1A1B1C5D8757006FFF7BFB0D1D84B0C9" box="[151,223,1227,1252]" gridcol="0" gridrow="5" pageId="12" pageNumber="445">T4PW</th>
<td id="1A1B1C5D8757006FFEEDFB0D1CACB0C9" box="[257,503,1227,1252]" gridcol="1" gridrow="5" pageId="12" pageNumber="445">0.58 ± 0.11 (0.43–0.76)</td>
<td id="1A1B1C5D8757006FFDF7FB0D1E50B0C9" box="[539,779,1227,1252]" gridcol="2" gridrow="5" pageId="12" pageNumber="445">0.51 ± 0.04 (0.48–0.54)</td>
<td id="1A1B1C5D8757006FFCC3FB0D1944B0C9" box="[815,1055,1227,1252]" gridcol="3" gridrow="5" pageId="12" pageNumber="445">0.57 ± 0.08 (0.48–0.70)</td>
<td id="1A1B1C5D8757006FFBA0FB0D1874B0C9" box="[1100,1327,1227,1252]" gridcol="4" gridrow="5" pageId="12" pageNumber="445">0.58 ± 0.3 (0.36–0.79)</td>
<td id="1A1B1C5D8757006FFAB5FB0D1886B0C9" box="[1369,1501,1227,1252]" gridcol="5" gridrow="5" pageId="12" pageNumber="445">0.9</td>
<td id="1A1B1C5D8757006FFA14FB0D1A55B0C9" box="[1528,1806,1227,1252]" gridcol="6" gridrow="5" pageId="12" pageNumber="445">1.27 ± 0.11 (1.19–1.34)</td>
<td id="1A1B1C5D8757006FF8C6FB0D1A80B0C9" box="[1834,2011,1227,1252]" gridcol="7" gridrow="5" pageId="12" pageNumber="445">0.87</td>
</tr>
</table>
</paragraph>
<paragraph id="E74577638756FF9DFF2BFF5019EDB56C" blockId="13.[151,1437,150,1041]" pageId="13" pageNumber="446">
<emphasis id="D58EAB718756FF9DFF2BFF501E3CB49D" bold="true" box="[199,871,150,177]" pageId="13" pageNumber="446">
Coloration in life (from photos, but not of the 
<typeStatus id="3841C9C18756FF9DFD1CFF501E01B49D" box="[752,858,150,176]" pageId="13" pageNumber="446" type="holotype">holotype</typeStatus>
).
</emphasis>
Dorsal surface of the head Drab (19) which fades from interorbital bar to Glaucus (272); a Vandyke Brown (282) irregular interorbital bar; dorsal surfaces of head and body with Vandyke Brown (282) markings; back Smoke Gray (267) with two ill-defined longitudinal bands Salmon Color (58); dorsal surface of arms with Salmon Color (58) spots; legs with transverse bands of Vandyke Brown (282) on a Salmon Color (58) to Smoke Gray (267) background; iris Light Yellow Ocher (13).
</paragraph>
<paragraph id="E74577638756FF9DFF2BFE8D19E7B580" blockId="13.[151,1437,150,1041]" pageId="13" pageNumber="446">
<emphasis id="D58EAB718756FF9DFF2BFE8D1E0AB549" bold="true" box="[199,849,330,357]" pageId="13" pageNumber="446">
Measurements of the 
<typeStatus id="3841C9C18756FF9DFE25FE8D1F6BB548" box="[457,560,331,357]" pageId="13" pageNumber="446" type="holotype">holotype</typeStatus>
(measurements in mm).
</emphasis>
SVL 18.7, HL 7.62, HW 7.16, EW 1.38, END 2.43, IOD 2.59, TD 1.86, EL 2.3, IND 2.02, FoL 5.23, HaL 4.21, IPTL 0.72, MPTL 0.8, OPTL 0.18, F3L 3.85, F3PW 0.64, F4PW 0.67, FeL 8.51, TL 9.29, IMTL 0.84, OMTL 0.56, FL 8.61, T4L 7.62, T4PW 0.76.
</paragraph>
<paragraph id="E74577638756FF9DFF2BFE701EE8B6A8" blockId="13.[151,1437,150,1041]" pageId="13" pageNumber="446">
<emphasis id="D58EAB718756FF9DFF2BFE701C64B5FD" bold="true" box="[199,319,438,464]" pageId="13" pageNumber="446">Variation.</emphasis>
<materialsCitation id="57927D3E8756FF9DFEA8FE711F96B634" ID-GBIF-Occurrence="4898140301" collectionCode="IBH" pageId="13" pageNumber="446" specimenCode="IBH 32711-32712" specimenCount="9" typeStatus="paratype">
Variations in the morphometric data of the type series and species compared are summarized in 
<tableCitation id="AA7842D88756FF9DFAB2FE711DF3B5D8" captionStart="TABLE 4" captionStartId="11.[152,244,462,488]" captionTargetBox="[162,2022,539,1680]" captionText="TABLE 4. Morphometric data (in mm) of the type series of Eleutherodactylus coelum sp. nov. and closely related species. N = sample size. For each character, the mean ± SD and range (in parentheses) are presented. Abbreviations for morphological characters are explained in the Materials and methods section." httpUri="http://table.plazi.org/id/B38527EB8750FF9BFF74FE081B16B621" pageId="13" pageNumber="446" tableUuid="B38527EB8750FF9BFF74FE081B16B621">Table 4</tableCitation>
. We observed little variation in coloration and no evident morphological differences among the 
<specimenCount id="F1FCBCEA8756FF9DFB1DFE1D18C2B5D8" box="[1265,1433,475,501]" count="9" pageId="13" pageNumber="446" type="generic" typeStatus="paratype">nine paratypes</specimenCount>
. The dorsal background of 
<specimenCode id="B75CDF188756FF9DFE25FE391FC0B634" box="[457,667,511,537]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 32711–32712</specimenCode>
and
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFD34FE391EF0B634" ID-GBIF-Occurrence="4898140308" box="[728,939,510,537]" collectionCode="IBH" isTailEnumeration="true" pageId="13" pageNumber="446" specimenCode="IBH 36349-36350" specimenCount="1" typeStatus="paratype">
<specimenCode id="B75CDF188756FF9DFD34FE391EF0B634" box="[728,939,510,537]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 36349–36350</specimenCode>
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFC59FE391CD0B610" ID-GBIF-Occurrence="4898140303" collectionCode="IBH" pageId="13" pageNumber="446" specimenCode="IBH 30477" specimenCount="1" typeStatus="holotype">
is paler than the 
<typeStatus id="3841C9C18756FF9DFB95FE391984B634" box="[1145,1247,511,537]" pageId="13" pageNumber="446" type="holotype">holotype</typeStatus>
. The interorbital bar in 
<specimenCode id="B75CDF188756FF9DFF33FDE51C07B610" box="[223,348,547,573]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 30477</specimenCode>
and
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFE7FFDE51F3FB610" ID-GBIF-Occurrence="4898140322" box="[403,612,546,573]" collectionCode="IBH" isTailEnumeration="true" pageId="13" pageNumber="446" specimenCode="IBH 36352-36353" specimenCount="1" typeStatus="holotype">
<specimenCode id="B75CDF188756FF9DFE7FFDE51F3FB610" box="[403,612,546,573]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 36352–36353</specimenCode>
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFD87FDE519E7B610" ID-GBIF-Occurrence="4898140305" box="[619,1212,547,573]" collectionCode="IBH" pageId="13" pageNumber="446" specimenCode="IBH 32711-32712" specimenCount="1" typeStatus="holotype">
is fainter than the 
<typeStatus id="3841C9C18756FF9DFCD9FDE51EC0B610" box="[821,923,547,573]" pageId="13" pageNumber="446" type="holotype">holotype</typeStatus>
; in the 
<specimenCode id="B75CDF188756FF9DFC00FDE519E7B610" box="[1004,1212,547,573]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 32711–32712</specimenCode>
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFB2FFDE51E35B64C" ID-GBIF-Occurrence="4898140319" collectionCode="IBH" pageId="13" pageNumber="446" specimenCode="IBH 36353-36354" specimenCount="1" typeStatus="holotype">
is narrow and well-defined. The two dorsal longitudinal bands in 
<specimenCode id="B75CDF188756FF9DFD7BFD811E30B64C" box="[663,875,582,609]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 36353–36354</specimenCode>
,
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFC99FD811EAFB64C" ID-GBIF-Occurrence="4898140318" box="[885,1012,583,609]" collectionCode="IBH" isTailEnumeration="true" pageId="13" pageNumber="446" specimenCode="IBH 30477" specimenCount="1" typeStatus="holotype">
<specimenCode id="B75CDF188756FF9DFC99FD811EAFB64C" box="[885,1012,583,609]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 30477</specimenCode>
</materialsCitation>
, 
<materialsCitation id="57927D3E8756FF9DFC13FD8119F1B64C" ID-GBIF-Occurrence="4898140306" box="[1023,1194,583,609]" collectionCode="IBH" pageId="13" pageNumber="446" specimenCode="IBH 30480" specimenCount="1" typeStatus="holotype">
and 
<specimenCode id="B75CDF188756FF9DFBC2FD8119F1B64C" box="[1070,1194,583,609]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 30480</specimenCode>
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFB5DFD811E73B6A8" ID-GBIF-Occurrence="4898140312" collectionCode="IBH" pageId="13" pageNumber="446" specimenCode="IBH 36533" specimenCount="1" typeStatus="holotype">
are not visible. There are no transverse dark bands on the legs of 
<specimenCode id="B75CDF188756FF9DFD90FDAD1FA2B6A8" box="[636,761,618,645]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 36533</specimenCode>
and
</materialsCitation>
<materialsCitation id="57927D3E8756FF9DFCC3FDAD1EF4B6A8" ID-GBIF-Occurrence="4898140321" box="[815,943,619,645]" collectionCode="IBH" isTailEnumeration="true" pageId="13" pageNumber="446" specimenCode="IBH 32711" specimenCount="1" typeStatus="holotype">
<specimenCode id="B75CDF188756FF9DFCC3FDAD1EF4B6A8" box="[815,943,619,645]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="13" pageNumber="446">IBH 32711</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AFE024E88756FF9EFF2BFD481F81B4D5" lastPageId="14" lastPageNumber="447" pageId="13" pageNumber="446" type="distribution">
<paragraph id="E74577638756FF9DFF2BFD48198CB03C" blockId="13.[151,1437,150,1041]" pageId="13" pageNumber="446">
<emphasis id="D58EAB718756FF9DFF2BFD481F15B685" bold="true" box="[199,590,654,681]" pageId="13" pageNumber="446">Distribution and natural history.</emphasis>
<taxonomicName id="20FA0CE08756FF9DFDBAFD491E23B684" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[598,888,655,681]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="446" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718756FF9DFDBAFD491E23B684" box="[598,888,655,681]" italics="true" pageId="13" pageNumber="446">Eleutherodactylus coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718756FF9DFC93FD491E82B684" bold="true" box="[895,985,655,681]" pageId="13" pageNumber="446">
<taxonomicNameLabel id="CEBD160A8756FF9DFC93FD491E82B684" box="[895,985,655,681]" pageId="13" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known for the locality Alta Cima and from Gómez Farías, in the El Cielo Biosphere Reserve, in southwestern of the state of 
<collectingRegion id="253EB9818756FF9DFBB4FD751987B6E0" box="[1112,1244,691,717]" country="Mexico" name="Tamaulipas" pageId="13" pageNumber="446">Tamaulipas</collectingRegion>
, 
<collectingCountry id="9FED37F38756FF9DFB0AFD751867B6E0" box="[1254,1340,691,717]" name="Mexico" pageId="13" pageNumber="446">Mexico</collectingCountry>
(
<figureCitation id="7FC16BE68756FF9DFAA6FD7518D0B6E0" box="[1354,1419,691,717]" captionStart="FIGURE 1" captionStartId="1.[151,250,1923,1947]" captionTargetBox="[158,1427,1001,1891]" captionTargetId="figure-295@1.[151,1436,993,1900]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of some members of the Eleutherodactylus longipes species clade in eastern Mexico. State names are indicated." figureDoi="http://doi.org/10.5281/zenodo.12190345" httpUri="https://zenodo.org/record/12190345/files/figure.png" pageId="13" pageNumber="446">Fig. 1</figureCitation>
), in the foothills of the Sierra Madre Oriental known as Sierra de Cucharas and Sierra Chiquita. The species occurs from low to moderate elevations (approximately 
<quantity id="2002DA868756FF9DFD57FD3C1E13B738" box="[699,840,762,789]" metricMagnitude="2" metricUnit="m" metricValue="8.66" metricValueMax="13.819999999999999" metricValueMin="3.5" pageId="13" pageNumber="446" unit="m" value="866.0" valueMax="1382.0" valueMin="350.0">350–1382 m</quantity>
elevation) in the subtropical evergreen and mountain cloud forests. Specimens have been found on top of bush leaves (
<figureCitation id="7FC16BE68756FF9DFC9BFCD91EE0B714" box="[887,955,798,825]" captionStart="FIGURE 5" captionStartId="13.[151,250,1978,2002]" captionTargetBox="[249,1339,1137,1954]" captionTargetId="figure-453@13.[249,1339,1137,1954]" captionTargetPageId="13" captionText="FIGURE 5. Similar habitat where Eleutherodactylus coelum sp. nov. individuals were found. Photo: Omar Becerra Soria." figureDoi="http://doi.org/10.5281/zenodo.12190355" httpUri="https://zenodo.org/record/12190355/files/figure.png" pageId="13" pageNumber="446">Fig. 5</figureCitation>
), under rocks along a dry river, and on top of water containers that local people name canoas. It must be a nocturnal species like its congeners. The individuals collected in 
<date id="934451A38756FF9DFECDFCA11C82B7AC" box="[289,473,871,897]" pageId="13" pageNumber="446" value="2018-09">September 2018</date>
were seen during the day under rocks and were not emitting any advertisement calls; however, this species is likely to be acoustically active like other species in the 
<taxonomicName id="20FA0CE08756FF9DFBAAFC4D1993B788" box="[1094,1224,907,933]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="446" phylum="Chordata" rank="species" species="longipes">
<emphasis id="D58EAB718756FF9DFBAAFC4D190CB789" box="[1094,1111,907,932]" italics="true" pageId="13" pageNumber="446">E</emphasis>
. 
<emphasis id="D58EAB718756FF9DFB85FC4D1993B788" box="[1129,1224,907,933]" italics="true" pageId="13" pageNumber="446">longipes</emphasis>
</taxonomicName>
species clade and other congeners. In 
<specimenCount id="F1FCBCEA8756FF9DFE9FFC691F42B7E4" box="[371,537,943,969]" count="2" pageId="13" pageNumber="446" type="generic">two specimens</specimenCount>
(IBH36349–36350), we found remnants of insects (coleopterans and formicids), malacostracan (isopods), and arachnids (spiders and pseudoscorpions). It occurs in sympatry with 
<taxonomicName id="20FA0CE08756FF9DFB08FC151C5EB03C" authorityName="Rovito &amp; Parra-Olea" authorityYear="2015" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="446" phylum="Chordata" rank="species" species="cieloensis">
<emphasis id="D58EAB718756FF9DFB08FC151C5EB03C" italics="true" pageId="13" pageNumber="446">Chiropterotriton cieloensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="20FA0CE08756FF9DFEF8FC311CD0B03D" authorityName="Rabb" authorityYear="1958" box="[276,395,1015,1041]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="446" phylum="Chordata" rank="species" species="cracens">
<emphasis id="D58EAB718756FF9DFEF8FC311C7CB03C" box="[276,295,1015,1041]" italics="true" pageId="13" pageNumber="446">C</emphasis>
. 
<emphasis id="D58EAB718756FF9DFED8FC311CD0B03D" box="[308,395,1015,1040]" italics="true" pageId="13" pageNumber="446">cracens</emphasis>
</taxonomicName>
, 
<taxonomicName id="20FA0CE08756FF9DFE75FC311FF0B03C" baseAuthorityName="Walker" baseAuthorityYear="1955" box="[409,683,1015,1041]" class="Amphibia" family="Plethodontidae" genus="Aquiloeurycea" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="446" phylum="Chordata" rank="species" species="scandens">
<emphasis id="D58EAB718756FF9DFE75FC311FF0B03C" box="[409,683,1015,1041]" italics="true" pageId="13" pageNumber="446">Aquiloeurycea scandens</emphasis>
</taxonomicName>
, and 
<taxonomicName id="20FA0CE08756FF9DFD04FC311988B03C" authority="(Rovito &amp; Parra-Olea 2015)" baseAuthorityName="Rovito &amp; Parra-Olea" baseAuthorityYear="2015" box="[744,1235,1014,1041]" class="Amphibia" family="Plethodontidae" genus="Isthmura" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="446" phylum="Chordata" rank="species" species="bellii">
<emphasis id="D58EAB718756FF9DFD04FC311ED5B03C" box="[744,910,1015,1041]" italics="true" pageId="13" pageNumber="446">Isthmura bellii</emphasis>
(
<bibRefCitation id="836B0A928756FF9DFC70FC311990B03C" author="Rovito, S. M. &amp; Parra-Olea, G." box="[924,1227,1014,1041]" pageId="13" pageNumber="446" pagination="57 - 74" refId="ref11456" refString="Rovito, S. M. &amp; Parra-Olea, G. (2015) Two new species of Chiropterotriton (Caudata: Plethodontidae) from northern Mexico. Zootaxa, 4048 (1), 57 - 74. https: // doi. org / 10.11646 / zootaxa. 4048.1.3" type="journal article" year="2015">Rovito &amp; Parra-Olea 2015</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="B38527EB8756FF9DFF7BF87C1834B3FF" ID-DOI="http://doi.org/10.5281/zenodo.12190355" ID-Zenodo-Dep="12190355" box="[151,1391,1978,2002]" httpUri="https://zenodo.org/record/12190355/files/figure.png" pageId="13" pageNumber="446" startId="13.[151,250,1978,2002]" targetBox="[249,1339,1137,1954]" targetPageId="13" targetType="figure">
<paragraph id="E74577638756FF9DFF7BF87C1834B3FF" blockId="13.[151,1391,1978,2002]" box="[151,1391,1978,2002]" pageId="13" pageNumber="446">
<emphasis id="D58EAB718756FF9DFF7BF87C1C48B3FF" bold="true" box="[151,275,1978,2002]" pageId="13" pageNumber="446">FIGURE 5.</emphasis>
Similar habitat where 
<taxonomicName id="20FA0CE08756FF9DFE16F87C1E5AB3FF" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[506,769,1978,2002]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="446" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718756FF9DFE16F87C1E5AB3FF" box="[506,769,1978,2002]" italics="true" pageId="13" pageNumber="446">Eleutherodactylus coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718756FF9DFCEBF87C1E0CB3FF" bold="true" box="[775,855,1978,2002]" pageId="13" pageNumber="446">
<taxonomicNameLabel id="CEBD160A8756FF9DFCEBF87C1E0CB3FF" box="[775,855,1978,2002]" pageId="13" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
individuals were found. Photo: Omar Becerra Soria.
</paragraph>
</caption>
<paragraph id="E74577638755FF9EFF2BFF501F81B4D5" blockId="14.[151,1437,150,321]" pageId="14" pageNumber="447">
<emphasis id="D58EAB718755FF9EFF2BFF501CE6B49D" bold="true" box="[199,445,150,176]" pageId="14" pageNumber="447">Conservation status.</emphasis>
We recommend that 
<taxonomicName id="20FA0CE08755FF9EFD43FF511E7BB49D" authority="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea, 2024" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[687,800,150,177]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="447" phylum="Chordata" rank="species" species="coelum" status="sp. nov.">
<emphasis id="D58EAB718755FF9EFD43FF511F9BB49D" box="[687,704,151,176]" italics="true" pageId="14" pageNumber="447">E</emphasis>
. 
<emphasis id="D58EAB718755FF9EFD23FF501E7BB49D" box="[719,800,150,176]" italics="true" pageId="14" pageNumber="447">coelum</emphasis>
</taxonomicName>
<emphasis id="D58EAB718755FF9EFCC4FF501ED8B49D" bold="true" box="[808,899,150,176]" pageId="14" pageNumber="447">
<taxonomicNameLabel id="CEBD160A8755FF9EFCC4FF501ED8B49D" box="[808,899,150,176]" pageId="14" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
it be designated as Critically Endangered (CR) based on criterion B1ab(iv) (extent of occurrence &lt;
<quantity id="2002DA868755FF9EFD23FF7C1E7FB4F9" box="[719,804,186,212]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="447" unit="km" value="100.0">100 km</quantity>
<superScript id="108FDA2B8755FF9EFCCFFF7C1E70B4E5" attach="left" box="[803,811,186,200]" fontSize="6" pageId="14" pageNumber="447">2</superScript>
, and number of localities) of the International Union for the Conservation of Nature Red List (
<bibRefCitation id="836B0A928755FF9EFDD1FF181F95B4D5" author="IUCN Standards &amp; Petitions Committee" box="[573,718,222,248]" pageId="14" pageNumber="447" refId="ref10240" refString="IUCN Standards and Petitions Committee. (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 25 September 2023)" type="book" year="2022">IUCN, 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AFE024E88755FF9EFF2BFEC41ECCB56D" pageId="14" pageNumber="447" type="etymology">
<paragraph id="E74577638755FF9EFF2BFEC41ECCB56D" blockId="14.[151,1437,150,321]" pageId="14" pageNumber="447">
<emphasis id="D58EAB718755FF9EFF2BFEC41C15B531" bold="true" box="[199,334,258,284]" pageId="14" pageNumber="447">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="20FA0CE08755FF9EFDD2FEC41FD4B531" authorityName="Hernández-Austria &amp; García-Castillo &amp; Parra-Olea" authorityYear="2024" box="[574,655,258,284]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="447" phylum="Chordata" rank="species" species="coelum">
<emphasis id="D58EAB718755FF9EFDD2FEC41FD4B531" box="[574,655,258,284]" italics="true" pageId="14" pageNumber="447">coelum</emphasis>
</taxonomicName>
(in Latin) means heaven in allusion to the Reserva de la Biosfera El Cielo, where the species occurs (El Cielo means heaven in Spanish).
</paragraph>
</subSubSection>
</treatment>
</document>