426 lines
54 KiB
XML
426 lines
54 KiB
XML
<document id="5CA24CE595EE78E807700D8454CFFC54" ID-CLB-Dataset="5979" ID-DOI="10.11646/zootaxa.4990.2.12" ID-GBIF-Dataset="9538fac8-39c0-4fe6-90d5-4b86b118d0d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5026053" ID-ZooBank="62188F0F-D601-4B40-B1BE-68811367D0AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624537475477" checkinUser="plazi" docAuthor="Fuhrmann, Juares & Smith, Andrew B. T." docDate="2021" docId="6A4C87E4FF81652E16C947C6FD6E750E" docLanguage="en" docName="zootaxa.4990.2.12.pdf" docOrigin="Zootaxa 4990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Compsodactylus mendax Fuhrmann & Smith 2021, new species" docType="treatment" docVersion="5" lastPageNumber="391" masterDocId="9675FF9CFF80652A165E4545FFBE7327" masterDocTitle="Compsodactylus Fuhrmann, 2012 (Coleoptera: Scarabaeidae: Melolonthinae) distributional notes and a new species" masterLastPageNumber="393" masterPageNumber="387" pageNumber="388" updateTime="1699020239842" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="53DFC216CE6222AF9780B042E7CD453B">Compsodactylus Fuhrmann, 2012 (Coleoptera: Scarabaeidae: Melolonthinae) distributional notes and a new species</mods:title>
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<mods:name id="AF9C1971E125BBA72E62F2A8797EE93A" type="personal">
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<mods:namePart id="A917335A00065DC76C63CA206AA676AC">Fuhrmann, Juares</mods:namePart>
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<mods:affiliation id="51998560F49952C2E768694347D4021C">Museu de Zoologia da Universidade de São Paulo, 04218 - 970 São Paulo-SP, Brazil.</mods:affiliation>
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<mods:namePart id="CF69F7E603F2FC9EDD93A080E9BFF912">Smith, Andrew B. T.</mods:namePart>
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<mods:affiliation id="0ADB31F23A181A8156387296EF02A1DA">Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada.</mods:affiliation>
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<mods:title id="A9F0D7766347C311344E9D094C934D11">Zootaxa</mods:title>
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<mods:date id="CD21AD36F6B0EF8F23B6D6EC6524FFE1">2021</mods:date>
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<mods:number id="4A3F267A1B81570980E4CA9CBE0218F5">2021-06-21</mods:number>
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<treatment id="6A4C87E4FF81652E16C947C6FD6E750E" ID-DOI="http://doi.org/10.5281/zenodo.5026061" ID-GBIF-Taxon="183124986" ID-Zenodo-Dep="5026061" LSID="urn:lsid:plazi:treatment:6A4C87E4FF81652E16C947C6FD6E750E" httpUri="http://treatment.plazi.org/id/6A4C87E4FF81652E16C947C6FD6E750E" lastPageId="4" lastPageNumber="391" pageId="1" pageNumber="388">
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<subSubSection id="AAFF6579FF81652B16C947C6FCDD71B9" box="[151,867,643,670]" pageId="1" pageNumber="388" type="nomenclature">
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<paragraph id="E25A36F2FF81652B16C947C6FCDD71B9" blockId="1.[151,867,643,707]" box="[151,867,643,670]" pageId="1" pageNumber="388">
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<heading id="B912819EFF81652B16C947C6FCDD71B9" bold="true" box="[151,867,643,670]" fontSize="11" level="1" pageId="1" pageNumber="388" reason="1">
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<emphasis id="D091EAE0FF81652B16C947C6FCDD71B9" bold="true" box="[151,867,643,670]" pageId="1" pageNumber="388">
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<taxonomicName id="25E54D71FF81652B16C947C6FD7971B9" authority="Fuhrmann & Smith" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[151,711,643,670]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="mendax" status="sp. nov.">
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<emphasis id="D091EAE0FF81652B16C947C6FE7A71B9" bold="true" box="[151,452,643,670]" italics="true" pageId="1" pageNumber="388">Compsodactylus mendax</emphasis>
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Fuhrmann & Smith
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</taxonomicName>
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,
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<taxonomicNameLabel id="CBA2579BFF81652B148D47C1FCDD71B9" box="[723,867,644,670]" pageId="1" pageNumber="388" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="AAFF6579FF81652B16C947EDFE0171E4" box="[151,447,680,707]" pageId="1" pageNumber="388" type="description">
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<paragraph id="E25A36F2FF81652B16C947EDFE0171E4" blockId="1.[151,867,643,707]" box="[151,447,680,707]" pageId="1" pageNumber="388">
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(
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<figureCitation id="7ADE2A77FF81652B16C147EDFEB271E5" box="[159,268,680,706]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="1" pageNumber="388">Figs. 1−8</figureCitation>
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,
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||
<figureCitation id="7ADE2A77FF81652B174747EDFEDD71E4" box="[281,355,680,707]" captionStart="FIGURES 9−18" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[164,1405,198,1843]" captionTargetId="figure-17@3.[151,1435,181,1864]" captionTargetPageId="3" captionText="FIGURES 9−18. Compsodactylus, posterior leg of males; 9−10, C. argentinus (femur-tibia, apex of tibia); 11−12, C. parvulus (femur-tibia, apex of tibia); 13−14, C. martinezi (femur-tibia, apex of tibia); 15−16, C. mendax new species (femur-tibia, apex of tibia); 17−18, C. vallejoi (femur-tibia, apex of tibia; modified from Figueroa & Neita-Moreno 2019). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="1" pageNumber="388">15−16</figureCitation>
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,
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<figureCitation id="7ADE2A77FF81652B173047EDFE0971E4" box="[366,439,680,707]" captionStart-0="FIGURES 19−25" captionStart-1="FIGURE 26" captionStartId-0="4.[151,264,1119,1143]" captionStartId-1="5.[151,250,1630,1654]" captionTargetBox-0="[165,1421,187,1088]" captionTargetBox-1="[158,1431,732,1602]" captionTargetId-0="figure-22@4.[160,1427,181,1096]" captionTargetId-1="figure-916@5.[151,1436,726,1607]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 19−25. Compsodactylus mendax new species; 19, parameres, distal; 20−21, aedeagus (lateral, laterodistal), 22, genital ring of males; 23−26, terminalia of females (dorsal, lateral, ventral). gcd, distal piece of gonocoxite; gcp; proximal piece of gonocoxite; ppt, paraprocts; gst, gonostyle; spc, spiculum gastrale; st9, abdominal sternite IX; tg9, abdominal tergite IX. Scale = 0.5 mm." captionText-1="FIGURE 26. Compsodactylus, map of species records. Scale = 500 km." figureDoi-0="http://doi.org/10.5281/zenodo.5026065" figureDoi-1="http://doi.org/10.5281/zenodo.5026067" httpUri-0="https://zenodo.org/record/5026065/files/figure.png" httpUri-1="https://zenodo.org/record/5026067/files/figure.png" pageId="1" pageNumber="388">19−26</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="AAFF6579FF81652B16C947B5FB557075" pageId="1" pageNumber="388" type="materials_examined">
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<paragraph id="E25A36F2FF81652B16C947B5FB557075" blockId="1.[151,1437,752,2038]" pageId="1" pageNumber="388">
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<emphasis id="D091EAE0FF81652B16C947B5FEFF702D" bold="true" box="[151,321,752,778]" pageId="1" pageNumber="388">Type material</emphasis>
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||
.
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||
<materialsCitation id="528D3CAFFF81652B170C47B5FB597075" ID-GBIF-Occurrence="3309805305" collectingDate="2011-03-30" collectingDateMax="2011-03-31" collectingDateMin="2011-03-30" collectionCode="FSCA, CMNC" collectorName="J. B. Heppner & C. Carrera" country="Peru" elevation="1035" location="Figs" municipality="Male" pageId="1" pageNumber="388" specimenCount="6" stateProvince="Lima" typeStatus="holotype">
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<collectingMunicipality id="023EAC88FF81652B170C47B5FE32702D" box="[338,396,752,778]" pageId="1" pageNumber="388">Male</collectingMunicipality>
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<typeStatus id="3D5E8850FF81652B17CB47B5FE49702D" box="[405,503,752,778]" pageId="1" pageNumber="388" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="84F4AE37FF81652B145747B5FDED702D" box="[521,595,752,778]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="388" type="Museum">FSCA</collectionCode>
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,
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<figureCitation id="7ADE2A77FF81652B143F47B5FD6E702D" box="[609,720,752,778]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="1" pageNumber="388">Figs. 1−4</figureCitation>
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),
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||
<typeStatus id="3D5E8850FF81652B14B647B5FCFC702D" box="[744,834,752,778]" pageId="1" pageNumber="388" type="allotype">allotype</typeStatus>
|
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(
|
||
<collectionCode id="84F4AE37FF81652B150A47B5FC20702D" box="[852,926,752,778]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="388" type="Museum">FSCA</collectionCode>
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,
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||
<location id="E73A6029FF81652B15F347B5FC5F702D" LSID="urn:lsid:plazi:treatment:6A4C87E4FF81652E16C947C6FD6E750E:E73A6029FF81652B15F347B5FC5F702D" box="[941,993,752,778]" country="Peru" municipality="Male" name="Figs" pageId="1" pageNumber="388" stateProvince="Lima">Figs</location>
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; 6−8), and male
|
||
<typeStatus id="3D5E8850FF81652B12F947B4FAB9702C" box="[1191,1287,753,779]" pageId="1" pageNumber="388" type="paratype">paratype</typeStatus>
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(
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||
<collectionCode id="84F4AE37FF81652B134747B5FAD1702D" box="[1305,1391,752,778]" pageId="1" pageNumber="388">CMNC</collectionCode>
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||
) all labeled (
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||
<figureCitation id="7ADE2A77FF81652B16A64651FE837008" box="[248,317,788,815]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="1" pageNumber="388">Fig. 5</figureCitation>
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||
): “
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||
<collectingCountry id="9AF27662FF81652B17004651FE127009" box="[350,428,788,814]" name="Peru" pageId="1" pageNumber="388">PERU</collectingCountry>
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||
: Dept.
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||
<collectingRegion id="2021F810FF81652B17A24651FD897009" box="[508,567,788,814]" country="Peru" name="Lima" pageId="1" pageNumber="388">Lima</collectingRegion>
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/ Sta. Eulalia
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<quantity id="251D9B17FF81652B14864651FC987008" box="[728,806,788,815]" metricMagnitude="3" metricUnit="m" metricValue="1.035" pageId="1" pageNumber="388" unit="m" value="1035.0">
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<elevation id="69C8D1C1FF81652B14864651FC987008" box="[728,806,788,815]" metricMagnitude="3" metricUnit="m" metricValue="1.035" pageId="1" pageNumber="388" unit="m" value="1035.0">1035m</elevation>
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</quantity>
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/
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<date id="965B1032FF81652B15634651FC447009" box="[829,1018,788,815]" pageId="1" pageNumber="388" value="2011-03-30" valueMax="2011-03-31" valueMin="2011-03-30">
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<collectingDate id="861FE9DAFF81652B15634651FC447009" box="[829,1018,788,815]" pageId="1" pageNumber="388" value="2011-03-30" valueMax="2011-03-31" valueMin="2011-03-30">30−31 Mar 2011</collectingDate>
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</date>
|
||
/
|
||
<collectorName id="4F105324FF81652B124F4650FB1A7008" box="[1041,1188,789,815]" pageId="1" pageNumber="388">J.B. Heppner</collectorName>
|
||
&
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||
<collectorName id="4F105324FF81652B12944651FAFB7009" box="[1226,1349,788,814]" pageId="1" pageNumber="388">C. Carrera</collectorName>
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||
”. These specimens also have our corresponding red
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||
<typeStatus id="3D5E8850FF81652B14DF467DFD5D7075" box="[641,739,824,850]" pageId="1" pageNumber="388" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="3D5E8850FF81652B1544467DFCCA7075" box="[794,884,824,850]" pageId="1" pageNumber="388" type="allotype">allotype</typeStatus>
|
||
labels and yellow
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||
<typeStatus id="3D5E8850FF81652B1218467CFB187074" box="[1094,1190,825,851]" pageId="1" pageNumber="388" type="paratype">paratype</typeStatus>
|
||
label
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AAFF6579FF81652B16994619FC657720" pageId="1" pageNumber="388" type="diagnosis">
|
||
<paragraph id="E25A36F2FF81652B16994619FC657720" blockId="1.[151,1437,752,2038]" pageId="1" pageNumber="388">
|
||
<emphasis id="D091EAE0FF81652B16994619FE847051" bold="true" box="[199,314,860,886]" pageId="1" pageNumber="388">Diagnosis</emphasis>
|
||
. Length greater than 8.0 mm; dorsal surface with metallic green reflections; antennae with 10 antennomers; clypeus elongate with expanded apex in males (
|
||
<figureCitation id="7ADE2A77FF81652B150546C5FC1C70BD" box="[859,930,896,922]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="1" pageNumber="388">Fig. 4</figureCitation>
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||
); clypeus with short, spiny setae along apex and lateral edges in males; apex of protarsomere I 2 times wider than apex of protarsomere II in males; elytral striae distinctly punctate; male metatibial apex with an inner truncate, twisted process (
|
||
<figureCitation id="7ADE2A77FF81652B1241468DFBCD70C4" box="[1055,1139,968,995]" captionStart="FIGURES 9−18" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[164,1405,198,1843]" captionTargetId="figure-17@3.[151,1435,181,1864]" captionTargetPageId="3" captionText="FIGURES 9−18. Compsodactylus, posterior leg of males; 9−10, C. argentinus (femur-tibia, apex of tibia); 11−12, C. parvulus (femur-tibia, apex of tibia); 13−14, C. martinezi (femur-tibia, apex of tibia); 15−16, C. mendax new species (femur-tibia, apex of tibia); 17−18, C. vallejoi (femur-tibia, apex of tibia; modified from Figueroa & Neita-Moreno 2019). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="1" pageNumber="388">Fig. 16</figureCitation>
|
||
); genitalia of females with gonostyle, and gonocoxite about 2.5 times longer than wide (
|
||
<figureCitation id="7ADE2A77FF81652B151D46A9FC717720" box="[835,975,1004,1031]" captionStart="FIGURES 19−25" captionStartId="4.[151,264,1119,1143]" captionTargetBox="[165,1421,187,1088]" captionTargetId="figure-22@4.[160,1427,181,1096]" captionTargetPageId="4" captionText="FIGURES 19−25. Compsodactylus mendax new species; 19, parameres, distal; 20−21, aedeagus (lateral, laterodistal), 22, genital ring of males; 23−26, terminalia of females (dorsal, lateral, ventral). gcd, distal piece of gonocoxite; gcp; proximal piece of gonocoxite; ppt, paraprocts; gst, gonostyle; spc, spiculum gastrale; st9, abdominal sternite IX; tg9, abdominal tergite IX. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5026065" httpUri="https://zenodo.org/record/5026065/files/figure.png" pageId="1" pageNumber="388">Figs. 23−25</figureCitation>
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).
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AAFF6579FF81652B16994155FDB676FC" pageId="1" pageNumber="388" type="materials_examined">
|
||
<paragraph id="E25A36F2FF81652B16994155FDB676FC" blockId="1.[151,1437,752,2038]" pageId="1" pageNumber="388">
|
||
<emphasis id="D091EAE0FF81652B16994155FE7D770D" bold="true" box="[199,451,1040,1066]" pageId="1" pageNumber="388">
|
||
<typeStatus id="3D5E8850FF81652B16994155FE8A770D" box="[199,308,1040,1066]" pageId="1" pageNumber="388" type="holotype">Holotype</typeStatus>
|
||
description
|
||
</emphasis>
|
||
. Length 9.0 mm, width 4.0 mm at the middle of elytra. Dorsal and ventral surface of head and thorax (except lateral edges of pronotum) black with metallic green reflections (
|
||
<figureCitation id="7ADE2A77FF81652B12204171FB557769" box="[1150,1259,1076,1102]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="1" pageNumber="388">Figs. 1−4</figureCitation>
|
||
); dorsal surface of elytra and lateral edges of pronotum yellowish brown with light metallic green reflections; ventral surface of abdomen shiny reddish brown, ventrite
|
||
<collectionCode id="84F4AE37FF81652B14124138FDD477B0" box="[588,618,1149,1175]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="388">VI</collectionCode>
|
||
with anterior dark stripe; legs bicolored, reddish brown with dark sections and markings. Head with clypeus elongate, strongly expanded at apex (
|
||
<figureCitation id="7ADE2A77FF81652B15E641E5FC42779D" box="[952,1020,1184,1210]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="1" pageNumber="388">Fig. 4</figureCitation>
|
||
), surface glabrous medially and with short, spiny setae along apex and lateral edges. Frons surface with rugose punctures and densely setose, setae long, thin. Interocular distance 5 times wider than dorsal eye width. Antenna with 10 antennomeres, club longer than antennomeres II–
|
||
<collectionCode id="84F4AE37FF81652B17044048FEC97600" box="[346,375,1293,1319]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="388">VI</collectionCode>
|
||
combined. Pronotum surface rugose with long, dense setae, anterior margin with some minute, spiny setae. Elytra surface with moderately dense setae, striae distinctly punctate. Pygidium and ventral surface with dense punctures and hair-like setae. Protibia with 2 outer teeth, protarsomere I greatly enlarged with a padlike granulate ventral surface. Metafemur not strongly enlarged (about 2.1−2.4 times longer than wide,
|
||
<figureCitation id="7ADE2A77FF81652B1364403DFA3076B4" box="[1338,1422,1400,1427]" captionStart="FIGURES 9−18" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[164,1405,198,1843]" captionTargetId="figure-17@3.[151,1435,181,1864]" captionTargetPageId="3" captionText="FIGURES 9−18. Compsodactylus, posterior leg of males; 9−10, C. argentinus (femur-tibia, apex of tibia); 11−12, C. parvulus (femur-tibia, apex of tibia); 13−14, C. martinezi (femur-tibia, apex of tibia); 15−16, C. mendax new species (femur-tibia, apex of tibia); 17−18, C. vallejoi (femur-tibia, apex of tibia; modified from Figueroa & Neita-Moreno 2019). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="1" pageNumber="388">Fig. 15</figureCitation>
|
||
), inner margin without spines or carina and with some spiny setae. Metatibial apex with an inner, twisted process; metatibial spurs absent (
|
||
<figureCitation id="7ADE2A77FF81652B17F94085FE4576FC" box="[423,507,1472,1499]" captionStart="FIGURES 9−18" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[164,1405,198,1843]" captionTargetId="figure-17@3.[151,1435,181,1864]" captionTargetPageId="3" captionText="FIGURES 9−18. Compsodactylus, posterior leg of males; 9−10, C. argentinus (femur-tibia, apex of tibia); 11−12, C. parvulus (femur-tibia, apex of tibia); 13−14, C. martinezi (femur-tibia, apex of tibia); 15−16, C. mendax new species (femur-tibia, apex of tibia); 17−18, C. vallejoi (femur-tibia, apex of tibia; modified from Figueroa & Neita-Moreno 2019). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="1" pageNumber="388">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AAFF6579FF81652B169940A1FB5A7594" pageId="1" pageNumber="388" type="description">
|
||
<paragraph id="E25A36F2FF81652B169940A1FB5A7594" blockId="1.[151,1437,752,2038]" pageId="1" pageNumber="388">
|
||
<emphasis id="D091EAE0FF81652B169940A1FE8676D9" bold="true" box="[199,312,1508,1534]" pageId="1" pageNumber="388">Variation</emphasis>
|
||
. The
|
||
<typeStatus id="3D5E8850FF81652B172040A1FE6676D9" box="[382,472,1508,1534]" pageId="1" pageNumber="388" type="allotype">allotype</typeStatus>
|
||
is
|
||
<quantity id="251D9B17FF81652B17A140A1FDE676D8" box="[511,600,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="388" unit="mm" value="8.5">8.5 mm</quantity>
|
||
in length with the following sexually dimorphic characters: clypeus not elongate, without glabrous area, and with apex subquadrate, not expanded; frons with short, spiny setae; protarsomere I cylindrical, not enlarged; metatibial apex without inner process and with 2 spurs. Female terminalia bearing a pair of elongate paraprocts; gonocoxite about 2.5 longer than wide and divided in a proximal and a distal piece; a lateral plate between paraproct and proximal piece of gonocoxite is present, gonostyle present (
|
||
<figureCitation id="7ADE2A77FF81652B12DF4331FAB075A8" box="[1153,1294,1652,1679]" captionStart="FIGURES 19−25" captionStartId="4.[151,264,1119,1143]" captionTargetBox="[165,1421,187,1088]" captionTargetId="figure-22@4.[160,1427,181,1096]" captionTargetPageId="4" captionText="FIGURES 19−25. Compsodactylus mendax new species; 19, parameres, distal; 20−21, aedeagus (lateral, laterodistal), 22, genital ring of males; 23−26, terminalia of females (dorsal, lateral, ventral). gcd, distal piece of gonocoxite; gcp; proximal piece of gonocoxite; ppt, paraprocts; gst, gonostyle; spc, spiculum gastrale; st9, abdominal sternite IX; tg9, abdominal tergite IX. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5026065" httpUri="https://zenodo.org/record/5026065/files/figure.png" pageId="1" pageNumber="388">Figs. 23−25</figureCitation>
|
||
). The elytral suture is yellow in
|
||
<typeStatus id="3D5E8850FF81652B173243DDFE707595" box="[364,462,1688,1714]" pageId="1" pageNumber="388" type="holotype">holotype</typeStatus>
|
||
and black with metallic green reflections in the
|
||
<typeStatus id="3D5E8850FF81652B15B543DDFBFB7595" box="[1003,1093,1688,1714]" pageId="1" pageNumber="388" type="allotype">allotype</typeStatus>
|
||
and
|
||
<typeStatus id="3D5E8850FF81652B122543DCFB617594" box="[1147,1247,1689,1715]" pageId="1" pageNumber="388" type="paratype">paratype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AAFF6579FF81652B169943F9FD4D75DC" pageId="1" pageNumber="388" type="distribution">
|
||
<paragraph id="E25A36F2FF81652B169943F9FD4D75DC" blockId="1.[151,1437,752,2038]" pageId="1" pageNumber="388">
|
||
<emphasis id="D091EAE0FF81652B169943F9FEE775F1" bold="true" box="[199,345,1724,1750]" pageId="1" pageNumber="388">Distribution</emphasis>
|
||
. The
|
||
<typeStatus id="3D5E8850FF81652B17C443F9FE4175F1" box="[410,511,1724,1750]" pageId="1" pageNumber="388" type="holotype">holotype</typeStatus>
|
||
,
|
||
<typeStatus id="3D5E8850FF81652B145443F9FDD975F1" box="[522,615,1724,1750]" pageId="1" pageNumber="388" type="allotype">allotype</typeStatus>
|
||
, and a single
|
||
<typeStatus id="3D5E8850FF81652B14A143F8FCE175F0" box="[767,863,1725,1751]" pageId="1" pageNumber="388" type="paratype">paratype</typeStatus>
|
||
were collected in Santa Eulalia,
|
||
<collectingRegion id="2021F810FF81652B129143F9FAB475F1" box="[1231,1290,1724,1750]" country="Peru" name="Lima" pageId="1" pageNumber="388">Lima</collectingRegion>
|
||
Department,
|
||
<collectingCountry id="9AF27662FF81652B16C943A4FF7475DC" box="[151,202,1761,1787]" name="Peru" pageId="1" pageNumber="388">Peru</collectingCountry>
|
||
(approximately
|
||
<geoCoordinate id="87D15035FF81652B17DA43A5FE5375DD" box="[388,493,1760,1786]" degrees="11.902" direction="south" orientation="latitude" pageId="1" pageNumber="388" precision="55" value="-11.902">11.902°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="87D15035FF81652B17A643A5FDD375DC" box="[504,621,1760,1787]" degrees="76.666" direction="west" orientation="longitude" pageId="1" pageNumber="388" precision="55" value="-76.666">76.666°W</geoCoordinate>
|
||
) at
|
||
<quantity id="251D9B17FF81652B14C843A5FD5075DC" box="[662,750,1760,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.035" pageId="1" pageNumber="388" unit="m" value="1035.0">1035 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AAFF6579FF81652B16994241FD5D7465" pageId="1" pageNumber="388" type="etymology">
|
||
<paragraph id="E25A36F2FF81652B16994241FD5D7465" blockId="1.[151,1437,752,2038]" pageId="1" pageNumber="388">
|
||
<emphasis id="D091EAE0FF81652B16994241FEF67439" bold="true" box="[199,328,1796,1822]" pageId="1" pageNumber="388">Etymology</emphasis>
|
||
. From the Latin
|
||
<taxonomicName id="25E54D71FF81652B14514241FDD87439" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[527,614,1796,1822]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF81652B14514241FDD87439" box="[527,614,1796,1822]" italics="true" pageId="1" pageNumber="388">mendax</emphasis>
|
||
</taxonomicName>
|
||
meaning liar, in reference to the elongate clypeus being reminiscent of Pinocchio’s nose. This name is a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AAFF6579FF81652E16994209FD6E750E" lastPageId="4" lastPageNumber="391" pageId="1" pageNumber="388" type="discussion">
|
||
<paragraph id="E25A36F2FF81652816994209FC4C73DE" blockId="1.[151,1437,752,2038]" lastBlockId="2.[151,1436,151,249]" lastPageId="2" lastPageNumber="389" pageId="1" pageNumber="388">
|
||
<emphasis id="D091EAE0FF81652B16994209FE8A7441" bold="true" box="[199,308,1868,1894]" pageId="1" pageNumber="388">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="25E54D71FF81652B171F4209FDE67441" authority="Fuhrmann & Smith, 2021" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[321,600,1868,1894]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="mendax" status="sp. nov.">
|
||
<emphasis id="D091EAE0FF81652B171F4209FDE67441" box="[321,600,1868,1894]" italics="true" pageId="1" pageNumber="388">Compsodactylus mendax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CBA2579BFF81652B14004208FD5A7441" box="[606,740,1868,1895]" pageId="1" pageNumber="388" rank="species">new species</taxonomicNameLabel>
|
||
is easily distinguished from the other four species in the genus because the clypeal apex is enlarged in males while the other species have a trapezoidal clypeus (
|
||
<figureCitation id="7ADE2A77FF81652B128B4235FAA474AD" box="[1237,1306,1904,1930]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="1" pageNumber="388">Fig. 4</figureCitation>
|
||
); the dorsal surface has green metallic reflections and antennae have 10 antennomeres in females, a character combination not found in other species.
|
||
<taxonomicName id="25E54D71FF81652B17C042FDFD0B74F5" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[414,693,1976,2002]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF81652B17C042FDFD0B74F5" box="[414,693,1976,2002]" italics="true" pageId="1" pageNumber="388">Compsodactylus mendax</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="25E54D71FF81652B151442FDFBD974F5" authority="(Frey, 1972)" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[842,1127,1976,2003]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="D091EAE0FF81652B151442FDFCE374F5" box="[842,861,1976,2002]" italics="true" pageId="1" pageNumber="388">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF81652B153542FCFC6B74F5" box="[875,981,1977,2002]" italics="true" pageId="1" pageNumber="388">martinezi</emphasis>
|
||
(Frey, 1972)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="25E54D71FF81652B12C142FDFEC374D1" authority="Figueroa & Neita-Moreno, 2019" authorityName="Figueroa & Neita-Moreno" authorityYear="2019" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="vallejoi">
|
||
<emphasis id="D091EAE0FF81652B12C142FDFB0C74F5" box="[1183,1202,1976,2002]" italics="true" pageId="1" pageNumber="388">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF81652B129E42FDFAAA74F5" box="[1216,1300,1976,2002]" italics="true" pageId="1" pageNumber="388">vallejoi</emphasis>
|
||
<bibRefCitation id="86744B03FF81652B134542FDFEC374D1" author="Figueroa, L. & Neita-Moreno, J. C." pageId="1" pageNumber="388" pagination="587 - 591" refId="ref3915" refString="Figueroa, L. & Neita-Moreno, J. C. (2019) A new Compsodactylus Fuhrmann (Coleoptera: Scarabaeidae: Melolonthinae) from Peru. Zootaxa, 4560 (3), 587 - 591. https: // doi. org / 10.11646 / zootaxa. 4560.3.12" type="journal article" year="2019">
|
||
Figueroa & Neita-Moreno,
|
||
<quantity id="251D9B17FF81652B171B4299FEC374D1" box="[325,381,2012,2038]" metricMagnitude="1" metricUnit="m" metricValue="5.12826" pageId="1" pageNumber="388" unit="in" value="2019.0">2019</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in that all three species have the apical spine of metatibia thick and projecting perpendicular to length of the tibiae in males (
|
||
<figureCitation id="7ADE2A77FF826528144C45D2FD237396" box="[530,669,151,177]" captionStart="FIGURES 9−18" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[164,1405,198,1843]" captionTargetId="figure-17@3.[151,1435,181,1864]" captionTargetPageId="3" captionText="FIGURES 9−18. Compsodactylus, posterior leg of males; 9−10, C. argentinus (femur-tibia, apex of tibia); 11−12, C. parvulus (femur-tibia, apex of tibia); 13−14, C. martinezi (femur-tibia, apex of tibia); 15−16, C. mendax new species (femur-tibia, apex of tibia); 17−18, C. vallejoi (femur-tibia, apex of tibia; modified from Figueroa & Neita-Moreno 2019). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="2" pageNumber="389">Figs. 13−18</figureCitation>
|
||
) and females have gonostyles.
|
||
<taxonomicName id="25E54D71FF826528125245D2FAF87397" baseAuthorityName="Moser" baseAuthorityYear="1919" box="[1036,1350,151,177]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="argentinus">
|
||
<emphasis id="D091EAE0FF826528125245D2FAF87397" box="[1036,1350,151,177]" italics="true" pageId="2" pageNumber="389">Compsodactylus argentinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="25E54D71FF82652813DD45D2FF4773F2" baseAuthorityName="Frey" baseAuthorityYear="1970" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="D091EAE0FF82652813DD45D2FA287396" box="[1411,1430,151,177]" italics="true" pageId="2" pageNumber="389">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF82652816C945FEFF4773F2" box="[151,249,187,213]" italics="true" pageId="2" pageNumber="389">parvulus</emphasis>
|
||
</taxonomicName>
|
||
differ from these three species in that the apical spine of metatibia is thin and projecting parallel to the length of the tibia in males (
|
||
<figureCitation id="7ADE2A77FF826528178C459AFDF273DE" box="[466,588,223,249]" captionStart="FIGURES 9−18" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[164,1405,198,1843]" captionTargetId="figure-17@3.[151,1435,181,1864]" captionTargetPageId="3" captionText="FIGURES 9−18. Compsodactylus, posterior leg of males; 9−10, C. argentinus (femur-tibia, apex of tibia); 11−12, C. parvulus (femur-tibia, apex of tibia); 13−14, C. martinezi (femur-tibia, apex of tibia); 15−16, C. mendax new species (femur-tibia, apex of tibia); 17−18, C. vallejoi (femur-tibia, apex of tibia; modified from Figueroa & Neita-Moreno 2019). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="2" pageNumber="389">Figs. 9−12</figureCitation>
|
||
) and females do not have gonostyles.
|
||
</paragraph>
|
||
<caption id="B69A667AFF82652816C942FEFCFF74D0" ID-DOI="http://doi.org/10.5281/zenodo.5026059" ID-Zenodo-Dep="5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="2" pageNumber="389" startId="2.[151,264,1979,2003]" targetBox="[151,1436,270,1953]" targetPageId="2">
|
||
<paragraph id="E25A36F2FF82652816C942FEFCFF74D0" blockId="2.[151,1436,1979,2039]" pageId="2" pageNumber="389">
|
||
<emphasis id="D091EAE0FF82652816C942FEFE8674F4" bold="true" box="[151,312,1979,2003]" pageId="2" pageNumber="389">FIGURES 1−8</emphasis>
|
||
.
|
||
<taxonomicName id="25E54D71FF826528171842FEFDFB74F4" authority="Fuhrmann & Smith, 2021" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[326,581,1979,2003]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="mendax" status="sp. nov.">
|
||
<emphasis id="D091EAE0FF826528171842FEFDFB74F4" box="[326,581,1979,2003]" italics="true" pageId="2" pageNumber="389">Compsodactylus mendax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CBA2579BFF826528141342F9FD7574F4" box="[589,715,1979,2003]" pageId="2" pageNumber="389" rank="species">new species</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="D091EAE0FF826528148642FEFCBE74F4" bold="true" box="[728,768,1979,2003]" pageId="2" pageNumber="389">1−5</emphasis>
|
||
, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label);
|
||
<emphasis id="D091EAE0FF82652816C9429AFF0174D0" bold="true" box="[151,191,2015,2039]" pageId="2" pageNumber="389">6−8</emphasis>
|
||
, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B69A667AFF83652916C9421AFAE07498" ID-DOI="http://doi.org/10.5281/zenodo.5026063" ID-Zenodo-Dep="5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="3" pageNumber="390" startId="3.[151,264,1887,1911]" targetBox="[164,1405,198,1843]" targetPageId="3">
|
||
<paragraph id="E25A36F2FF83652916C9421AFAE07498" blockId="3.[151,1437,1887,1984]" pageId="3" pageNumber="390">
|
||
<emphasis id="D091EAE0FF83652916C9421AFEFD7450" bold="true" box="[151,323,1887,1911]" pageId="3" pageNumber="390">FIGURES 9−18</emphasis>
|
||
.
|
||
<taxonomicName id="25E54D71FF83652917114225FE48745F" authorityName="Fuhrmann" authorityYear="2012" box="[335,502,1888,1912]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D091EAE0FF83652917114225FE48745F" box="[335,502,1888,1912]" italics="true" pageId="3" pageNumber="390">Compsodactylus</emphasis>
|
||
</taxonomicName>
|
||
, posterior leg of males;
|
||
<emphasis id="D091EAE0FF83652914B3421AFC9C7450" bold="true" box="[749,802,1887,1911]" pageId="3" pageNumber="390">9−10</emphasis>
|
||
,
|
||
<taxonomicName id="25E54D71FF83652915714225FC097450" baseAuthorityName="Moser" baseAuthorityYear="1919" box="[815,951,1888,1911]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="argentinus">
|
||
<emphasis id="D091EAE0FF83652915714225FCFE7450" box="[815,832,1888,1911]" italics="true" pageId="3" pageNumber="390">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF83652915124225FC097450" box="[844,951,1888,1911]" italics="true" pageId="3" pageNumber="390">argentinus</emphasis>
|
||
</taxonomicName>
|
||
(femur-tibia, apex of tibia);
|
||
<emphasis id="D091EAE0FF8365291287421AFAA77450" bold="true" box="[1241,1305,1887,1911]" pageId="3" pageNumber="390">11−12</emphasis>
|
||
,
|
||
<taxonomicName id="25E54D71FF83652913784225FA23745F" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[1318,1437,1888,1912]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="D091EAE0FF83652913784225FA897450" box="[1318,1335,1888,1911]" italics="true" pageId="3" pageNumber="390">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF836529131A4225FA23745F" box="[1348,1437,1888,1912]" italics="true" pageId="3" pageNumber="390">parvulus</emphasis>
|
||
</taxonomicName>
|
||
(femur-tibia, apex of tibia);
|
||
<emphasis id="D091EAE0FF83652917EA42C6FE4B74BC" bold="true" box="[436,501,1923,1947]" pageId="3" pageNumber="390">13−14</emphasis>
|
||
,
|
||
<taxonomicName id="25E54D71FF836529145C42C1FD3E74BC" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[514,640,1924,1947]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="D091EAE0FF836529145C42C1FDAD74BC" box="[514,531,1924,1947]" italics="true" pageId="3" pageNumber="390">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF836529147E42C1FD3E74BC" box="[544,640,1924,1947]" italics="true" pageId="3" pageNumber="390">martinezi</emphasis>
|
||
</taxonomicName>
|
||
(femur-tibia, apex of tibia);
|
||
<emphasis id="D091EAE0FF83652915FD42C6FC5A74BC" bold="true" box="[931,996,1923,1947]" pageId="3" pageNumber="390">15−16</emphasis>
|
||
,
|
||
<taxonomicName id="25E54D71FF83652915AF42C1FBE074BB" authority="Fuhrmann & Smith, 2021" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[1009,1118,1924,1948]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="mendax" status="sp. nov.">
|
||
<emphasis id="D091EAE0FF83652915AF42C1FBBC74BC" box="[1009,1026,1924,1947]" italics="true" pageId="3" pageNumber="390">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF836529125142C1FBE074BB" box="[1039,1118,1924,1948]" italics="true" pageId="3" pageNumber="390">mendax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CBA2579BFF836529123B42C1FB6174BB" box="[1125,1247,1924,1948]" pageId="3" pageNumber="390" rank="species">new species</taxonomicNameLabel>
|
||
(femur-tibia, apex of tibia);
|
||
<emphasis id="D091EAE0FF83652916AB42E2FE897498" bold="true" box="[245,311,1959,1983]" pageId="3" pageNumber="390">17−18</emphasis>
|
||
,
|
||
<taxonomicName id="25E54D71FF836529171D42EDFE1374E7" authorityName="Figueroa & Neita-Moreno" authorityYear="2019" box="[323,429,1960,1984]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="vallejoi">
|
||
<emphasis id="D091EAE0FF836529171D42EDFEEA7498" box="[323,340,1960,1983]" italics="true" pageId="3" pageNumber="390">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF836529173F42EDFE1374E7" box="[353,429,1960,1984]" italics="true" pageId="3" pageNumber="390">vallejoi</emphasis>
|
||
</taxonomicName>
|
||
(femur-tibia, apex of tibia; modified from
|
||
<bibRefCitation id="86744B03FF836529150142EDFB1874E7" author="Figueroa, L. & Neita-Moreno, J. C." box="[863,1190,1960,1984]" pageId="3" pageNumber="390" pagination="587 - 591" refId="ref3915" refString="Figueroa, L. & Neita-Moreno, J. C. (2019) A new Compsodactylus Fuhrmann (Coleoptera: Scarabaeidae: Melolonthinae) from Peru. Zootaxa, 4560 (3), 587 - 591. https: // doi. org / 10.11646 / zootaxa. 4560.3.12" type="journal article" year="2019">Figueroa & Neita-Moreno 2019</bibRefCitation>
|
||
). Scale = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B69A667AFF84652E16C9411AFE8077C4" ID-DOI="http://doi.org/10.5281/zenodo.5026065" ID-Zenodo-Dep="5026065" httpUri="https://zenodo.org/record/5026065/files/figure.png" pageId="4" pageNumber="391" startId="4.[151,264,1119,1143]" targetBox="[165,1421,187,1088]" targetPageId="4">
|
||
<paragraph id="E25A36F2FF84652E16C9411AFE8077C4" blockId="4.[151,1437,1119,1252]" pageId="4" pageNumber="391">
|
||
<emphasis id="D091EAE0FF84652E16C9411AFEED7750" bold="true" box="[151,339,1119,1143]" pageId="4" pageNumber="391">FIGURES 19−25</emphasis>
|
||
.
|
||
<taxonomicName id="25E54D71FF84652E173D4125FDDD775F" authority="Fuhrmann & Smith, 2021" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[355,611,1120,1144]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="mendax" status="sp. nov.">
|
||
<emphasis id="D091EAE0FF84652E173D4125FDDD775F" box="[355,611,1120,1144]" italics="true" pageId="4" pageNumber="391">Compsodactylus mendax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CBA2579BFF84652E14334125FD53775F" box="[621,749,1120,1144]" pageId="4" pageNumber="391" rank="species">new species</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="D091EAE0FF84652E14A5411AFCAA7750" bold="true" box="[763,788,1119,1143]" pageId="4" pageNumber="391">19</emphasis>
|
||
, parameres, distal;
|
||
<emphasis id="D091EAE0FF84652E15BA411AFB9B7750" bold="true" box="[996,1061,1119,1143]" pageId="4" pageNumber="391">20−21</emphasis>
|
||
, aedeagus (lateral, laterodistal),
|
||
<emphasis id="D091EAE0FF84652E1323411AFA287750" bold="true" box="[1405,1430,1119,1143]" pageId="4" pageNumber="391">22</emphasis>
|
||
, genital ring of males;
|
||
<emphasis id="D091EAE0FF84652E172C41C6FE0D77BC" bold="true" box="[370,435,1155,1179]" pageId="4" pageNumber="391">23−26</emphasis>
|
||
, terminalia of females (dorsal, lateral, ventral).
|
||
<emphasis id="D091EAE0FF84652E15CC41C1FC0677BB" bold="true" box="[914,952,1156,1180]" pageId="4" pageNumber="391">gcd</emphasis>
|
||
, distal piece of gonocoxite;
|
||
<emphasis id="D091EAE0FF84652E128D41C1FB4777BB" bold="true" box="[1235,1273,1156,1180]" pageId="4" pageNumber="391">gcp</emphasis>
|
||
; proximal piece of gonocoxite;
|
||
<emphasis id="D091EAE0FF84652E176A41EDFEE777E7" bold="true" box="[308,345,1192,1216]" pageId="4" pageNumber="391">ppt</emphasis>
|
||
, paraprocts;
|
||
<emphasis id="D091EAE0FF84652E17BE41EDFE4177E7" bold="true" box="[480,511,1192,1216]" pageId="4" pageNumber="391">gst</emphasis>
|
||
, gonostyle;
|
||
<emphasis id="D091EAE0FF84652E142141EDFD1C77E7" bold="true" box="[639,674,1192,1216]" pageId="4" pageNumber="391">spc</emphasis>
|
||
, spiculum gastrale;
|
||
<emphasis id="D091EAE0FF84652E152D41E2FC2C7798" bold="true" box="[883,914,1191,1215]" pageId="4" pageNumber="391">st9</emphasis>
|
||
, abdominal sternite IX;
|
||
<emphasis id="D091EAE0FF84652E12CE41E2FB0F7798" bold="true" box="[1168,1201,1191,1215]" pageId="4" pageNumber="391">tg9</emphasis>
|
||
, abdominal tergite IX. Scale = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E25A36F2FF84652E16994057FD6E750E" blockId="4.[151,1436,1298,1577]" pageId="4" pageNumber="391">
|
||
<taxonomicName id="25E54D71FF84652E16994057FE63760B" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[199,477,1298,1324]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF84652E16994057FE63760B" box="[199,477,1298,1324]" italics="true" pageId="4" pageNumber="391">Compsodactylus mendax</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="25E54D71FF84652E148C4057FCE2760B" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[722,860,1298,1324]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="D091EAE0FF84652E148C4057FD5B760B" box="[722,741,1298,1324]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E14AC4056FCE2760B" box="[754,860,1299,1324]" italics="true" pageId="4" pageNumber="391">martinezi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="25E54D71FF84652E15D14057FBBC760B" authorityName="Figueroa & Neita-Moreno" authorityYear="2019" box="[911,1026,1298,1324]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="vallejoi">
|
||
<emphasis id="D091EAE0FF84652E15D14057FC1C760B" box="[911,930,1298,1324]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E15F04057FBBC760B" box="[942,1026,1298,1324]" italics="true" pageId="4" pageNumber="391">vallejoi</emphasis>
|
||
</taxonomicName>
|
||
by (characters of 1−
|
||
<taxonomicName id="25E54D71FF84652E12BB4057FAD0760B" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1253,1390,1298,1324]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="D091EAE0FF84652E12BB4057FB46760B" box="[1253,1272,1298,1324]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E135A4056FAD0760B" box="[1284,1390,1299,1324]" italics="true" pageId="4" pageNumber="391">martinezi</emphasis>
|
||
</taxonomicName>
|
||
and 2−
|
||
<taxonomicName id="25E54D71FF84652E16EB4073FE957677" authorityName="Figueroa & Neita-Moreno" authorityYear="2019" box="[181,299,1334,1360]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="vallejoi">
|
||
<emphasis id="D091EAE0FF84652E16EB4073FF767677" box="[181,200,1334,1360]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E16894073FE957677" box="[215,299,1334,1360]" italics="true" pageId="4" pageNumber="391">vallejoi</emphasis>
|
||
</taxonomicName>
|
||
given in brackets): dorsal surface with metallic green reflections (1−as
|
||
<taxonomicName id="25E54D71FF84652E12044073FB6D7677" authorityName="Fuhrmann & Smith" authorityYear="2021" box="[1114,1235,1334,1360]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF84652E12044073FBD37677" box="[1114,1133,1334,1360]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E12224073FB6D7677" box="[1148,1235,1334,1360]" italics="true" pageId="4" pageNumber="391">mendax</emphasis>
|
||
</taxonomicName>
|
||
; 2−without green reflections); antennae with 10 antennomeres (1−nine antennomeres; 2−as
|
||
<taxonomicName id="25E54D71FF84652E15E1401FFA8B7653" authority=": Jhon C. Neita Moreno" box="[959,1333,1370,1396]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF84652E15E1401FFC6C7653" box="[959,978,1370,1396]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E1583401FFB8A7653" box="[989,1076,1370,1396]" italics="true" pageId="4" pageNumber="391">mendax</emphasis>
|
||
: Jhon C. Neita Moreno
|
||
</taxonomicName>
|
||
, personal communication); male protarsomere I greatly enlarged (1−slightly enlarged; 2−as
|
||
<taxonomicName id="25E54D71FF84652E1212403BFB7976BF" box="[1100,1223,1406,1432]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF84652E1212403BFBE176BF" box="[1100,1119,1406,1432]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E122E403BFB7976BF" box="[1136,1223,1406,1432]" italics="true" pageId="4" pageNumber="391">mendax</emphasis>
|
||
</taxonomicName>
|
||
); male metafemur slightly enlarged and approximately 2.1−2.4 times longer than wide (1 and 2−greatly enlarged and approximately 1.5−1.7 times longer than wide;
|
||
<figureCitation id="7ADE2A77FF84652E14524083FD7C76C7" box="[524,706,1478,1505]" captionStart="FIGURES 9−18" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[164,1405,198,1843]" captionTargetId="figure-17@3.[151,1435,181,1864]" captionTargetPageId="3" captionText="FIGURES 9−18. Compsodactylus, posterior leg of males; 9−10, C. argentinus (femur-tibia, apex of tibia); 11−12, C. parvulus (femur-tibia, apex of tibia); 13−14, C. martinezi (femur-tibia, apex of tibia); 15−16, C. mendax new species (femur-tibia, apex of tibia); 17−18, C. vallejoi (femur-tibia, apex of tibia; modified from Figueroa & Neita-Moreno 2019). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026063" httpUri="https://zenodo.org/record/5026063/files/figure.png" pageId="4" pageNumber="391">Figs. 13, 15, 17</figureCitation>
|
||
); male metatarsus somewhat filiform (1−somewhat moniliform; 2−as
|
||
<taxonomicName id="25E54D71FF84652E168D40AFFEF57523" box="[211,331,1514,1540]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF84652E168D40AFFF587523" box="[211,230,1514,1540]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E16AA40AFFEF57523" box="[244,331,1514,1540]" italics="true" pageId="4" pageNumber="391">mendax</emphasis>
|
||
</taxonomicName>
|
||
); male pygidium with hair-like setae (1−with scale-like setae; 2−as
|
||
<taxonomicName id="25E54D71FF84652E121A40AFFB027523" box="[1092,1212,1514,1540]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="D091EAE0FF84652E121A40AFFBE97523" box="[1092,1111,1514,1540]" italics="true" pageId="4" pageNumber="391">C</emphasis>
|
||
.
|
||
<emphasis id="D091EAE0FF84652E123B40AFFB027523" box="[1125,1212,1514,1540]" italics="true" pageId="4" pageNumber="391">mendax</emphasis>
|
||
</taxonomicName>
|
||
); females with long gonocoxite (
|
||
<figureCitation id="7ADE2A77FF84652E177D434BFE13750E" box="[291,429,1550,1577]" captionStart="FIGURES 19−25" captionStartId="4.[151,264,1119,1143]" captionTargetBox="[165,1421,187,1088]" captionTargetId="figure-22@4.[160,1427,181,1096]" captionTargetPageId="4" captionText="FIGURES 19−25. Compsodactylus mendax new species; 19, parameres, distal; 20−21, aedeagus (lateral, laterodistal), 22, genital ring of males; 23−26, terminalia of females (dorsal, lateral, ventral). gcd, distal piece of gonocoxite; gcp; proximal piece of gonocoxite; ppt, paraprocts; gst, gonostyle; spc, spiculum gastrale; st9, abdominal sternite IX; tg9, abdominal tergite IX. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5026065" httpUri="https://zenodo.org/record/5026065/files/figure.png" pageId="4" pageNumber="391">Figs. 23−25</figureCitation>
|
||
;
|
||
<figureCitation id="7ADE2A77FF84652E17E7434BFE7C750F" box="[441,450,1550,1576]" captionStart="FIGURES 1−8" captionStartId="2.[151,264,1979,2003]" captionTargetBox="[151,1436,270,1953]" captionTargetId="figure-74@2.[151,1436,270,1953]" captionTargetPageId="2" captionText="FIGURES 1−8. Compsodactylus mendax new species; 1−5, male holotype (dorsal, dorsolateral, lateral, dorsofrontal, label); 6−8, female allotype (dorsal, dorsolateral, lateral). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5026059" httpUri="https://zenodo.org/record/5026059/files/figure.png" pageId="4" pageNumber="391">1</figureCitation>
|
||
−wide; 2−comma-like).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |