173 lines
27 KiB
XML
173 lines
27 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3682289" ID-GBIF-Dataset="10c257cd-4869-4667-86fb-9ff9b671ada1" ID-GBIF-Taxon="162247148" ID-Zenodo-Dep="3682289" approvedBy="poa" checkinTime="1582296023719" checkinUser="tatiana" docAuthor="Gustavo R. S. Ruiz & Antonio D. Brescovit" docDate="2008" docId="6A0A87C9A661FF8E48EBBC9AB9B8F911" docLanguage="en" docName="RevBrasZool.25.1.139-147.pdf.imf" docOrigin="Revista Brasileira de Zoologia 25 (1)" docStyle="DocumentStyle{}" docTitle="Helvetia Peckham & Peckham 1894" docType="treatment" docVersion="8" lastPageNumber="140" masterDocId="9633FFB1A660FF8F4827BD28BB36FFEB" masterDocTitle="Revision of Helvetia (Araneae: Salticidae: Heliophaninae)" masterLastPageNumber="147" masterPageNumber="139" pageId="1" pageNumber="140" updateTime="1643574341443" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of Helvetia (Araneae: Salticidae: Heliophaninae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gustavo R. S. Ruiz</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Antonio D. Brescovit</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Revista Brasileira de Zoologia</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2008</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2008-03-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>25</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>139</mods:start>
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<mods:end>147</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/S0101-81752008000100018</mods:identifier>
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<mods:identifier type="GBIF-Dataset">10c257cd-4869-4667-86fb-9ff9b671ada1</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3678502</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3682289" ID-GBIF-Taxon="162247148" ID-Zenodo-Dep="3682289" LSID="urn:lsid:plazi:treatment:6A0A87C9A661FF8E48EBBC9AB9B8F911" httpUri="http://treatment.plazi.org/id/6A0A87C9A661FF8E48EBBC9AB9B8F911" lastPageNumber="140" pageId="1" pageNumber="140">
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<subSubSection box="[204,648,434,462]" pageId="1" pageNumber="140" type="nomenclature">
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<paragraph blockId="1.[204,648,434,462]" box="[204,648,434,462]" pageId="1" pageNumber="140">
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<heading bold="true" box="[204,648,434,462]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="140" reason="3">
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<taxonomicName authority="Peckham & Peckham, 1894" authorityName="Peckham & Peckham" authorityYear="1894" box="[204,648,434,462]" class="Arachnida" family="Salticidae" genus="Helvetia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[204,292,434,462]" italics="true" pageId="1" pageNumber="140">Helvetia</emphasis>
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<bibRefCitation author="Peckham & Peckham" box="[303,648,434,462]" firstAuthor="Peckham" journalOrPublisher="Occasional Papers of the Natural History Society of Wisconsin" pageId="1" pageNumber="140" pagination="85 - 156" part="2" refId="ref6708" refString="PECKHAM, G. W. & E. G. PECKHAM. 1894. Spiders of the Marptusa group. Occasional Papers of the Natural History Society of Wisconsin 2: 85 - 156." title="Spiders of the Marptusa group" type="journal article" year="1894">Peckham & Peckham, 1894</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="140" type="type_taxon">
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<paragraph blockId="1.[113,738,474,614]" pageId="1" pageNumber="140">
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<treatmentCitationGroup pageId="1" pageNumber="140">
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<taxonomicName authority="Peckham & Peckham, 1894: 119" authorityName="Peckham & Peckham" authorityPageNumber="119" authorityYear="1894" box="[113,509,474,494]" class="Arachnida" family="Salticidae" genus="Helvetia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis box="[113,189,475,494]" italics="true" pageId="1" pageNumber="140">Helvetia</emphasis>
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<treatmentCitation author="PECKHAM, G. W. & E. G. PECKHAM" box="[194,509,474,494]" page="119" pageId="1" pageNumber="140" year="1894">
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<bibRefCitation author="Peckham & Peckham" box="[194,509,474,494]" firstAuthor="Peckham" journalOrPublisher="Occasional Papers of the Natural History Society of Wisconsin" pageId="1" pageNumber="140" pagination="85 - 156" part="2" refId="ref6708" refString="PECKHAM, G. W. & E. G. PECKHAM. 1894. Spiders of the Marptusa group. Occasional Papers of the Natural History Society of Wisconsin 2: 85 - 156." title="Spiders of the Marptusa group" type="journal article" year="1894">Peckham & Peckham, 1894: 119</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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(
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<typeStatus box="[520,564,475,494]" pageId="1" pageNumber="140">type</typeStatus>
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species by mono- typy:
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<taxonomicName authority="Peckham & Peckham, 1894" authorityName="Peckham & Peckham" authorityYear="1894" box="[211,726,504,524]" class="Arachnida" family="Salticidae" genus="Helvetia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="santarema">
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<emphasis box="[211,411,505,524]" italics="true" pageId="1" pageNumber="140">Helvetia santarema</emphasis>
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<treatmentCitation author="Peckham & Peckham" box="[421,726,504,524]" pageId="1" pageNumber="140" year="1984">
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<bibRefCitation author="Peckham & Peckham" box="[421,726,504,524]" firstAuthor="Peckham" journalOrPublisher="Occasional Papers of the Natural History Society of Wisconsin" pageId="1" pageNumber="140" pagination="85 - 156" part="2" refId="ref6708" refString="PECKHAM, G. W. & E. G. PECKHAM. 1894. Spiders of the Marptusa group. Occasional Papers of the Natural History Society of Wisconsin 2: 85 - 156." title="Spiders of the Marptusa group" type="journal article" year="1894">Peckham & Peckham, 1894</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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);
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<treatmentCitation author="Platnick" box="[145,293,534,554]" pageId="1" pageNumber="140" year="2007">
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<bibRefCitation author="Platnick" box="[145,293,534,554]" firstAuthor="Platnick" journalOrPublisher="American Museum of Natural History, Electronic Database" pageId="1" pageNumber="140" publicationUrl="http://research.amnh.org/entomology/spiders/catalog/index.html" refId="ref6744" refString="PLATNICK, N. I. 2007. The world spider catalog. Version 7.5. American Museum of Natural History, Electronic Database available at http: // research. amnh. org / entomology / spiders / catalog / index. html" title="The world spider catalog. Version 7.5" type="url" year="2007">Platnick, 2007</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="140" type="reference_group">
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<paragraph blockId="1.[113,738,474,614]" pageId="1" pageNumber="140">
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<treatmentCitationGroup pageId="1" pageNumber="140">
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<taxonomicName authority="Simon, 1901 a: 607" authorityName="Simon" authorityPageNumber="607" authorityYear="1901" box="[113,375,564,584]" class="Arachnida" family="Salticidae" genus="Deloripa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis box="[113,191,564,583]" italics="true" pageId="1" pageNumber="140">Deloripa</emphasis>
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<treatmentCitation author="SIMON, E." box="[196,375,564,584]" page="607" pageId="1" pageNumber="140" year="1901">
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<bibRefCitation author="Simon" box="[196,375,564,584]" firstAuthor="Simon" journalOrPublisher="Paris" pageId="1" pageNumber="140" pagination="381 - 668" part="2" refId="ref6817" refString="SIMON, E. 1901 a. Histoire naturelle des araignees. Paris, vol. 2, p. 381 - 668." title="Histoire naturelle des araignees" type="journal article" year="1901" yearSuffix="a">Simon, 1901a: 607</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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(
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<typeStatus box="[387,430,565,584]" pageId="1" pageNumber="140">type</typeStatus>
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species by original designation:
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<taxonomicName authority="Simon, 1901" authorityName="Simon" authorityYear="1901" box="[145,447,594,614]" class="Arachnida" family="Salticidae" genus="Deloripa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="semialba">
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<emphasis box="[145,312,594,613]" italics="true" pageId="1" pageNumber="140">Deloripa semialba</emphasis>
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<treatmentCitation author="Simon" box="[319,447,594,614]" pageId="1" pageNumber="140" year="1901">
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<bibRefCitation author="Simon" box="[319,447,594,614]" firstAuthor="Simon" journalOrPublisher="Paris" pageId="1" pageNumber="140" pagination="381 - 668" part="2" refId="ref6817" refString="SIMON, E. 1901 a. Histoire naturelle des araignees. Paris, vol. 2, p. 381 - 668." title="Histoire naturelle des araignees" type="journal article" year="1901">Simon, 1901</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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);
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<treatmentCitation author="Platnick" box="[464,608,594,614]" pageId="1" pageNumber="140" year="2007">
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<bibRefCitation author="Platnick" box="[464,608,594,614]" firstAuthor="Platnick" journalOrPublisher="American Museum of Natural History, Electronic Database" pageId="1" pageNumber="140" publicationUrl="http://research.amnh.org/entomology/spiders/catalog/index.html" refId="ref6744" refString="PLATNICK, N. I. 2007. The world spider catalog. Version 7.5. American Museum of Natural History, Electronic Database available at http: // research. amnh. org / entomology / spiders / catalog / index. html" title="The world spider catalog. Version 7.5" type="url" year="2007">Platnick, 2007</bibRefCitation>
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</treatmentCitation>
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.
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<emphasis bold="true" box="[618,715,594,613]" pageId="1" pageNumber="140">Syn. nov.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="140" type="diagnosis">
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<paragraph blockId="1.[113,738,631,1786]" pageId="1" pageNumber="140">
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Diagnosis. The neotropical genus
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<taxonomicName authorityName="Peckham & Peckham" authorityYear="1894" box="[516,593,632,651]" class="Arachnida" family="Salticidae" genus="Helvetia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus">
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<emphasis box="[516,593,632,651]" italics="true" pageId="1" pageNumber="140">Helvetia</emphasis>
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</taxonomicName>
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can be distinguished from all the remaining genera of the subfamily
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<taxonomicName authorityName="Petrunkevitch" authorityYear="1928" box="[113,261,691,711]" class="Arachnida" family="Salticidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="140" phylum="Arthropoda" rank="subFamily" subFamily="Heliophaninae">Heliophaninae</taxonomicName>
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by having a trapezoid carapace, which is much wider at the posterior region than it is at the region of the anterior eye row (
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<figureCitation box="[292,348,751,771]" captionStart="Figures 4-10" captionStartId="4.[189,256,1381,1401]" captionTargetBox="[201,1451,197,1372]" captionTargetId="figure@4.[197,1446,189,1376]" captionTargetPageId="4" captionText="Figures 4-10. Epigynum of (4-5) Helvetia cancrimana: (4) ventral view̡ (5) dorsal view; (6-7) Helvetia labiata: (6) ventral view̡ (7) dorsal view; (8-10) Helvetia semialba: (8) female holotype̡ dorsal view̡ (9) epigynum̡ vental view̡ (10) dorsal view. Bars = 0.25 mm̡ except for fig. 8 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3678506" httpUri="https://zenodo.org/record/3678506/files/figure.png" pageId="1" pageNumber="140">Fig. 8</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="140" type="description">
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<paragraph blockId="1.[113,738,631,1786]" pageId="1" pageNumber="140">
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Description. Small spiders (about
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<quantity box="[502,579,781,801]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="140" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3-6 mm</quantity>
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) with low, elongated carapace and long abdomen (
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<figureCitation box="[484,541,810,830]" captionStart="Figures 1-3" captionStartId="2.[189,256,1210,1230]" captionTargetBox="[189,1474,195,1202]" captionTargetId="figure@2.[265,1399,186,715]" captionTargetPageId="2" captionText="Figures 1-3. Helvetia cancrimana: (1) male̡ laterodorsal view̡ arrows show the stridulatory apparatus on femur I and carapace; (2) stridulatory apparatus̡ rugose portion of lateral carapace; (3) modified femoral setae of the stridulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.3678504" httpUri="https://zenodo.org/record/3678504/files/figure.png" pageId="1" pageNumber="140">Fig. 1</figureCitation>
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). Abdominal color patterns include pairs of light patches on a dark background, which can be fused forming transverse stripes (
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<figureCitation box="[610,668,870,890]" captionStart="Figures 1-3" captionStartId="2.[189,256,1210,1230]" captionTargetBox="[189,1474,195,1202]" captionTargetId="figure@2.[265,1399,186,715]" captionTargetPageId="2" captionText="Figures 1-3. Helvetia cancrimana: (1) male̡ laterodorsal view̡ arrows show the stridulatory apparatus on femur I and carapace; (2) stridulatory apparatus̡ rugose portion of lateral carapace; (3) modified femoral setae of the stridulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.3678504" httpUri="https://zenodo.org/record/3678504/files/figure.png" pageId="1" pageNumber="140">Fig. 1</figureCitation>
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), dark abdomen with a large patch of light scales covering the entire abdomen and longitudinal stripes of dark scales on a light background (
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<figureCitation box="[199,254,960,980]" captionStart="Figures 4-10" captionStartId="4.[189,256,1381,1401]" captionTargetBox="[201,1451,197,1372]" captionTargetId="figure@4.[197,1446,189,1376]" captionTargetPageId="4" captionText="Figures 4-10. Epigynum of (4-5) Helvetia cancrimana: (4) ventral view̡ (5) dorsal view; (6-7) Helvetia labiata: (6) ventral view̡ (7) dorsal view; (8-10) Helvetia semialba: (8) female holotype̡ dorsal view̡ (9) epigynum̡ vental view̡ (10) dorsal view. Bars = 0.25 mm̡ except for fig. 8 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3678506" httpUri="https://zenodo.org/record/3678506/files/figure.png" pageId="1" pageNumber="140">Fig. 8</figureCitation>
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). Chelicerae about one third of carapace length, almost vertical (
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<figureCitation box="[275,329,989,1009]" captionStart="Figures 1-3" captionStartId="2.[189,256,1210,1230]" captionTargetBox="[189,1474,195,1202]" captionTargetId="figure@2.[265,1399,186,715]" captionTargetPageId="2" captionText="Figures 1-3. Helvetia cancrimana: (1) male̡ laterodorsal view̡ arrows show the stridulatory apparatus on femur I and carapace; (2) stridulatory apparatus̡ rugose portion of lateral carapace; (3) modified femoral setae of the stridulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.3678504" httpUri="https://zenodo.org/record/3678504/files/figure.png" pageId="1" pageNumber="140">Fig. 1</figureCitation>
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), with two teeth on promargin and one on retromargin. Legs Iusually strongest and darkest, strongly developed in some species (
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<figureCitation box="[384,436,1049,1069]" captionStart="Figures 1-3" captionStartId="2.[189,256,1210,1230]" captionTargetBox="[189,1474,195,1202]" captionTargetId="figure@2.[265,1399,186,715]" captionTargetPageId="2" captionText="Figures 1-3. Helvetia cancrimana: (1) male̡ laterodorsal view̡ arrows show the stridulatory apparatus on femur I and carapace; (2) stridulatory apparatus̡ rugose portion of lateral carapace; (3) modified femoral setae of the stridulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.3678504" httpUri="https://zenodo.org/record/3678504/files/figure.png" pageId="1" pageNumber="140">Fig. 1</figureCitation>
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). Palps have the heliophanine bump and can have a short embolus, originating distally on the prolateral side of the tegulum and a short RTA (
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<figureCitation captionStart="Figures 15-20" captionStartId="6.[189,256,1173,1193]" captionTargetBox="[352,1306,201,1167]" captionTargetId="figure@6.[344,1309,189,1172]" captionTargetPageId="6" captionText="Figures 15-20. Left palp of (15-16) Helvetia galianoae: (15) ventral view̡ (16) retrolateral view; (17-18) Helvetia humillima: (17) ventral view̡ (18) retrolateral view; (19-20) Helvetia stridulans: (19) vental view̡ (20) retrolareal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678513" httpUri="https://zenodo.org/record/3678513/files/figure.png" pageId="1" pageNumber="140">Figs 15 and 16</figureCitation>
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), or can exhibit extensive modifications, such as a very long embolus associated with a counterclockwise rotation of the tegulum in the left palp (
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<figureCitation box="[368,438,1199,1219]" captionStart="Figures 15-20" captionStartId="6.[189,256,1173,1193]" captionTargetBox="[352,1306,201,1167]" captionTargetId="figure@6.[344,1309,189,1172]" captionTargetPageId="6" captionText="Figures 15-20. Left palp of (15-16) Helvetia galianoae: (15) ventral view̡ (16) retrolateral view; (17-18) Helvetia humillima: (17) ventral view̡ (18) retrolateral view; (19-20) Helvetia stridulans: (19) vental view̡ (20) retrolareal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678513" httpUri="https://zenodo.org/record/3678513/files/figure.png" pageId="1" pageNumber="140">Fig. 19</figureCitation>
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), grooves on the cymbium to hold the tip of the curling embolus (
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<figureCitation box="[480,549,1229,1249]" captionStart="Figures 15-20" captionStartId="6.[189,256,1173,1193]" captionTargetBox="[352,1306,201,1167]" captionTargetId="figure@6.[344,1309,189,1172]" captionTargetPageId="6" captionText="Figures 15-20. Left palp of (15-16) Helvetia galianoae: (15) ventral view̡ (16) retrolateral view; (17-18) Helvetia humillima: (17) ventral view̡ (18) retrolateral view; (19-20) Helvetia stridulans: (19) vental view̡ (20) retrolareal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678513" httpUri="https://zenodo.org/record/3678513/files/figure.png" pageId="1" pageNumber="140">Fig. 18</figureCitation>
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), bifid RTA, with a developed ventral branch (
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<figureCitation box="[391,461,1258,1278]" captionStart="Figures 15-20" captionStartId="6.[189,256,1173,1193]" captionTargetBox="[352,1306,201,1167]" captionTargetId="figure@6.[344,1309,189,1172]" captionTargetPageId="6" captionText="Figures 15-20. Left palp of (15-16) Helvetia galianoae: (15) ventral view̡ (16) retrolateral view; (17-18) Helvetia humillima: (17) ventral view̡ (18) retrolateral view; (19-20) Helvetia stridulans: (19) vental view̡ (20) retrolareal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678513" httpUri="https://zenodo.org/record/3678513/files/figure.png" pageId="1" pageNumber="140">Fig. 20</figureCitation>
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) or with a very developed dorsal branch (see
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<bibRefCitation author="GALIANO" box="[309,446,1288,1308]" firstAuthor="GALIANO" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="1" pageNumber="140" pagination="51 - 56" part="35" refId="ref6535" refString="GALIANO, M. E. 1976. Dos nuevas especies del genero Helvetia Peckham, 1894 (Araneae, Salticidae). Revista de la Sociedad Entomologica Argentina 35: 51 - 56." title="Dos nuevas especies del genero Helvetia Peckham, 1894 (Araneae, Salticidae)" type="journal article" year="1976">GALIANO 1976</bibRefCitation>
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, fig. 10). Epigyna vary from very short copulatory ducts (
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<figureCitation box="[404,526,1318,1338]" captionStart="Figures 4-10" captionStartId="4.[189,256,1381,1401]" captionTargetBox="[201,1451,197,1372]" captionTargetId="figure@4.[197,1446,189,1376]" captionTargetPageId="4" captionText="Figures 4-10. Epigynum of (4-5) Helvetia cancrimana: (4) ventral view̡ (5) dorsal view; (6-7) Helvetia labiata: (6) ventral view̡ (7) dorsal view; (8-10) Helvetia semialba: (8) female holotype̡ dorsal view̡ (9) epigynum̡ vental view̡ (10) dorsal view. Bars = 0.25 mm̡ except for fig. 8 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3678506" httpUri="https://zenodo.org/record/3678506/files/figure.png" pageId="1" pageNumber="140">Figs 4 and 5</figureCitation>
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) to coiled, very long copulatory ducts (
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<figureCitation box="[295,399,1348,1368]" captionStart="Figures 11-14" captionStartId="5.[113,180,1154,1174]" captionTargetBox="[248,1259,194,1141]" captionTargetId="figure@5.[242,1263,181,1150]" captionTargetPageId="5" captionText="Figures 11-14. Epigynum of (11-12) Helvetia santarema: (11) ventral view̡ (12) dorsal view; (13-14) Helvetia albovittata: (13) ventral view̡ (14) dorsal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678508" httpUri="https://zenodo.org/record/3678508/files/figure.png" pageId="1" pageNumber="140">Figs 11-14</figureCitation>
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), on whose medial part there is a pair of glandular swollen areas; copulatory openings are usually well separated from each other (
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<figureCitation box="[486,599,1408,1428]" captionStart="Figures 4-10" captionStartId="4.[189,256,1381,1401]" captionTargetBox="[201,1451,197,1372]" captionTargetId="figure@4.[197,1446,189,1376]" captionTargetPageId="4" captionText="Figures 4-10. Epigynum of (4-5) Helvetia cancrimana: (4) ventral view̡ (5) dorsal view; (6-7) Helvetia labiata: (6) ventral view̡ (7) dorsal view; (8-10) Helvetia semialba: (8) female holotype̡ dorsal view̡ (9) epigynum̡ vental view̡ (10) dorsal view. Bars = 0.25 mm̡ except for fig. 8 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3678506" httpUri="https://zenodo.org/record/3678506/files/figure.png" pageId="1" pageNumber="140">Figs 4, 6, 9</figureCitation>
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, and 13), but can be found close together, forming a median small atrium (
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<figureCitation box="[120,189,1468,1488]" captionStart="Figures 11-14" captionStartId="5.[113,180,1154,1174]" captionTargetBox="[248,1259,194,1141]" captionTargetId="figure@5.[242,1263,181,1150]" captionTargetPageId="5" captionText="Figures 11-14. Epigynum of (11-12) Helvetia santarema: (11) ventral view̡ (12) dorsal view; (13-14) Helvetia albovittata: (13) ventral view̡ (14) dorsal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678508" httpUri="https://zenodo.org/record/3678508/files/figure.png" pageId="1" pageNumber="140">Fig. 11</figureCitation>
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) or on the internal wall of a very large, deep atrium (
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<figureCitation box="[119,276,1497,1517]" captionStart="Figures 21-24" captionStartId="7.[114,181,1116,1136]" captionTargetBox="[254,1255,198,1101]" captionTargetId="figure@7.[249,1256,189,1104]" captionTargetPageId="7" captionText="Figures 21-24. Epigynum of (21-22) Helvetia rinaldiae: (21) ventral view̡ (22) dorsal view; (23-24) Helvetia roeweri: (23) ventral view̡ (24) dorsal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678517" httpUri="https://zenodo.org/record/3678517/files/figure.png" pageId="1" pageNumber="140">Figs 21 and 22</figureCitation>
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); the epigynal pockets can be fused (
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<figureCitation box="[665,725,1497,1517]" captionStart="Figures 4-10" captionStartId="4.[189,256,1381,1401]" captionTargetBox="[201,1451,197,1372]" captionTargetId="figure@4.[197,1446,189,1376]" captionTargetPageId="4" captionText="Figures 4-10. Epigynum of (4-5) Helvetia cancrimana: (4) ventral view̡ (5) dorsal view; (6-7) Helvetia labiata: (6) ventral view̡ (7) dorsal view; (8-10) Helvetia semialba: (8) female holotype̡ dorsal view̡ (9) epigynum̡ vental view̡ (10) dorsal view. Bars = 0.25 mm̡ except for fig. 8 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3678506" httpUri="https://zenodo.org/record/3678506/files/figure.png" pageId="1" pageNumber="140">Fig. 4</figureCitation>
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), medially placed (
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<figureCitation box="[285,421,1527,1547]" captionStart="Figures 4-10" captionStartId="4.[189,256,1381,1401]" captionTargetBox="[201,1451,197,1372]" captionTargetId="figure@4.[197,1446,189,1376]" captionTargetPageId="4" captionText="Figures 4-10. Epigynum of (4-5) Helvetia cancrimana: (4) ventral view̡ (5) dorsal view; (6-7) Helvetia labiata: (6) ventral view̡ (7) dorsal view; (8-10) Helvetia semialba: (8) female holotype̡ dorsal view̡ (9) epigynum̡ vental view̡ (10) dorsal view. Bars = 0.25 mm̡ except for fig. 8 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3678506" httpUri="https://zenodo.org/record/3678506/files/figure.png" pageId="1" pageNumber="140">Figs 7 and 10</figureCitation>
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) or separated into a pair of lateral pockets (
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<figureCitation box="[243,390,1557,1577]" captionStart="Figures 11-14" captionStartId="5.[113,180,1154,1174]" captionTargetBox="[248,1259,194,1141]" captionTargetId="figure@5.[242,1263,181,1150]" captionTargetPageId="5" captionText="Figures 11-14. Epigynum of (11-12) Helvetia santarema: (11) ventral view̡ (12) dorsal view; (13-14) Helvetia albovittata: (13) ventral view̡ (14) dorsal view. Bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3678508" httpUri="https://zenodo.org/record/3678508/files/figure.png" pageId="1" pageNumber="140">Figs 12 and 14</figureCitation>
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), which can be connected in some species. The stridulatory apparatus consists in this group of a series of three modified setae dorsally on the distal portion of the femur I (
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<figureCitation box="[238,359,1647,1667]" captionStart="Figures 1-3" captionStartId="2.[189,256,1210,1230]" captionTargetBox="[189,1474,195,1202]" captionTargetId="figure@2.[265,1399,186,715]" captionTargetPageId="2" captionText="Figures 1-3. Helvetia cancrimana: (1) male̡ laterodorsal view̡ arrows show the stridulatory apparatus on femur I and carapace; (2) stridulatory apparatus̡ rugose portion of lateral carapace; (3) modified femoral setae of the stridulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.3678504" httpUri="https://zenodo.org/record/3678504/files/figure.png" pageId="1" pageNumber="140">Figs 1 and 3</figureCitation>
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) and a granulose area laterally on the anterior portion of the carapace (
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<figureCitation box="[442,564,1677,1697]" captionStart="Figures 1-3" captionStartId="2.[189,256,1210,1230]" captionTargetBox="[189,1474,195,1202]" captionTargetId="figure@2.[265,1399,186,715]" captionTargetPageId="2" captionText="Figures 1-3. Helvetia cancrimana: (1) male̡ laterodorsal view̡ arrows show the stridulatory apparatus on femur I and carapace; (2) stridulatory apparatus̡ rugose portion of lateral carapace; (3) modified femoral setae of the stridulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.3678504" httpUri="https://zenodo.org/record/3678504/files/figure.png" pageId="1" pageNumber="140">Figs 1 and 2</figureCitation>
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), extending from the anterior lateral eyes to the region behind the posterior lateral eyes.
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</paragraph>
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</subSubSection>
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<subSubSection box="[174,654,1766,1786]" pageId="1" pageNumber="140" type="distribution">
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<paragraph blockId="1.[113,738,631,1786]" box="[174,654,1766,1786]" pageId="1" pageNumber="140">Distribution. Known only from South America.</paragraph>
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</subSubSection>
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</treatment>
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</document> |