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<document id="E28261C320FE33CC362ACED9F1CA7A5A" ID-DOI="10.13133/2284-4880/525" ID-ISSN="2284-4880" ID-Zenodo-Dep="12763456" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721241537173" checkinUser="felipe" docAuthor="Kasparek, Max" docDate="2021" docId="681A87E0932BA50AFF12FB009D55F9DA" docLanguage="en" docName="FragEntomol.53.2.333-346.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/525" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Exanthidium guichardi Pasteels 1980" docType="treatment" docVersion="2" lastPageNumber="341" masterDocId="9423FF98932CA502FF9AFFB49E56FFF6" masterDocTitle="The bee genus Pseudoanthidium: revision of the subgenus Exanthidium with the description of a new species (Apoidea: Megachilidae)" masterLastPageNumber="346" masterPageNumber="333" pageNumber="340" updateTime="1721243763790" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="B9F31F7FDEC65A0CC917C666DB4454AA">The bee genus Pseudoanthidium: revision of the subgenus Exanthidium with the description of a new species (Apoidea: Megachilidae)</mods:title>
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<mods:namePart id="E6B5EE617CDE706E98A19E855F5EAC91">Kasparek, Max</mods:namePart>
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<mods:title id="56DE544A94760E4041F48918D41141BC">Fragmenta entomologica</mods:title>
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<mods:date id="D55852FD805C1F5D96495D3BB48A3450">2021</mods:date>
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<mods:number id="90D01E121F57E7F55EEB31211A210CB4">2021-11-30</mods:number>
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<mods:number id="36EDA9C3D1298F376CAB0C35E1204C8E">53</mods:number>
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<mods:number id="F162D6C920C06A656F843A8D58BEBD97">2</mods:number>
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<mods:url id="6CDD10B927AFEE7EB224CD132BA417EE">http://dx.doi.org/10.13133/2284-4880/525</mods:url>
</mods:location>
<mods:classification id="480BF63B48D77AAF7631D1DE16A19000">journal article</mods:classification>
<mods:identifier id="90EB6154A09B4A9A5E5BF537E4742472" type="DOI">10.13133/2284-4880/525</mods:identifier>
<mods:identifier id="635B85B89B9449774C5864ED6B6BAA05" type="ISSN">2284-4880</mods:identifier>
<mods:identifier id="BE5D85882D163DC13B633CD48EB8EC9C" type="Zenodo-Dep">12763456</mods:identifier>
<mods:identifier id="753EF77891FCAF53D9CD3EBE149D75FE" type="ZooBank">230E3A6B-A580-400B-B8E3-3811E030B906</mods:identifier>
</mods:mods>
<treatment id="681A87E0932BA50AFF12FB009D55F9DA" LSID="urn:lsid:plazi:treatment:681A87E0932BA50AFF12FB009D55F9DA" httpUri="http://treatment.plazi.org/id/681A87E0932BA50AFF12FB009D55F9DA" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340">
<subSubSection id="A8A9657D932BA505FF12FB009C4BFB3A" box="[136,541,1204,1228]" pageId="7" pageNumber="340" type="nomenclature">
<paragraph id="E00C36F6932BA505FF12FB009C4BFB3A" blockId="7.[136,541,1204,1228]" box="[136,541,1204,1228]" pageId="7" pageNumber="340">
<heading id="BB44819A932BA505FF12FB009C4BFB3A" box="[136,541,1204,1228]" fontSize="10" level="2" pageId="7" pageNumber="340" reason="5">
<taxonomicName id="27B34D75932BA505FF12FB009C4BFB3A" ID-CoL="8SP7R" authority="Pasteels, 1980" authorityName="Pasteels" authorityYear="1980" box="[136,541,1204,1228]" class="Insecta" family="Megachilidae" genus="Exanthidium" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="guichardi">
<emphasis id="D2C7EAE4932BA505FF12FB009F28FB3A" box="[136,382,1204,1228]" italics="true" pageId="7" pageNumber="340">Exanthidium guichardi</emphasis>
<bibRefCitation id="84224B07932BA505FE1EFB009C4BFB3A" author="Pasteels J. J." box="[388,541,1204,1228]" pageId="7" pageNumber="340" pagination="91 - 98" refId="ref8834" refString="Pasteels J. J. 1980. Notes sur les Megachilidae (Hymenoptera, Apoidea) d'Arabie et des regions desertiques limitrophes. 2. Especes nouvelles d'Anthidiinae. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 116: 91 - 98." type="journal article" year="1980">Pasteels, 1980</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A8A9657D932BA50AFF34FB609D55F9DA" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340" type="materials_examined">
<paragraph id="E00C36F6932BA505FF34FB609C31FA9A" blockId="7.[135,760,1236,1932]" pageId="7" pageNumber="340">
<materialsCitation id="50DB3CAB932BA505FF34FB609C31FA9A" collectingDate="1976-03-05" collectorName="Wadi Ouryat, Ag. &amp; K. M. Guichard" country="Oman" county="Male" latitude="22.916666" location="Ag. St." longLatPrecision="1258" longitude="57.15" municipality="Wadi Ouryat" pageId="7" pageNumber="340" specimenCount="1" stateProvince="Ad Dakhiliyah" typeStatus="holotype">
<emphasis id="D2C7EAE4932BA505FF34FB609F44FB1A" box="[174,274,1236,1260]" italics="true" pageId="7" pageNumber="340">
<typeStatus id="3F088854932BA505FF34FB609F58FB1A" box="[174,270,1236,1260]" pageId="7" pageNumber="340" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="096D4E7A932BA505FE83FB609F06FB1A" box="[281,336,1236,1260]" pageId="7" pageNumber="340">Male</collectingCounty>
,
<collectingCountry id="98A47666932BA505FEC6FB609FC9FB1A" box="[348,415,1236,1260]" name="Oman" pageId="7" pageNumber="340">Oman</collectingCountry>
:
<collectorName id="4D465320932BA505FE30FB609C34FB1A" box="[426,610,1236,1260]" pageId="7" pageNumber="340">
<collectingMunicipality id="0068AC8C932BA505FE30FB609C67FB1A" box="[426,561,1236,1260]" pageId="7" pageNumber="340">Wadi Ouryat</collectingMunicipality>
,
<location id="E56C602D932BA505FDA6FB609C34FB1A" LSID="urn:lsid:plazi:treatment:681A87E0932BA50AFF12FB009D55F9DA:E56C602D932BA505FDA6FB609C34FB1A" box="[572,610,1236,1260]" country="Oman" county="Male" latitude="22.916666" longLatPrecision="1258" longitude="57.15" municipality="Wadi Ouryat" name="Ag. St." pageId="7" pageNumber="340" stateProvince="Ad Dakhiliyah">Ag.</location>
</collectorName>
St.,
<date id="940D1036932BA505FD0EFB609CA3FB1A" box="[660,757,1236,1260]" pageId="7" pageNumber="340" value="1976-03-05">
<collectingDate id="8449E9DE932BA505FD0EFB609CA3FB1A" box="[660,757,1236,1260]" pageId="7" pageNumber="340" value="1976-03-05">5.iii.1976</collectingDate>
</date>
(
<collectingRegion id="2277F814932BA505FF15FB409FCBFAFB" box="[143,413,1268,1293]" country="Oman" name="Ad Dakhiliyah" pageId="7" pageNumber="340">ad-Dachiliyya governorate</collectingRegion>
, approx.
<geoCoordinate id="85875031932BA505FE69FB409C1BFAFA" box="[499,589,1268,1292]" degrees="22" direction="north" minutes="55" orientation="latitude" pageId="7" pageNumber="340" precision="925" value="22.916666">22°55’N</geoCoordinate>
,
<geoCoordinate id="85875031932BA505FDCEFB409CFAFAFA" box="[596,684,1268,1292]" degrees="57" direction="east" minutes="09" orientation="longitude" pageId="7" pageNumber="340" precision="925" value="57.15">57°09’E</geoCoordinate>
),
<collectorName id="4D465320932BA505FD20FB419EB0FADA" pageId="7" pageNumber="340">K.M. Guichard</collectorName>
leg.; https://data.nhm.ac.uk/dataset/collection-specimens/resource/05ff2255-c38a-40c9-b657-4ccb55ab2feb/record/9096058; photograph (
<figureCitation id="78882A73932BA505FE3BFAE09FA6FA9A" box="[417,496,1364,1388]" captionStart="Fig" captionStartId="6.[151,182,1885,1904]" captionTargetBox="[151,1451,171,1862]" captionTargetId="figure-9@6.[151,1451,171,1862]" captionTargetPageId="6" captionText="Fig. 9 – Pseudoanthidium guichardi Pasteels, 1980. Male. A, Holotype from Oman (NHMUK); B-H, Male from United Arab Emirates (DEI); A-B, Habitus ventral; C-D, Head and mesosoma in dorsal view; E, Head in frontal view; F, Apical terga; G, Habitus lateral; H, Apex of metasoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.12763484" httpUri="https://zenodo.org/record/12763484/files/figure.png" pageId="7" pageNumber="340">Fig. 9A</figureCitation>
) examined.
</materialsCitation>
</paragraph>
<paragraph id="E00C36F6932BA505FF34FAC09FE0F9DA" blockId="7.[135,760,1236,1932]" pageId="7" pageNumber="340">
<emphasis id="D2C7EAE4932BA505FF34FAC09F9BFA7A" box="[174,461,1396,1420]" italics="true" pageId="7" pageNumber="340">Other material examined.</emphasis>
<materialsCitation id="50DB3CAB932BA505FE42FAC09EF7F9FA" collectingDate="2005-04-08" collectionCode="DEI" collectorName="Deposited &amp; A. van Harten" country="United Arab Emirates" latitude="24.083334" location="UNITED ARAB EMIRATES" longLatPrecision="1252" longitude="55.666668" pageId="7" pageNumber="340" specimenCode="DEI130" specimenCount="1" specimenCount-male="1">
<emphasis id="D2C7EAE4932BA505FE42FAC09E9AFA5A" bold="true" pageId="7" pageNumber="340">
<collectingCountry id="98A47666932BA505FE42FAC09E9AFA5A" name="United Arab Emirates" pageId="7" pageNumber="340">UNITED ARAB EMIRATES</collectingCountry>
</emphasis>
:
<specimenCount id="F6B5FD7F932BA505FF47FA209F5CFA5A" box="[221,266,1428,1452]" count="1" pageId="7" pageNumber="340" type="male">1 ♂</specimenCount>
, al-Ain al-Fayda (
<geoCoordinate id="85875031932BA505FE4FFA209C62FA5A" box="[469,564,1428,1452]" degrees="24" direction="north" minutes="05" orientation="latitude" pageId="7" pageNumber="340" precision="925" value="24.083334">24°05’N</geoCoordinate>
,
<geoCoordinate id="85875031932BA505FDD8FA209CC9FA5A" box="[578,671,1428,1452]" degrees="55" direction="east" minutes="40" orientation="longitude" pageId="7" pageNumber="340" precision="925" value="55.666668">55°40’E</geoCoordinate>
),
<date id="940D1036932BA505FD2FFA209EEBFA3A" pageId="7" pageNumber="340" value="2005-04-08">
<collectingDate id="8449E9DE932BA505FD2FFA209EEBFA3A" pageId="7" pageNumber="340" value="2005-04-08">8 Apr 2005</collectingDate>
</date>
(listed by
<bibRefCitation id="84224B07932BA505FEA3FA009FEEFA3A" author="Dathe H. H." box="[313,440,1460,1484]" pageId="7" pageNumber="340" pagination="335 - 432" refId="ref8181" refString="Dathe H. H. 2009. Order Hymenoptera, superfamily Apoidea. Families Colletidae, Andrenidae, Halictidae, Melittidae, Megachilidae and Apidae. Arthropod Fauna of the UAE, 2: 335 - 432." type="journal article" year="2009">Dathe 2009</bibRefCitation>
under
<taxonomicName id="27B34D75932BA505FD93FA019C38FA3A" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[521,622,1460,1484]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FD93FA019C38FA3A" box="[521,622,1460,1484]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
).
<collectorName id="4D465320932BA505FD12FA009CA2FA3A" box="[648,756,1460,1484]" pageId="7" pageNumber="340">Deposited</collectorName>
in the
<collectorName id="4D465320932BA505FF51FA609F35FA1A" box="[203,355,1492,1516]" pageId="7" pageNumber="340">A. van Harten</collectorName>
collection in the
<collectionCode id="86A2AE33932BA505FD85FA619C1DFA1A" box="[543,587,1493,1516]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="7" pageNumber="340" type="Other type of research institution/biorepository">DEI</collectionCode>
(
<specimenCode id="B0159E8D932BA505FDC1FA609CE4FA1A" box="[603,690,1492,1516]" collectionCode="DEI" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="7" pageNumber="340" type="Other type of research institution/biorepository">DEI130</specimenCode>
) (
<figureCitation id="78882A73932BA505FD53FA609ECFF9FA" captionStart="Fig" captionStartId="6.[151,182,1885,1904]" captionTargetBox="[151,1451,171,1862]" captionTargetId="figure-9@6.[151,1451,171,1862]" captionTargetPageId="6" captionText="Fig. 9 – Pseudoanthidium guichardi Pasteels, 1980. Male. A, Holotype from Oman (NHMUK); B-H, Male from United Arab Emirates (DEI); A-B, Habitus ventral; C-D, Head and mesosoma in dorsal view; E, Head in frontal view; F, Apical terga; G, Habitus lateral; H, Apex of metasoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.12763484" httpUri="https://zenodo.org/record/12763484/files/figure.png" pageId="7" pageNumber="340">Fig. 9</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="50DB3CAB932BA505FF31FA409FE4F9DA" collectingDate="2015-03-09" collectionCode="S, OLL" collectorName="M. Snizek" country="Oman" location="Dank" pageId="7" pageNumber="340" specimenCount="1" specimenCount-female="1">
<emphasis id="D2C7EAE4932BA505FF31FA409EA9F9FA" bold="true" box="[171,255,1524,1548]" pageId="7" pageNumber="340">
<collectingCountry id="98A47666932BA505FF31FA409EA9F9FA" box="[171,255,1524,1548]" name="Oman" pageId="7" pageNumber="340">OMAN</collectingCountry>
</emphasis>
:
<specimenCount id="F6B5FD7F932BA505FE97FA409F78F9FA" box="[269,302,1524,1548]" count="1" pageId="7" pageNumber="340" type="female">1♀</specimenCount>
,
<locationDeviation id="F1BE2FD4932BA505FEA1FA419FE4F9FA" box="[315,434,1524,1548]" location="Ibri" pageId="7" pageNumber="340">NW of Ibri</locationDeviation>
,
<collectionCode id="86A2AE33932BA505FE27FA409F9AF9FA" box="[445,460,1524,1548]" pageId="7" pageNumber="340">S</collectionCode>
of
<location id="E56C602D932BA505FE74FA409C7CF9FA" LSID="urn:lsid:plazi:treatment:681A87E0932BA50AFF12FB009D55F9DA:E56C602D932BA505FE74FA409C7CF9FA" box="[494,554,1524,1548]" country="Oman" name="Dank" pageId="7" pageNumber="340">Dank</location>
,
<date id="940D1036932BA505FDAEFA409C9DF9FA" box="[564,715,1524,1548]" pageId="7" pageNumber="340" value="2015-03-09">
<collectingDate id="8449E9DE932BA505FDAEFA409C9DF9FA" box="[564,715,1524,1548]" pageId="7" pageNumber="340" value="2015-03-09">9 March 2015</collectingDate>
</date>
,
<collectorName id="4D465320932BA505FD4CFA419E87F9DA" pageId="7" pageNumber="340">M. Snižek</collectorName>
leg. (
<figureCitation id="78882A73932BA505FE97F9A09F0CF9DA" box="[269,346,1556,1580]" captionStart="Fig" captionStartId="7.[137,169,710,729]" captionTargetBox="[136,756,180,690]" captionTargetId="figure-776@7.[136,759,177,690]" captionTargetPageId="7" captionText="Fig. 10 – Pseudoanthidium guichardi Pasteels, 1980. Female from Oman (OLL)." figureDoi="http://doi.org/10.5281/zenodo.12763488" httpUri="https://zenodo.org/record/12763488/files/figure.png" pageId="7" pageNumber="340">Fig. 10</figureCitation>
) (
<collectionCode id="86A2AE33932BA505FEEBF9A09FFCF9DA" box="[369,426,1556,1580]" pageId="7" pageNumber="340">OLL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E00C36F6932BA505FF34F9809C50F91A" blockId="7.[135,760,1236,1932]" pageId="7" pageNumber="340">
<emphasis id="D2C7EAE4932BA505FF34F9809FE0F9BA" box="[174,438,1588,1612]" italics="true" pageId="7" pageNumber="340">Material not examined</emphasis>
.
<emphasis id="D2C7EAE4932BA505FE54F9809E9AF99A" bold="true" pageId="7" pageNumber="340">
<collectingCountry id="98A47666932BA505FE54F9809E9AF99A" name="United Arab Emirates" pageId="7" pageNumber="340">UNITED ARAB EMIRATES</collectingCountry>
</emphasis>
:
<specimenCount id="F6B5FD7F932BA505FF40F9E09F54F99A" box="[218,258,1620,1644]" count="1" pageId="7" pageNumber="340" type="female">1 ♀</specimenCount>
, Fagsha,
<collectingRegion id="2277F814932BA505FEF2F9E09F92F99A" box="[360,452,1620,1644]" country="United Arab Emirates" name="Al Fujayrah" pageId="7" pageNumber="340">Fujairah</collectingRegion>
,
<date id="940D1036932BA505FE54F9E09C00F99A" box="[462,598,1620,1644]" pageId="7" pageNumber="340" value="1988-04-21">21 Apr 1988</date>
, I. Hamer leg., D. Baker det. (Snow Entomological Collection, Ohio), www.gbif.org/occurrence/785011857 (also listed by
<bibRefCitation id="84224B07932BA505FD2EF9209E97F93A" author="Dathe H. H." pageId="7" pageNumber="340" pagination="335 - 432" refId="ref8181" refString="Dathe H. H. 2009. Order Hymenoptera, superfamily Apoidea. Families Colletidae, Andrenidae, Halictidae, Melittidae, Megachilidae and Apidae. Arthropod Fauna of the UAE, 2: 335 - 432." type="journal article" year="2009">Dathe 2009</bibRefCitation>
). This material is listed under
<taxonomicName id="27B34D75932BA505FD92F9019C3CF93A" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[520,618,1716,1740]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FD92F9019C3CF93A" box="[520,618,1716,1740]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
and is tentatively assigned here to
<taxonomicName id="27B34D75932BA505FEE4F9619FA9F91A" authorityName="Pasteels" authorityYear="1980" box="[382,511,1748,1772]" class="Insecta" family="Megachilidae" genus="Exanthidium" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="guichardi">
<emphasis id="D2C7EAE4932BA505FEE4F9619FA9F91A" box="[382,511,1748,1772]" italics="true" pageId="7" pageNumber="340">P. guichardi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E00C36F6932BA505FF34F9409B73FEDA" blockId="7.[135,760,1236,1932]" lastBlockId="7.[815,1445,180,1932]" pageId="7" pageNumber="340">
Structurally similar to
<taxonomicName id="27B34D75932BA505FE3AF9419C54F8FA" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[416,514,1780,1804]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FE3AF9419C54F8FA" box="[416,514,1780,1804]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
, but entire integument light yellow except for scutellum and face (dominant ground colour in
<taxonomicName id="27B34D75932BA505FED9F8819FF2F8BA" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[323,420,1844,1868]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FED9F8819FF2F8BA" box="[323,420,1844,1868]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
black). Yellow tergal bands interrupted in the middle in
<taxonomicName id="27B34D75932BA505FE2BF8E19C40F89A" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[433,534,1876,1900]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FE2BF8E19C40F89A" box="[433,534,1876,1900]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
, uninterrupted in
<taxonomicName id="27B34D75932BA505FD7AF8E19EB9F87A" authorityName="Pasteels" authorityYear="1980" class="Insecta" family="Megachilidae" genus="Exanthidium" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="guichardi">
<emphasis id="D2C7EAE4932BA505FD7AF8E19EB9F87A" italics="true" pageId="7" pageNumber="340">P. guichardi</emphasis>
</taxonomicName>
. In
<taxonomicName id="27B34D75932BA505FE81F8C19FCAF87A" authorityName="Pasteels" authorityYear="1980" box="[283,412,1908,1932]" class="Insecta" family="Megachilidae" genus="Exanthidium" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="guichardi">
<emphasis id="D2C7EAE4932BA505FE81F8C19FCAF87A" box="[283,412,1908,1932]" italics="true" pageId="7" pageNumber="340">P. guichardi</emphasis>
</taxonomicName>
, the tergal discs have an almost step-like elevation over the depressions with a clear, albeit somewhat jagged border line (best seen medially on T2- T4). By contrast, the transition from the disc to the depression is inconspicuous and smooth in
<taxonomicName id="27B34D75932BA505FB23FEA09B48FEDA" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[1209,1310,276,300]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FB23FEA09B48FEDA" box="[1209,1310,276,300]" italics="true" pageId="7" pageNumber="340">A. enslini</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E00C36F6932BA505FCCCFE809B62FD9A" blockId="7.[815,1445,180,1932]" pageId="7" pageNumber="340">
<emphasis id="D2C7EAE4932BA505FCCCFE809DDBFEBA" box="[854,909,308,332]" italics="true" pageId="7" pageNumber="340">Male</emphasis>
(
<figureCitation id="78882A73932BA505FC05FE809DB4FEBA" box="[927,994,308,332]" captionStart="Fig" captionStartId="6.[151,182,1885,1904]" captionTargetBox="[151,1451,171,1862]" captionTargetId="figure-9@6.[151,1451,171,1862]" captionTargetPageId="6" captionText="Fig. 9 – Pseudoanthidium guichardi Pasteels, 1980. Male. A, Holotype from Oman (NHMUK); B-H, Male from United Arab Emirates (DEI); A-B, Habitus ventral; C-D, Head and mesosoma in dorsal view; E, Head in frontal view; F, Apical terga; G, Habitus lateral; H, Apex of metasoma in ventral view." figureDoi="http://doi.org/10.5281/zenodo.12763484" httpUri="https://zenodo.org/record/12763484/files/figure.png" pageId="7" pageNumber="340">Fig. 9</figureCitation>
).
<quantity id="274B9B13932BA505FC60FE819A68FEBA" box="[1018,1086,309,332]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="340" unit="mm" value="7.0">7 mm</quantity>
.
<emphasis id="D2C7EAE4932BA505FBD1FE809AD8FEBA" box="[1099,1166,308,332]" italics="true" pageId="7" pageNumber="340">Head:</emphasis>
Light yellow except for black in the upper face from above antennal sockets to hind ocelli; yellow spot below anterior ocellus (present in
<collectionCode id="86A2AE33932BA505FAF4FEC19BCCFE7A" box="[1390,1434,373,396]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="7" pageNumber="340" type="Other type of research institution/biorepository">DEI</collectionCode>
specimen, absent in
<typeStatus id="3F088854932BA505FB8AFE209A24FE5A" box="[1040,1138,404,428]" pageId="7" pageNumber="340" type="holotype">holotype</typeStatus>
); clypeus bell-shaped; head densely punctate throughout except for marginal areas of clypeus which are smooth; dense, long, silvery, adpressed hair covering face; vertex and gena almost hairless; mandible pale yellow, sparsely punctate, with three black teeth; scape yellow; flagella reddish-brown, upper side of distal flagella grey-brown; preoccipital ridge angulate.
</paragraph>
<paragraph id="E00C36F6932BA505FCCCFDC19BF3FC7A" blockId="7.[815,1445,180,1932]" pageId="7" pageNumber="340">
<emphasis id="D2C7EAE4932BA505FCCCFDC19D87FD7A" box="[854,977,629,652]" italics="true" pageId="7" pageNumber="340">
<taxonomicName id="27B34D75932BA505FCCCFDC19D9DFD7A" box="[854,971,629,652]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="340" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
Light yellow; scutum black with anterolateral yellow stripe in
<typeStatus id="3F088854932BA505FB92FD209A3EFD5A" box="[1032,1128,660,684]" pageId="7" pageNumber="340" type="holotype">holotype</typeStatus>
, but in
<collectionCode id="86A2AE33932BA505FB2EFD219AB6FD5A" box="[1204,1248,661,684]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="7" pageNumber="340" type="Other type of research institution/biorepository">DEI</collectionCode>
specimen, anteri- or third entirely reddish brown and laterally yellow; pronotal lobe with high lamella; scutellum black anteromedially (
<typeStatus id="3F088854932BA505FCADFD409DCEFCFA" box="[823,920,756,780]" pageId="7" pageNumber="340" type="holotype">holotype</typeStatus>
) and yellow posteriorly (entirely yellow with only some inconspicuous reddish tone anteromedially in
<collectionCode id="86A2AE33932BA505FCAAFC819D0AFCBA" box="[816,860,821,844]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="7" pageNumber="340" type="Other type of research institution/biorepository">DEI</collectionCode>
specimen); posteromedially slightly depressed; posterior margin transparent, jagged; widely rounded in dorsal view; sharp-edged in lateral view; axilla triangular, yellow.
</paragraph>
<paragraph id="E00C36F6932BA505FCCCFC219B21FADA" blockId="7.[815,1445,180,1932]" pageId="7" pageNumber="340">
<emphasis id="D2C7EAE4932BA505FCCCFC219D98FC5A" box="[854,974,917,940]" italics="true" pageId="7" pageNumber="340">Metasoma:</emphasis>
Yellow except for the declivous anterior part of T1 which is black in the
<typeStatus id="3F088854932BA505FB16FC009ABFFC3A" box="[1164,1257,948,972]" pageId="7" pageNumber="340" type="holotype">holotype</typeStatus>
and light brown in the
<collectionCode id="86A2AE33932BA505FCECFC619DF4FC1A" box="[886,930,981,1004]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="7" pageNumber="340" type="Other type of research institution/biorepository">DEI</collectionCode>
specimen; tergal bands on discs unbroken; depressions yellowish-brown; marginal areas smooth, transparent and slightly turned upwards; discs conspicuously elevated over depressions, the two areas separated by a distinct jagged line; punctation on discs dense and coarse, on depressions dense and fine; T4 bulging laterally, T5 with blunt tooth laterally, T6 with flat, transparent hooklike tooth laterally; T6 bulging mediolaterally; T7 long with deep median emargination; margins broadly transparent; S6 laterally with strongly sclerotised wings; apex thin and transparent, bilobed; apex of (hidden) S8 truncate.
</paragraph>
<paragraph id="E00C36F6932BA505FCCCFA809B27F9DA" blockId="7.[815,1445,180,1932]" pageId="7" pageNumber="340">
<emphasis id="D2C7EAE4932BA505FCCCFA809DF0FABA" box="[854,934,1332,1356]" italics="true" pageId="7" pageNumber="340">Female</emphasis>
(
<figureCitation id="78882A73932BA505FC22FA809A5DFABA" box="[952,1035,1332,1356]" captionStart="Fig" captionStartId="7.[137,169,710,729]" captionTargetBox="[136,756,180,690]" captionTargetId="figure-776@7.[136,759,177,690]" captionTargetPageId="7" captionText="Fig. 10 – Pseudoanthidium guichardi Pasteels, 1980. Female from Oman (OLL)." figureDoi="http://doi.org/10.5281/zenodo.12763488" httpUri="https://zenodo.org/record/12763488/files/figure.png" pageId="7" pageNumber="340">Fig. 10</figureCitation>
). Structurally very similar to the female of
<emphasis id="D2C7EAE4932BA505FC1FFAE19DBAFA9A" box="[901,1004,1364,1388]" italics="true" pageId="7" pageNumber="340">
<taxonomicName id="27B34D75932BA505FC1FFAE19DBEFA9A" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[901,1000,1364,1388]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">P. enslini</taxonomicName>
,
</emphasis>
but distinguished by colouration: almost entirely yellow and light reddish-brown except for black maculation around ocelli and between ocelli and antennal sockets, and some black remnants at anterior and posterior margin of scutum. Face and vertex light reddish-brown, lower face including clypeus, and gena yellow; T1-T5 with broad yellow bands and light brown depressions.
</paragraph>
<paragraph id="E00C36F6932BA50AFCCCF9809FE2FA5A" blockId="7.[815,1445,180,1932]" lastBlockId="8.[151,772,1364,1580]" lastPageId="8" lastPageNumber="341" pageId="7" pageNumber="340">
<emphasis id="D2C7EAE4932BA505FCCCF9809DECF9BA" box="[854,954,1588,1612]" italics="true" pageId="7" pageNumber="340">Remarks.</emphasis>
The species is distinguished from
<taxonomicName id="27B34D75932BA505FAA3F9819BCAF9BA" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[1337,1436,1588,1612]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FAA3F9819BCAF9BA" box="[1337,1436,1588,1612]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
according to
<bibRefCitation id="84224B07932BA505FC24F9E09A34F99A" author="Pasteels J. J." box="[958,1122,1620,1644]" pageId="7" pageNumber="340" pagination="91 - 98" refId="ref8834" refString="Pasteels J. J. 1980. Notes sur les Megachilidae (Hymenoptera, Apoidea) d'Arabie et des regions desertiques limitrophes. 2. Especes nouvelles d'Anthidiinae. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 116: 91 - 98." type="journal article" year="1980">Pasteels (1980)</bibRefCitation>
by a bilobed
<collectionCode id="86A2AE33932BA505FB63F9E09B5EF99A" box="[1273,1288,1620,1644]" pageId="7" pageNumber="340">S</collectionCode>
6, a relatively broad gena and a relatively long distance of the hind ocellus from the preoccipital ridge. However, both species have a bilobed
<collectionCode id="86A2AE33932BA505FC3AF9009DF9F93A" box="[928,943,1716,1740]" pageId="7" pageNumber="340">S</collectionCode>
6, only slightly less emarginate in
<taxonomicName id="27B34D75932BA505FAABF9019BC3F93A" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[1329,1429,1716,1740]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FAABF9019BC3F93A" box="[1329,1429,1716,1740]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
. Also the difference of the width of the gena as compared to the width of the eye is moderate and the difference in the postocelli-preoccipital distance could not be confirmed when compared with
<taxonomicName id="27B34D75932BA505FB83F8819A2CF8BA" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[1049,1146,1844,1868]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="340" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE4932BA505FB83F8819A2CF8BA" box="[1049,1146,1844,1868]" italics="true" pageId="7" pageNumber="340">P. enslini</emphasis>
</taxonomicName>
<typeStatus id="3F088854932BA505FB1BF8809AB7F8BA" box="[1153,1249,1844,1868]" pageId="7" pageNumber="340" type="holotype">holotype</typeStatus>
. These traits may thus fall into individual variation and do not have diagnostic value. Also additional morphological differences such as the presence of a lateral bulge on T6 vs. a blunt lateral tooth in
<taxonomicName id="27B34D759324A50AFF76FAC19F1CFA7A" baseAuthorityName="Alfken" baseAuthorityYear="1928" box="[236,330,1396,1420]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="D2C7EAE49324A50AFF76FAC19F1CFA7A" box="[236,330,1396,1420]" italics="true" pageId="8" pageNumber="341">P. enslini</emphasis>
</taxonomicName>
cannot be used as specific characters until the full variation is known.
</paragraph>
<caption id="B4CC667E9324A50AFF0DFCA99C92FCBD" ID-DOI="http://doi.org/10.5281/zenodo.12763490" ID-Zenodo-Dep="12763490" httpUri="https://zenodo.org/record/12763490/files/figure.png" pageId="8" pageNumber="341" startId="8.[151,181,797,816]" targetBox="[151,771,179,774]" targetPageId="8" targetType="figure">
<paragraph id="E00C36F69324A50AFF0DFCA99C92FCBD" blockId="8.[150,772,797,843]" pageId="8" pageNumber="341">
<emphasis id="D2C7EAE49324A50AFF0DFCA99E88FCC6" bold="true" box="[151,222,797,816]" pageId="8" pageNumber="341">Fig. 11 –</emphasis>
<taxonomicName id="27B34D759324A50AFF7AFCA99FE7FCC6" baseAuthorityName="Morawitz" baseAuthorityYear="1894" box="[224,433,797,816]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="pictipes">
<emphasis id="D2C7EAE49324A50AFF7AFCA99FE7FCC6" box="[224,433,797,816]" italics="true" pageId="8" pageNumber="341">Pseudoanthidium pictipes</emphasis>
</taxonomicName>
. Male holotype (ZISP).
<emphasis id="D2C7EAE49324A50AFDEAFCA99C29FCC6" bold="true" box="[624,639,797,816]" pageId="8" pageNumber="341">A</emphasis>
, Habitus dorsal;
<emphasis id="D2C7EAE49324A50AFF0CFC8C9EF2FCBD" bold="true" box="[150,164,824,843]" pageId="8" pageNumber="341">B</emphasis>
, Head in frontal view;
<emphasis id="D2C7EAE49324A50AFEC7FC8C9F3AFCBD" bold="true" box="[349,364,824,843]" pageId="8" pageNumber="341">C</emphasis>
, Apical terga. Photographs:Y.Astafurova.
</paragraph>
</caption>
<caption id="B4CC667E9324A50AFF0DFB4C9C09FAD0" ID-DOI="http://doi.org/10.5281/zenodo.12763492" ID-Zenodo-Dep="12763492" httpUri="https://zenodo.org/record/12763492/files/figure.png" pageId="8" pageNumber="341" startId="8.[151,182,1272,1291]" targetBox="[151,771,871,1250]" targetPageId="8" targetType="figure">
<paragraph id="E00C36F69324A50AFF0DFB4C9C09FAD0" blockId="8.[150,771,1272,1318]" pageId="8" pageNumber="341">
<emphasis id="D2C7EAE49324A50AFF0DFB4C9EB5FAFD" bold="true" box="[151,227,1272,1291]" pageId="8" pageNumber="341">Fig. 12 –</emphasis>
<taxonomicName id="27B34D759324A50AFF7CFB4C9F94FAFC" baseAuthorityName="Morawitz" baseAuthorityYear="1894" box="[230,450,1272,1291]" class="Insecta" family="Megachilidae" genus="Pseudoanthidium" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="341" phylum="Arthropoda" rank="species" species="pictipes">
<emphasis id="D2C7EAE49324A50AFF7CFB4C9F94FAFC" box="[230,450,1272,1291]" italics="true" pageId="8" pageNumber="341">Pseudoanthidium pictipes</emphasis>
</taxonomicName>
. Specialised hairs with hooked apices on S3 of the male. Photograph: Y.Astafurova (ZISP).
</paragraph>
</caption>
<paragraph id="E00C36F69324A50AFF27FA009C11FA1A" blockId="8.[151,772,1364,1580]" pageId="8" pageNumber="341">
Attempts to retrieve DNA from the
<collectingCountry id="98A476669324A50AFDA1FA009C24FA3A" box="[571,626,1460,1484]" name="United Arab Emirates" pageId="8" pageNumber="341">UAE</collectingCountry>
specimen for a genetic barcoding were not successful.
</paragraph>
<paragraph id="E00C36F69324A50AFF27FA409D55F9DA" blockId="8.[151,772,1364,1580]" pageId="8" pageNumber="341">
<emphasis id="D2C7EAE49324A50AFF27FA409F11F9FA" box="[189,327,1524,1548]" italics="true" pageId="8" pageNumber="341">Distribution.</emphasis>
Eastern Arabian Peninsula around the Gulf of
<collectingCountry id="98A476669324A50AFF73F9A09F7CF9DA" box="[233,298,1556,1580]" name="Oman" pageId="8" pageNumber="341">Oman</collectingCountry>
(
<collectingCountry id="98A476669324A50AFEADF9A09F2CF9DA" box="[311,378,1556,1580]" name="Oman" pageId="8" pageNumber="341">Oman</collectingCountry>
and
<collectingCountry id="98A476669324A50AFE36F9A09CC2F9DA" box="[428,660,1556,1580]" name="United Arab Emirates" pageId="8" pageNumber="341">United Arab Emirates</collectingCountry>
) (
<figureCitation id="78882A739324A50AFD32F9A09CA0F9DA" box="[680,758,1556,1580]" captionStart="Fig" captionStartId="10.[150,181,833,852]" captionTargetBox="[151,1451,166,816]" captionTargetId="figure-460@10.[151,1451,166,816]" captionTargetPageId="10" captionText="Fig. 15 – Distribution of Pseudoanthidium deserticolum sp.n., P. enslini, P. pictipes, and P. wahrmanicum." figureDoi="http://doi.org/10.5281/zenodo.12763500" httpUri="https://zenodo.org/record/12763500/files/figure.png" pageId="8" pageNumber="341">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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