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<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.5.63435" ID-GBIF-Dataset="bfba5087-74d4-44e8-b783-94db25e3442a" ID-GBIF-Taxon="184941061" ID-Pensoft-Pub="2535-0730-2-193" ID-Pensoft-UUID="21773559522D5D9DA4B1A7DA51E4F2A6" ID-ZooBank="984E15D880E04B7DA84F92BB0AD4EA73" ModsDocID="2535-0730-5-2-193" checkinTime="1629846914602" checkinUser="pensoft" docAuthor="Zahiri, Reza, Nazari, Vazrick, Rajaei, Hossein, Wiemers, Martin, Fatahi, Maryam, Seidel, Matthias, Dalsgaard, Thure &amp; Husemann, Martin" docDate="2021" docId="674F5F9AE73B50208B76FC7A29A0B151" docLanguage="en" docName="EvolutSyst 5(2): 193-261" docOrigin="Evolutionary Systematics 5 (2)" docPubDate="2021-08-20" docSource="http://dx.doi.org/10.3897/evolsyst.5.63435" docTitle="Melitaea ignasiti de Sagarra 1926" docType="treatment" docVersion="3" id="21773559522D5D9DA4B1A7DA51E4F2A6" lastPageNumber="193" masterDocId="21773559522D5D9DA4B1A7DA51E4F2A6" masterDocTitle="? An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH): Part II. superfamily Papilionoidea" masterLastPageNumber="261" masterPageNumber="193" pageId="0" pageNumber="193" updateTime="1643429667741" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>? An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH): Part II. superfamily Papilionoidea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zahiri, Reza</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6274-6973</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany &amp; Canadian Food Inspection Agency, Ottawa Plant Laboratory, Entomology Unit, Bldg. 18, 960 Carling Ave., K 1 A 0 C 6, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nazari, Vazrick</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9064-8959</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rajaei, Hossein</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3940-3734</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiemers, Martin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5272-3903</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fatahi, Maryam</mods:namePart>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4913-8778</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dalsgaard, Thure</mods:namePart>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Husemann, Martin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5536-6681</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>193</mods:start>
<mods:end>261</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/evolsyst.5.63435</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.5.63435</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0730-2-193</mods:identifier>
<mods:identifier type="ZooBank">984E15D880E04B7DA84F92BB0AD4EA73</mods:identifier>
<mods:identifier type="Pensoft-UUID">21773559522D5D9DA4B1A7DA51E4F2A6</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="184941061" LSID="urn:lsid:plazi:treatment:674F5F9AE73B50208B76FC7A29A0B151" httpUri="http://treatment.plazi.org/id/674F5F9AE73B50208B76FC7A29A0B151" lastPageNumber="193" pageId="0" pageNumber="193">
<subSubSection pageId="0" pageNumber="193" type="nomenclature">
<paragraph pageId="0" pageNumber="193">
?117.
<taxonomicName LSID="674F5F9A-E73B-5020-8B76-FC7A29A0B151" authority="de Sagarra, 1926" authorityName="de Sagarra" authorityYear="1926" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea ignasiti" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="ignasiti">Melitaea ignasiti de Sagarra, 1926</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="original combination">
<paragraph pageId="0" pageNumber="193">Original combination.</paragraph>
<paragraph pageId="0" pageNumber="193">
&quot;
<taxonomicName authority="ignasiti" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea trivia subsp. rassa" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="trivia" subSpecies="rassa">Melitaea trivia Schiff., rassa ignasiti</taxonomicName>
, nova rassa&quot; de Sagarra, 1926 Butl. Inst. Catal. Hist. Nat. (2)6: 131.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="current combination">
<paragraph pageId="0" pageNumber="193">Current combination.</paragraph>
<paragraph pageId="0" pageNumber="193">
<emphasis bold="true" pageId="0" pageNumber="193">
<taxonomicName authorityName="Sagarra" authorityYear="1926" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea trivia subsp. ignasiti" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="trivia" subSpecies="ignasiti">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea trivia ignasiti</emphasis>
</taxonomicName>
de Sagarra, 1926
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="current status">
<paragraph pageId="0" pageNumber="193">Current status.</paragraph>
<paragraph pageId="0" pageNumber="193">Valid subspecies.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="type material">
<paragraph pageId="0" pageNumber="193">Type material.</paragraph>
<paragraph pageId="0" pageNumber="193">
Paralectotypes 1?1? (ZMH 827660-827661) (Fig.
<figureCitation captionStart="Plate 16" captionStartId="F16" captionText="Plate 16. 110. Melitaea athalia tenuicula Verity, 1919; 111. Melitaea varia varissima Verity, 1914; 112. Melitaea cinxia riloensis Zuellich, 1936; 113. Melitaea deione ab. mediofasciata Bubacek, 1926; 114. Melitaea diamina ab. johni Stauder, 1922; 115. Melitaea trivia balcanica Zuellich, 1936; 116. Melitaea trivia catapelioides Stauder, 1918; 117. Melitaea trivia ignasiti de Sagarra, 1926. a. Dorsal view, b. ventral view." httpUri="https://binary.pensoft.net/fig/579725" pageId="0" pageNumber="193">117</figureCitation>
). &quot;CATALONIA / Seva 700 m. / 3.8.25 Romei&quot; // &quot;
<taxonomicName genus="M." lsidName="M. trivia" pageId="0" pageNumber="193" rank="species" species="trivia">
<emphasis italics="true" pageId="0" pageNumber="193">M. trivia</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="de Sagarra" authorityYear="1926" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea ignasiti" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="ignasiti">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea ignasiti</emphasis>
</taxonomicName>
Sag. / co-type? Querci&quot; // &quot;ZMH 827660&quot;; &quot;CATALONIA / Seva 700 m. / 3.8.25 Romei&quot; // &quot;Sammlung / R.
<normalizedToken originalValue="Jähnig">Jaehnig</normalizedToken>
/ Eing. Nr.29,1949&quot; // &quot;
<taxonomicName genus="M." lsidName="M. trivia" pageId="0" pageNumber="193" rank="species" species="trivia">
<emphasis italics="true" pageId="0" pageNumber="193">M. trivia</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="de Sagarra" authorityYear="1926" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea ignasiti" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="ignasiti">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea ignasiti</emphasis>
</taxonomicName>
Sag. / co-type? Querci&quot; //
<normalizedToken originalValue="“L.28”">&quot;L.28&quot;</normalizedToken>
// &quot;ZMH 827661&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="type locality">
<paragraph pageId="0" pageNumber="193">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="193">
Spain: Catalonia,
<normalizedToken originalValue="“Martorell”">&quot;Martorell&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="remarks">
<paragraph pageId="0" pageNumber="193">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="193">
<bibRefCitation author="de Sagarra, I" journalOrPublisher="Mitteilungen der Muenchner Entomologischen Gesellschaft" pageId="0" pageNumber="193" refId="B21" refString="de Sagarra, I, 1926. Anotacions a la lepidopterologia ibe'rica IV. Butlleti de la Institucio Catalana d'Historia Natural (2)6: 128-139." title="Anotacions a la lepidopterologia ibe'rica IV. Butlleti de la Institucio Catalana d'Historia Natural (2) 6: 128 - 139." year="1926">de Sagarra (1926)</bibRefCitation>
proposed this name as a &quot;rassa = race&quot; of
<taxonomicName genus="M." lsidName="M. trivia" pageId="0" pageNumber="193" rank="species" species="trivia">
<emphasis italics="true" pageId="0" pageNumber="193">M. trivia</emphasis>
</taxonomicName>
([Denis &amp; Schiffermullar], 1775). The term &quot;rassa = race&quot; which was employed by de Sagarra in 1926 was often used in the current sense of subspecies. As a result, the first line of article 45.6 (
<bibRefCitation author="ICZN" journalOrPublisher="Internationale Entomologische Zeitschrift" pageId="0" pageNumber="193" publicationUrl="http://www.iczn.org/iczn/index.jsp" refId="B47" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth edition. London. [available online at http://www.iczn.org/iczn/index.jsp" title="International Code of Zoological Nomenclature. Fourth edition. London. [available online at" url="http://www.iczn.org/iczn/index.jsp" year="1999">ICZN 1999</bibRefCitation>
) - the rank denoted by a species-group name following a binomen is subspecific - is applied here, it is thus available as subspecific.
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1941.tb01045.x" author="Higgins, LG" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="193" pagination="175 - 365" refId="B42" refString="Higgins, LG, 1941. An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London 91 (7): 175 - 365, DOI: https://doi.org/10.1111/j.1365-2311.1941.tb01045.x" title="An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera)." url="https://doi.org/10.1111/j.1365-2311.1941.tb01045.x" volume="91" year="1941">Higgins (1941)</bibRefCitation>
synonymised it with
<taxonomicName genus="M." lsidName="M. trivia subsp. pseudodidyma" pageId="0" pageNumber="193" rank="subSpecies" species="trivia" subSpecies="pseudodidyma">
<emphasis italics="true" pageId="0" pageNumber="193">M. trivia pseudodidyma</emphasis>
</taxonomicName>
Rebel, 1905 but he reconsidered it as a valid subspecies in his subsequent books.
<bibRefCitation author="Garcia-Barros, E" journalOrPublisher="Museo Nacional de Ciencias Naturales, CSIC. Madrid" pageId="0" pageNumber="193" refId="B31" refString="Garcia-Barros, E, Munguira, M, Stefanescu, C, Vives, MA, 2013. . LepidopteraPapilionoidea. In: Ramos MA et al. (Eds) Fauna Iberica, vol. 37. Museo Nacional de Ciencias Naturales, CSIC. Madrid" title=". LepidopteraPapilionoidea. In: Ramos MA et al. (Eds) Fauna Iberica, vol. 37." year="2013">
<normalizedToken originalValue="García-Barros">Garcia-Barros</normalizedToken>
et al. (2013)
</bibRefCitation>
made a comment in Fauna
<normalizedToken originalValue="Ibérica">Iberica</normalizedToken>
that &quot;the orange color of the Iberian specimens is lighter than those from other European areas; those have been attributed to subspecies &quot;
<taxonomicName authorityName="de Sagarra" authorityYear="1926" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea ignasiti" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="ignasiti">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea ignasiti</emphasis>
</taxonomicName>
de Sagarra, 1926&quot;, which, however, does not seem significantly different. Furthermore, it does not seem significantly differentiated from a genetic point of view (Wahlberg pers. comm.).&quot; The taxon
<taxonomicName authorityName="de Sagarra" authorityYear="1926" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea ignasiti" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="ignasiti">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea ignasiti</emphasis>
</taxonomicName>
was reinstated as a valid species by
<bibRefCitation author="van Oorschot, H" journalOrPublisher="Tshikolovets Publications, Pardubice" pageId="0" pageNumber="193" refId="B93" refString="van Oorschot, H, Coutsis, JG, 2014. The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae). Taxonomy and systematics with special reference to the male genitalia. Tshikolovets Publications, Pardubice" title="The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae). Taxonomy and systematics with special reference to the male genitalia." year="2014">van Oorschot and Coutsis (2014)</bibRefCitation>
, and this was accepted by Macia et al. (2015) who designated lectotype and paralectotypes. Therefore, specimens at ZMH are by default paralectotypes. However, most other recent authors (e.g.,
<bibRefCitation author="Kudrna, O" journalOrPublisher="Spolecnost pro Ochranu Motylu, Prachatice, CZ" pageId="0" pageNumber="193" refId="B57" refString="Kudrna, O, 2019. Distribution of Butterflies and Skippers in Europe (Lepidoptera: Rhopalocera, Grypocera). 24 years Mapping European Butterflies (1995-2019), Final Report. Spolecnost pro Ochranu Motylu, Prachatice, CZ" title="Distribution of Butterflies and Skippers in Europe (Lepidoptera: Rhopalocera, Grypocera). 24 years Mapping European Butterflies (1995 - 2019), Final Report." year="2019">Kudrna 2019</bibRefCitation>
) treat this taxon as a synonym to
<taxonomicName genus="M." lsidName="M. trivia" pageId="0" pageNumber="193" rank="species" species="trivia">
<emphasis italics="true" pageId="0" pageNumber="193">M. trivia</emphasis>
</taxonomicName>
, because there is no evidence for species status. Even Oorschot and Coutsis (2014) mention intermediates indicating hybridisation. It should be noted that the species name is not mentioned in the latest European checklist (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.811.28712" author="Wiemers, M" journalOrPublisher="ZooKeys" pageId="0" pageNumber="193" pagination="9 - 45" refId="B118" refString="Wiemers, M, Balletto, E, Dinca, V, Fric, ZF, Lamas, G, Lukhtanov, V, Munguira, ML, van Swaay, CAM, Vila, R, Vliegenthart, A, Wahlberg, N, Verovnik, R, 2018. An updated checklist of the European Butterflies (Lepidoptera, Papilionoidea). ZooKeys 811: 9 - 45, DOI: https://doi.org/10.3897/zookeys.811.28712" title="An updated checklist of the European Butterflies (Lepidoptera, Papilionoidea)." url="https://doi.org/10.3897/zookeys.811.28712" volume="811" year="2018">Wiemers et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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