161 lines
20 KiB
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161 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.70.107322" ID-Pensoft-Pub="1860-1324-2-291" ID-Pensoft-UUID="DD34414053665BD69D8E11CB57724004" ID-ZooBank="52692B916EE24AA0B9D639BDF3107D92" ModsDocID="1860-1324-70-2-291" checkinTime="1692351740246" checkinUser="pensoft" docAuthor="Sasakawa, Koji & Mitsuduka, Yoshiji" docDate="2023" docId="568F64789EBD5E9B963E70300D9C2638" docLanguage="en" docName="DeutEntomolZeit 70(2): 291-310" docOrigin="Deutsche Entomologische Zeitschrift 70 (2)" docPubDate="2023-08-17" docSource="http://dx.doi.org/10.3897/dez.70.107322" docTitle="Pterostichus (Nialoe) kamurosanus Sasakawa & Mitsuduka 2023, sp. nov." docType="treatment" docUuid="EC64C14A-C5A7-4161-BA26-77958C62BD19" docUuidSource="ZooBank" docVersion="1" id="DD34414053665BD69D8E11CB57724004" lastPageNumber="291" masterDocId="DD34414053665BD69D8E11CB57724004" masterDocTitle="A morphology-based revision and phylogenetic analysis of the Pterostichus macrogenys species group (Coleoptera, Carabidae) and implications for differentiation of the species group" masterLastPageNumber="310" masterPageNumber="291" pageNumber="291" updateTime="1692351740246" updateUser="pensoft">
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<mods:titleInfo id="AA7805917EFDCBD51644619024F3B4A0">
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<mods:title id="BAB6E854AF366369679C38A41156D73B">A morphology-based revision and phylogenetic analysis of the Pterostichus macrogenys species group (Coleoptera, Carabidae) and implications for differentiation of the species group</mods:title>
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<mods:name id="1888C1F3D4C01A1194BC131C62CF5067" type="personal">
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<mods:namePart id="2320BD9AC14AEFA89C4157F59F648842">Sasakawa, Koji</mods:namePart>
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<mods:affiliation id="FC18FD389EF74D8A300638DFE5C58045">Laboratory of Zoology, Department of Science Education, Faculty of Education, Chiba University, 1 - 33 Yayoi-cho, Inage-ku, Chiba-shi, Chiba 263 - 8522, Japan</mods:affiliation>
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<mods:nameIdentifier id="06FFEFD87F23C651E21FCF5099FD9ACE" type="email">ksasa@chiba-u.jp</mods:nameIdentifier>
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<mods:namePart id="2AE966F20182BB3D0E61DE09D5B8D3CF">Mitsuduka, Yoshiji</mods:namePart>
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<mods:affiliation id="A4143349C18785ACEF050D03A7CC683D">2 - 18 - 24 Hanatate Yamagata-shi, Yamagata 990 - 0067, Japan</mods:affiliation>
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<mods:typeOfResource id="FE9CCF04AD41BAD296BCE279290E53FE">text</mods:typeOfResource>
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<mods:title id="24B6C45E3924891D043EBBBA375C97C5">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part id="43E02CB7C24A7711DE2BEAF8F5EFCD93">
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<mods:date id="FD891D107E801FE9A40854066C8544C6">2023</mods:date>
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<mods:number id="E71FC18D26ACFDBA13189C6FCE5023AD">2023-08-17</mods:number>
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<mods:number id="180339C3D9C70B2FCB80BEA41C39329F">70</mods:number>
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<mods:number id="9FBD65F82A59914C71950732CFD2F9AE">2</mods:number>
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<subSubSection id="746C464F32EF8DE231C82989C2A6D39A" pageId="0" pageNumber="291" type="nomenclature">
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<paragraph id="711CF2905BBD6EDD8A1431265DBF4ED7" pageId="0" pageNumber="291">
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<taxonomicName id="C845DF7996D0D4883C5A51B265966FF8" LSID="https://zoobank.org/EC64C14A-C5A7-4161-BA26-77958C62BD19" authority="Sasakawa & Mitsuduka" authorityName="Sasakawa & Mitsuduka" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) kamurosanus" order="Coleoptera" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="kamurosanus" status="sp. nov." subGenus="Nialoe">Pterostichus (Nialoe) kamurosanus Sasakawa & Mitsuduka</taxonomicName>
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<taxonomicNameLabel id="74F65BFAA10EDA431EB23C2AAB11DFA2" pageId="0" pageNumber="291">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="1F9E6F38991108DBAA321055ED440C1F" pageId="0" pageNumber="291" type="description">
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<paragraph id="DF32C2E2ADD0B030AFAC05B03A2BCBAC" pageId="0" pageNumber="291">
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<figureCitation id="CA15A1CBD66451A5B293A51079F6FB89" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus dorsal view of the Pterostichus macrogenys species group. A. Pterostichus namahage sp. nov. holotype male from Haraikawa; B. P. kamurosanus sp. nov. holotype male from Ariya; C. P. atsumidakensis sp. nov. holotype male from Take-no-koshi; D. P. asahinus male from Momoyake." figureDoi="10.3897/dez.70.107322.figure2" httpUri="https://binary.pensoft.net/fig/893610" pageId="0" pageNumber="291">Figs 2B</figureCitation>
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<figureCitation id="FE6CB670CD5B1BC241B4CFC9729DC4C9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Endophallus of Pterostichus namahage sp. nov. holotype male from Haraikawa (A-D), P. kamurosanus sp. nov. holotype male from Ariya (E-H), P. atsumidakensis sp. nov. holotype male from Take-no-koshi (I-L) and P. asahinus male from Momoyake (M-P). Fully inflated endophallus in left lateral (A, E, I, M), apical (B, F, J, N), right lateral (C, G, K, O) and basal part in dorsal and apical part in ventral (D, H, L, P) views. Abbreviations: go - gonopore; lal - left apical lobe; lpb - left pigmented band; lpl - left pre-apical lobe; rpl - right pre-apical lobe." figureDoi="10.3897/dez.70.107322.figure4" httpUri="https://binary.pensoft.net/fig/893612" pageId="0" pageNumber="291">, 4E-H</figureCitation>
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<figureCitation id="7B2B69E9641BF95B6F308A9BE31E2EEB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Right paramere of Pterostichus namahage sp. nov. holotype male from Haraikawa (A), P. kamurosanus sp. nov. holotype male from Ariya (B), P. atsumidakensis sp. nov. holotype male from Take-no-koshi (C-E), P. asahinus male from Momoyake (F-H), P. chokaisanus male from Momoyake (I), P. chokaisanus male from Zao-chuo-kogen (J), P. kitakamisanus male from Odateminami (K, L) and P. kitakamisanus holotype male from Genbeidaira (M). Left lateral (A-C, F, I-K), right lateral (L, M), apical part in apical and basal part in ventral (D, G) and apical part in dorsal (E, H) views." figureDoi="10.3897/dez.70.107322.figure6" httpUri="https://binary.pensoft.net/fig/893614" pageId="0" pageNumber="291">, 6B</figureCitation>
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<figureCitation id="4E30316DB6E6916D3ADE346484CDF635" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genital membranous part of a Pterostichus kamurosanus sp. nov. paratype female from Ariya (A, B) and the P. atsumidakensis sp. nov. paratype female from Take-no-koshi (C, D). Dorsal (A, C) and ventral (B, D) views in everted condition of the vagina. Abbreviations: am - apophysis of the median oviduct; as - apophysis of the seminal canal; p - pigmentation on the innermost part of the vagina. Note that the two apophyses (am and as) are viewed through the membranous vagina." figureDoi="10.3897/dez.70.107322.figure7" httpUri="https://binary.pensoft.net/fig/893615" pageId="0" pageNumber="291">, 7A, B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="046C4AC1EBD42AC65E0374BC2FEE4C79" pageId="0" pageNumber="291" type="type material">
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<paragraph id="E1E3FE05108E1A439327441DE3E2C39E" pageId="0" pageNumber="291">Type materials.</paragraph>
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<paragraph id="589313AE11A3E2640BF39EAE7750E1C7" pageId="0" pageNumber="291">
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<emphasis id="6042AB3D5887A6C5AB89F67E12EA7699" bold="true" italics="true" pageId="0" pageNumber="291">Holotype</emphasis>
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, ♂, Ariya, alt. 403 m, Kaneyama-machi, Yamagata Prefecture, Japan (MGRS: 54SVJ49358 06377;
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<geoCoordinate id="06D9313A9376309CD3FF571D2A1CD68C" degrees="38.90482277" direction="north" orientation="latitude" precision="1" value="38.904823">38.90482277°N</geoCoordinate>
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,
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<geoCoordinate id="0D56368F06552A5839AD720A026BA50E" degrees="140.41594335" direction="east" orientation="longitude" precision="1" value="140.41594">140.41594335°E</geoCoordinate>
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), 18. vi-15. vii. 2021, subterranean baited trap, Y. Mitsuduka leg., in the collection of NSMT (Fig.
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<figureCitation id="6332707D9F8CF4AD8258BDE874FBD393" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus dorsal view of the Pterostichus macrogenys species group. A. Pterostichus namahage sp. nov. holotype male from Haraikawa; B. P. kamurosanus sp. nov. holotype male from Ariya; C. P. atsumidakensis sp. nov. holotype male from Take-no-koshi; D. P. asahinus male from Momoyake." figureDoi="10.3897/dez.70.107322.figure2" httpUri="https://binary.pensoft.net/fig/893610" pageId="0" pageNumber="291">2B</figureCitation>
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).
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<emphasis id="2D9338A1EC42F683C4FF58F443CC09A8" bold="true" italics="true" pageId="0" pageNumber="291">Paratypes</emphasis>
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, 4♀, same data as the holotype; 2♂2♀, Kamuro Forest Road, alt. 416 m, Ariya, Kaneyama-machi, Yamagata Prefecture, subterranean baited trap, Y. Mitsuduka leg. (1♂, 15-29. ix. 2015; 1♂2♀, 5. vii-21. viii. 2017).
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</paragraph>
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</subSubSection>
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<subSubSection id="918FA721ADC0A316A1E79B4B3A4CFB17" pageId="0" pageNumber="291" type="description">
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<paragraph id="DBDEB763199BB5027796FD43BFCAE84A" pageId="0" pageNumber="291">Description.</paragraph>
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<paragraph id="53FB6FBC7D8743CDA4CFCC5EBF29D029" pageId="0" pageNumber="291">
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<emphasis id="7864A87C111930E15D6EB403852F6965" bold="true" pageId="0" pageNumber="291">
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<emphasis id="8A3CAA2ACAADFCDEBD471F983D54D4BC" italics="true" pageId="0" pageNumber="291">Body length</emphasis>
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(
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<emphasis id="862C29EC146B5E219547F3EC7A56F216" italics="true" pageId="0" pageNumber="291">mm</emphasis>
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).
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</emphasis>
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♂ (n = 3), BLm 14.46-15.97, 15.19
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<normalizedToken id="D001CC8A9526F3A93DC9CCFAAE5163E9" originalValue="±">+/-</normalizedToken>
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0.64, BLl 13.15-14.43, 13.84
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<normalizedToken id="249B6FBF93763CF96047E26792E402CE" originalValue="±">+/-</normalizedToken>
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0.54, BLc 12.70-13.95, 13.37
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<normalizedToken id="7360A908E62154520E8A2617094145D5" originalValue="±">+/-</normalizedToken>
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0.52; ♀ (n = 6), BLm 14.19-15.08, 14.58
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<normalizedToken id="583E140DF36626D41C0A5783B958899F" originalValue="±">+/-</normalizedToken>
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0.46, BLl 13.10-13.63, 13.34
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<normalizedToken id="C2E58008BD43B01EF74574D396876DB6" originalValue="±">+/-</normalizedToken>
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0.27, BLc 12.60-13.21, 12.89
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<normalizedToken id="9267BB4416FC976E4BEBF58019C240EE" originalValue="±">+/-</normalizedToken>
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0.31.
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</paragraph>
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<paragraph id="2290FDC1CD26D3FBF11BB8A34A16D51E" pageId="0" pageNumber="291">
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<emphasis id="1F23B28176533845FA14D7AA4FA48430" bold="true" pageId="0" pageNumber="291">
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<emphasis id="CD84C7B89D8D1548EB6596F51F8BF4A8" italics="true" pageId="0" pageNumber="291">Head</emphasis>
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.
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</emphasis>
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Mandibular surface wrinkled with individual variation: short rudimentary wrinkles on both lateral sides, wrinkles on one lateral side, no wrinkles on either lateral side. Anterior-posterior length of tempora 1.7-1.8 times anterior-posterior length of eye. Surface of clypeus smooth in females, but irregular and slightly uneven in males. Antennal segment 2 with one or two setae, varying individually. Mentum tooth deeply bifid; width between paired apices apparently less than three times anterior-posterior length between level of mentum apices to level of innermost part of median notch.
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</paragraph>
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<paragraph id="7D58F7C42568922982F6F704C3C4C7E6" pageId="0" pageNumber="291">
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<emphasis id="61D39CD30E821C4EB95BAEF6DDE0A49D" bold="true" pageId="0" pageNumber="291">
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<emphasis id="99F4F5B0C719906192BF1532573F2832" italics="true" pageId="0" pageNumber="291">Pronotum</emphasis>
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.
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</emphasis>
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Lateral margin arcuate for apical 3/4, only slightly sinuate for basal 1/4; anterior margin emarginated, with contour arched more strongly than curvature of apical 3/4 of lateral margin; posterior margin emarginated at median area and arcuate at lateral areas, with curvature of median area distinctively greater than that of basal 1/4 of pronotal lateral margin and that of lateral areas same or only slightly greater than basal 1/4 of pronotal lateral margin; hind angles right-angled to slightly acute. Median line disappearing near pronotal anterior margin; in holotype male, posterior end disappears in front of transverse grooved area between laterobasal impressions, while in paratype female, posterior end reached pronotal posterior end. Transverse grooved area between laterobasal impressions concave to same degree as laterobasal impressions. Surface including laterobasal impressions smooth in most specimens; in some specimens, several punctations sparsely present near posterior ends of laterobasal impressions.
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</paragraph>
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<paragraph id="3FBEC76CD49689FA89C4E1D0F07C5452" pageId="0" pageNumber="291">
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<emphasis id="C6309B57DFC8AD0F6CA75E664A5D0CFB" bold="true" pageId="0" pageNumber="291">
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<emphasis id="78D8A2E6C9D2309D842EC4DE33E6B397" italics="true" pageId="0" pageNumber="291">Elytra</emphasis>
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.
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</emphasis>
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Scutellar-stria connected smoothly to stria 1 in most specimens; in some specimens, scutellar-stria disappears before connecting with stria 1, but its hypothetical extension smoothly connects to stria 1. One setigerous puncture on stria 1 at level of posterior end of scutellum. Posterior setigerous punctures on interval 3 at posterior 1/5-1/4.
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</paragraph>
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<paragraph id="A3361D6274F6AE4C8335456822642BFE" pageId="0" pageNumber="291">
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<emphasis id="C147F58659CD3EED0CCF69C5B992A060" bold="true" pageId="0" pageNumber="291">
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<emphasis id="1DFE2F76D688380B965F52663326297E" italics="true" pageId="0" pageNumber="291">Abdominal ventral sides</emphasis>
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.
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</emphasis>
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Sternum 7 of male very shallowly concave, with degree of concavity matching that of median area of sterna; shape of concavity transverse ellipse, but indistinct due to shallowness; posterior margin of ellipse aligns with sternum posterior margin; anterior-posterior length of concavity about 0.6 times anterior-posterior length of sternum 7; transverse, major axis of ellipse about 1.5 times that of longitudinal minor axis; area corresponding to minor axis very weakly raised, except near anterior and posterior ends.
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</paragraph>
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<paragraph id="6F74A9A46358730D0A1E982F467418B6" pageId="0" pageNumber="291">
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<emphasis id="5418A5867C17E281D5DF73D7E9E301B6" bold="true" pageId="0" pageNumber="291">
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<emphasis id="0F32874F1CBF5E9B01222A26E971C9EF" italics="true" pageId="0" pageNumber="291">Male genitalia</emphasis>
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.
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</emphasis>
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Endophallus with gonopore directed basal-ventrally; left pigmented band weakly sclerotised; right pre-apical lobe indistinct, only weakly swollen; left pre-apical lobe protrudes apically at apical part; left apical lobe short, not bifurcated (Fig.
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<figureCitation id="86713DF82361FE127CB3A294D67FF410" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Endophallus of Pterostichus namahage sp. nov. holotype male from Haraikawa (A-D), P. kamurosanus sp. nov. holotype male from Ariya (E-H), P. atsumidakensis sp. nov. holotype male from Take-no-koshi (I-L) and P. asahinus male from Momoyake (M-P). Fully inflated endophallus in left lateral (A, E, I, M), apical (B, F, J, N), right lateral (C, G, K, O) and basal part in dorsal and apical part in ventral (D, H, L, P) views. Abbreviations: go - gonopore; lal - left apical lobe; lpb - left pigmented band; lpl - left pre-apical lobe; rpl - right pre-apical lobe." figureDoi="10.3897/dez.70.107322.figure4" httpUri="https://binary.pensoft.net/fig/893612" pageId="0" pageNumber="291">4E-H</figureCitation>
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). Relative sizes of lobes: apical protrusion of left preapical lobe> left apical lobe apex >> right pre-apical lobe. Right paramere short, straight, with rounded apex (Fig.
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<figureCitation id="4D60CF4BC79B04BE1D6D7C17469C1624" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Right paramere of Pterostichus namahage sp. nov. holotype male from Haraikawa (A), P. kamurosanus sp. nov. holotype male from Ariya (B), P. atsumidakensis sp. nov. holotype male from Take-no-koshi (C-E), P. asahinus male from Momoyake (F-H), P. chokaisanus male from Momoyake (I), P. chokaisanus male from Zao-chuo-kogen (J), P. kitakamisanus male from Odateminami (K, L) and P. kitakamisanus holotype male from Genbeidaira (M). Left lateral (A-C, F, I-K), right lateral (L, M), apical part in apical and basal part in ventral (D, G) and apical part in dorsal (E, H) views." figureDoi="10.3897/dez.70.107322.figure6" httpUri="https://binary.pensoft.net/fig/893614" pageId="0" pageNumber="291">6B</figureCitation>
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).
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</paragraph>
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<paragraph id="7F0495C39CC26DC69ADCEA06B010E91E" pageId="0" pageNumber="291">
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<emphasis id="40EFD876413C5B3779D621787450A8ED" bold="true" italics="true" pageId="0" pageNumber="291">Female genitalia</emphasis>
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. Innermost part of vagina with pigmentation (Fig.
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<figureCitation id="566B1092BCA2777E3EDE4F168713F464" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genital membranous part of a Pterostichus kamurosanus sp. nov. paratype female from Ariya (A, B) and the P. atsumidakensis sp. nov. paratype female from Take-no-koshi (C, D). Dorsal (A, C) and ventral (B, D) views in everted condition of the vagina. Abbreviations: am - apophysis of the median oviduct; as - apophysis of the seminal canal; p - pigmentation on the innermost part of the vagina. Note that the two apophyses (am and as) are viewed through the membranous vagina." figureDoi="10.3897/dez.70.107322.figure7" httpUri="https://binary.pensoft.net/fig/893615" pageId="0" pageNumber="291">7A, B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="87097596D9D5F7298C0603BAECDE92AF" pageId="0" pageNumber="291" type="etymology">
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<paragraph id="62F20173331070D96BCDABFB29562F68" pageId="0" pageNumber="291">Etymology.</paragraph>
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<paragraph id="587C436694E191492A752175EDAEE6AC" pageId="0" pageNumber="291">
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The specific name is an adjective, derived from the Japanese noun Kamurosan, which refers to Mt. Kamuro, where the type specimens were collected and the Latin adjectival suffix -
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<emphasis id="A39F3B8C582D407D5F817A07DC06AE24" italics="true" pageId="0" pageNumber="291">anus</emphasis>
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(m), which means, when attached to a noun, "pertaining to".
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |