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<mods:title id="5A6AE68A998FCE9FB15D153005A6CF87">Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym</mods:title>
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<mods:title id="9D751060A2697F4D52FB728E0E1FFA1E">Journal of Natural History</mods:title>
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<heading id="854F81B7FFFA326B9B93497DFCD61712" bold="true" box="[428,887,1100,1126]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="1547" reason="3">
<taxonomicName id="19B84D58FFFA326B9B93497DFCD61712" ID-CoL="6JHMP" authority="James, 1969" authorityName="James" authorityYear="1969" box="[428,887,1100,1126]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="1547" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="ECCCEAC9FFFA326B9B93497DFCD61712" bold="true" box="[428,887,1100,1126]" pageId="35" pageNumber="1547">
<emphasis id="ECCCEAC9FFFA326B9B93497DFD741712" bold="true" box="[428,725,1100,1126]" italics="true" pageId="35" pageNumber="1547">Formosargus variegatus</emphasis>
<bibRefCitation id="BA294B2AFFFA326B98E2497DFCD61712" author="James MT" box="[733,887,1100,1126]" pageId="35" pageNumber="1547" pagination="339 - 350" refId="ref19004" refString="James MT. 1969. The soldier flies (Diptera, Stratiomyidae) collected in the Philippine Islands by the Noona Dan Expedition. Entomol Medd. 37 (4): 339 - 350." type="journal article" year="1969">James, 1969</bibRefCitation>
</emphasis>
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(
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)
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<treatmentCitation id="5F1910CAFFFA326B9A9F499DFDC717B1" author="James MT" box="[160,614,1196,1221]" page="342" pageId="35" pageNumber="1547" year="1969">
<taxonomicName id="19B84D58FFFA326B9A9F499DFDC717B1" ID-CoL="6JHMP" authority="James, 1969: 342" authorityName="James" authorityPageNumber="342" authorityYear="1969" box="[160,614,1196,1221]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="1547" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="ECCCEAC9FFFA326B9A9F499DFE0517B1" box="[160,420,1196,1221]" italics="true" pageId="35" pageNumber="1547">Formosargus variegatus</emphasis>
<bibRefCitation id="BA294B2AFFFA326B9B95499DFDC717B1" author="James MT" box="[426,614,1196,1221]" pageId="35" pageNumber="1547" pagination="339 - 350" refId="ref19004" refString="James MT. 1969. The soldier flies (Diptera, Stratiomyidae) collected in the Philippine Islands by the Noona Dan Expedition. Entomol Medd. 37 (4): 339 - 350." type="journal article" year="1969">James, 1969: 342</bibRefCitation>
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(orig. descr.).
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Type locality:
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,
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,
<collectingMunicipality id="3E63ACA1FFFA326B9AFF49FEFE10179C" box="[192,433,1231,1256]" pageId="35" pageNumber="1547">Mantalingajan Range</collectingMunicipality>
,
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.
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.
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<treatmentCitation id="5F1910CAFFFA326B9A9F49C3FD3F167F" author="Yang D &amp; Zhang TT &amp; Li Z" box="[160,670,1266,1291]" page="417" pageId="35" pageNumber="1547" year="2014">
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<emphasis id="ECCCEAC9FFFA326B9A9F49C3FE02167F" box="[160,419,1266,1291]" italics="true" pageId="35" pageNumber="1547">Formosargus variegates</emphasis>
<bibRefCitation id="BA294B2AFFFA326B9B9549C3FD3F167F" author="Yang D &amp; Zhang TT &amp; Li Z" box="[426,670,1266,1291]" pageId="35" pageNumber="1547" refId="ref20113" refString="Yang D, Zhang TT, Li Z. 2014. Stratiomyoidea of China. Beijing: China Agricultural University Press; p. 870." type="journal volume" year="2014">Yang et al., 2014: 417</bibRefCitation>
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. Incorrect subsequent spelling.
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</paragraph>
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Refs:
<bibRefCitation id="BA294B2AFFFA326B9AE44825FE2D1659" author="James MT" box="[219,396,1300,1325]" pageId="35" pageNumber="1547" pagination="14 - 42" refId="ref19040" refString="James MT. 1975. Family Stratiomyidae. In: Delfinado MD, Hardy DE, editors. A catalog of the diptera of the Oriental Region. Vol. II. suborder brachycera through division Aschiza, suborder cyclorrhapha. Vol. i - x. Honolulu: The University Press of Hawaii; p. 14 - 42." type="book chapter" year="1975">James, 1975: 18</bibRefCitation>
(cat.);
<bibRefCitation id="BA294B2AFFFA326B9BE84825FD151659" author="Woodley NE" box="[471,692,1300,1325]" pageId="35" pageNumber="1547" pagination="1 - 475" refId="ref19982" refString="Woodley NE. 2001. A world catalog of Stratiomyidae (Insecta: Diptera). Myia. 11: 1 - 475." type="journal article" year="2001">Woodley, 2001: 190</bibRefCitation>
(cat.).
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<emphasis id="ECCCEAC9FFFA326B9A9F4855FEB5160A" bold="true" box="[160,276,1380,1406]" italics="true" pageId="35" pageNumber="1547">Diagnosis</emphasis>
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Both scutal medial band and a pair of anterior lateral dark brown maculae discontinued at transverse suture; posterior part of scutum with a U-shaped dark brown macula; scutellum completely yellowish (
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); anepisternum widely dark brown, laterotergite and mediotergite dark brown. All hind tarsomeres yellowish.
</paragraph>
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<emphasis id="ECCCEAC9FFFA326B9A9F4B0FFEDE152C" bold="true" box="[160,383,1598,1624]" italics="true" pageId="35" pageNumber="1547">Material examined</emphasis>
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<emphasis id="ECCCEAC9FFFA326B9A9F4B50FEFA150F" bold="true" box="[160,347,1633,1659]" italics="true" pageId="35" pageNumber="1547">Type material.</emphasis>
<materialsCitation id="6ED03C86FFFA326B9B544B53FB2215B4" ID-GBIF-Occurrence="3965743327" collectingDate="1961-09-24" country="Philippines" elevation="600" location="Pinigisan" municipality="Mantalingajan" pageId="35" pageNumber="1547" specimenCount="1" specimenCount-male="1" stateProvince="Palawan" typeStatus="holotype">
<emphasis id="ECCCEAC9FFFA326B9B544B53FE51150F" bold="true" box="[363,496,1634,1659]" pageId="35" pageNumber="1547">
<typeStatus id="01038879FFFA326B9B544B53FE51150F" box="[363,496,1634,1659]" pageId="35" pageNumber="1547" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
(
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(ac)),
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labelled:
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,
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/
<collectingMunicipality id="3E63ACA1FFFA326B9A9F4BB4FEE115EA" box="[160,320,1669,1694]" pageId="35" pageNumber="1547">Mantalingajan</collectingMunicipality>
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<location id="DB676000FFFA326B9B774BB4FE0F15EA" LSID="urn:lsid:plazi:treatment:561187CDFFFA326E9B93497DFD851503:DB676000FFFA326B9B774BB4FE0F15EA" box="[328,430,1669,1694]" country="Philippines" municipality="Mantalingajan" name="Pinigisan" pageId="35" pageNumber="1547" stateProvince="Palawan">Pinigisan</location>
<quantity id="19409B3EFFFA326B9B884BB4FD8B15EA" box="[439,554,1669,1694]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="35" pageNumber="1547" unit="m" value="600.0">
<elevation id="5595D1E8FFFA326B9B884BB4FD8B15EA" box="[439,554,1669,1694]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="35" pageNumber="1547" unit="m" value="600.0">600 meter</elevation>
</quantity>
/
<date id="AA06101BFFFA326B980C4BB4FCB015EA" box="[563,785,1669,1694]" pageId="35" pageNumber="1547" value="1961-09-24">
<collectingDate id="BA42E9F3FFFA326B980C4BB4FCB015EA" box="[563,785,1669,1694]" pageId="35" pageNumber="1547" value="1961-09-24">24 September 1961</collectingDate>
</date>
/Noona Dan Exp. 6162 [printed on white paper]; Caught in/malaise-traps/inside forest [printed on white paper]; zmuc/
</materialsCitation>
00026038 [printed on white paper];
<taxonomicName id="19B84D58FFFF326E987A4907FD73173B" baseAuthorityName="James" baseAuthorityYear="1939" box="[581,722,1078,1103]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="1550" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFFF326E987A4907FD73173B" box="[581,722,1078,1103]" italics="true" pageId="38" pageNumber="1550">Formosargus</emphasis>
</taxonomicName>
/
<taxonomicName id="19B84D58FFFF326E98E44907FCED173B" authorityName="James" authorityYear="1969" box="[731,844,1078,1103]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="1550" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="ECCCEAC9FFFF326E98E44907FCED173B" box="[731,844,1078,1103]" italics="true" pageId="38" pageNumber="1550">variegatus</emphasis>
</taxonomicName>
/
<specimenCount id="C8BEFD52FFFF326E996A4906FCC8173A" box="[853,873,1079,1102]" pageId="38" pageNumber="1550" type="male"></specimenCount>
<emphasis id="ECCCEAC9FFFF326E994E4907FC15173B" box="[881,948,1078,1103]" italics="true" pageId="38" pageNumber="1550">James</emphasis>
/
<emphasis id="ECCCEAC9FFFF326E99824907FB93173B" box="[957,1074,1078,1103]" italics="true" pageId="38" pageNumber="1550">
<typeStatus id="01038879FFFF326E99824907FB93173B" box="[957,1074,1078,1103]" pageId="38" pageNumber="1550" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
[handwritten on red paper] (
<collectionCode id="B8A9AE1EFFFF326E9B9A4968FE511706" box="[421,496,1113,1138]" country="Venezuela" name="Universidad del Zulia" pageId="38" pageNumber="1550">UZMC</collectionCode>
). The
<typeStatus id="01038879FFFF326E980E4968FD341706" box="[561,661,1113,1138]" pageId="38" pageNumber="1550" type="holotype">holotype</typeStatus>
has the last four mid tarsomeres missing.
</paragraph>
</subSubSection>
<caption id="8AC76653FFFD326C9A9F4B65FE5B15FC" ID-DOI="http://doi.org/10.5281/zenodo.7318236" ID-Zenodo-Dep="7318236" httpUri="https://zenodo.org/record/7318236/files/figure.png" pageId="36" pageNumber="1548" startId="36.[160,225,1620,1643]" targetBox="[167,1148,145,1585]" targetPageId="36">
<paragraph id="DE0736DBFFFD326C9A9F4B65FE5B15FC" blockId="36.[160,1157,1620,1672]" pageId="36" pageNumber="1548">
<emphasis id="ECCCEAC9FFFD326C9A9F4B65FEAB151F" bold="true" box="[160,266,1620,1643]" pageId="36" pageNumber="1548">Figure 24.</emphasis>
<taxonomicName id="19B84D58FFFD326C9B2B4B65FE74151F" authorityName="Adisoemarto" authorityYear="1974" box="[276,469,1620,1643]" class="Insecta" family="Stratiomyidae" genus="Amsaria" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="1548" phylum="Arthropoda" rank="species" species="sagittocera">
<emphasis id="ECCCEAC9FFFD326C9B2B4B65FE74151F" box="[276,469,1620,1643]" italics="true" pageId="36" pageNumber="1548">Amsaria sagittocera</emphasis>
</taxonomicName>
, female, holotype [MZB]. (ab) Habitus, dorsal view. (cd) Habitus, lateral view. (e) Head, frontal view.
</paragraph>
</caption>
<caption id="8AC76653FFFC326D9A9F4B7CFDB515EA" ID-DOI="http://doi.org/10.5281/zenodo.7318238" ID-Zenodo-Dep="7318238" httpUri="https://zenodo.org/record/7318238/files/figure.png" pageId="37" pageNumber="1549" startId="37.[160,225,1613,1636]" targetBox="[174,1142,150,1573]" targetPageId="37">
<paragraph id="DE0736DBFFFC326D9A9F4B7CFDB515EA" blockId="37.[160,1156,1613,1694]" pageId="37" pageNumber="1549">
<emphasis id="ECCCEAC9FFFC326D9A9F4B7CFEA71510" bold="true" box="[160,262,1613,1636]" pageId="37" pageNumber="1549">Figure 25.</emphasis>
<taxonomicName id="19B84D58FFFC326D9B344B7CFE4F1510" authorityName="James" authorityYear="1969" box="[267,494,1613,1636]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="1549" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="ECCCEAC9FFFC326D9B344B7CFE4F1510" box="[267,494,1613,1636]" italics="true" pageId="37" pageNumber="1549">Formosargus variegatus</emphasis>
</taxonomicName>
, male, holotype [UZMC]. (a) Habitus, dorsal view. (b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen).
</paragraph>
</caption>
<caption id="8AC76653FFFF326E9A9F4E57FE3B10A1" ID-DOI="http://doi.org/10.5281/zenodo.7318240" ID-Zenodo-Dep="7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="38" pageNumber="1550" startId="38.[160,226,870,893]" targetBox="[168,1148,146,834]" targetPageId="38">
<paragraph id="DE0736DBFFFF326E9A9F4E57FE3B10A1" blockId="38.[160,1155,870,981]" pageId="38" pageNumber="1550">
<emphasis id="ECCCEAC9FFFF326E9A9F4E57FEAD1009" bold="true" box="[160,268,870,893]" pageId="38" pageNumber="1550">Figure 26.</emphasis>
Geographic distribution of
<taxonomicName id="19B84D58FFFF326E980E4E57FD0C1009" baseAuthorityName="James" baseAuthorityYear="1939" box="[561,685,870,893]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="1550" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCEAC9FFFF326E980E4E57FD0C1009" box="[561,685,870,893]" italics="true" pageId="38" pageNumber="1550">Formosargus</emphasis>
</taxonomicName>
. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the lines of faunal division are shown: 1. Wallaces Line; 2. Webers Line; 3. Lydekkers Line. Wallacea is the area that lies between Wallaces Line and Lydekkers Line.
</paragraph>
</caption>
<subSubSection id="96A26550FFFF326E9A9F499CFD7F179E" pageId="38" pageNumber="1550" type="distribution">
<paragraph id="DE0736DBFFFF326E9A9F499CFE1817B3" blockId="38.[160,734,1197,1258]" box="[160,441,1197,1223]" pageId="38" pageNumber="1550">
<emphasis id="ECCCEAC9FFFF326E9A9F499CFE1817B3" bold="true" box="[160,441,1197,1223]" italics="true" pageId="38" pageNumber="1550">Geographic distribution</emphasis>
</paragraph>
<paragraph id="DE0736DBFFFF326E9A9F49E0FD7F179E" blockId="38.[160,734,1197,1258]" box="[160,734,1233,1258]" pageId="38" pageNumber="1550">
<emphasis id="ECCCEAC9FFFF326E9A9F49E0FEFC179E" bold="true" box="[160,349,1233,1258]" pageId="38" pageNumber="1550">Oriental Region</emphasis>
.
<collectingCountry id="A6AF764BFFFF326E9B5549E0FE44179E" box="[362,485,1233,1258]" name="Philippines" pageId="38" pageNumber="1550">Philippines</collectingCountry>
(
<collectingRegion id="1C7CF839FFFF326E9BCD49E0FDF2179E" box="[498,595,1233,1258]" country="Philippines" name="Palawan" pageId="38" pageNumber="1550">Palawan</collectingRegion>
) (
<figureCitation id="46832A5EFFFF326E985749E0FD70179E" box="[616,721,1233,1258]" captionStart="Figure 26" captionStartId="38.[160,226,870,893]" captionTargetBox="[168,1148,146,834]" captionTargetId="figure-167@38.[163,1153,142,839]" captionTargetPageId="38" captionText="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the lines of faunal division are shown: 1. Wallaces Line; 2. Webers Line; 3. Lydekkers Line. Wallacea is the area that lies between Wallaces Line and Lydekkers Line." figureDoi="http://doi.org/10.5281/zenodo.7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="38" pageNumber="1550">Figure 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="96A26550FFFF326E9A9F4814FD851503" pageId="38" pageNumber="1550" type="discussion">
<paragraph id="DE0736DBFFFF326E9A9F4814FEBF164B" blockId="38.[160,1157,1317,1655]" box="[160,286,1317,1343]" pageId="38" pageNumber="1550">
<emphasis id="ECCCEAC9FFFF326E9A9F4814FEBF164B" bold="true" box="[160,286,1317,1343]" italics="true" pageId="38" pageNumber="1550">Comments</emphasis>
</paragraph>
<paragraph id="DE0736DBFFFF326E9A9F4878FD851503" blockId="38.[160,1157,1317,1655]" pageId="38" pageNumber="1550">
<taxonomicName id="19B84D58FFFF326E9A9F4878FE1B1616" authorityName="James" authorityYear="1969" box="[160,442,1353,1378]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="1550" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="ECCCEAC9FFFF326E9A9F4878FE1B1616" box="[160,442,1353,1378]" italics="true" pageId="38" pageNumber="1550">Formosargus variegatus</emphasis>
</taxonomicName>
has a scutum colour pattern similar to that seen in
<taxonomicName id="19B84D58FFFF326E9E4E4878FEAD16F1" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="1550" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
<emphasis id="ECCCEAC9FFFF326E9E4E4878FEAD16F1" italics="true" pageId="38" pageNumber="1550">F. trivittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFFF326E9B2B485DFED116F1" box="[276,368,1388,1413]" pageId="38" pageNumber="1550" rank="species">sp. nov.</taxonomicNameLabel>
, since both species have at least three dark bands over the anterior part of the scutum, with yellow between the bands and the end of each of posterior lateral band connected to the medial band posteriorly. In
<taxonomicName id="19B84D58FFFF326E99724880FC7D16BE" authorityName="James" authorityYear="1969" box="[845,988,1457,1482]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="1550" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="ECCCEAC9FFFF326E99724880FC7D16BE" box="[845,988,1457,1482]" italics="true" pageId="38" pageNumber="1550">F. variegatus</emphasis>
</taxonomicName>
, however, the medial band is completely interrupted at the transverse suture and the posterior lateral bands are strongly fused medially in the form of a semi-ellipse (
<figureCitation id="46832A5EFFFF326E9E3A48C7FBD7157B" box="[1029,1142,1526,1551]" captionStart="Figure 25" captionStartId="37.[160,225,1613,1636]" captionTargetBox="[174,1142,150,1573]" captionTargetId="figure-3@37.[163,1153,142,1581]" captionTargetPageId="37" captionText="Figure 25. Formosargus variegatus, male, holotype [UZMC]. (a) Habitus, dorsal view.(b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen)." figureDoi="http://doi.org/10.5281/zenodo.7318238" httpUri="https://zenodo.org/record/7318238/files/figure.png" pageId="38" pageNumber="1550">Figure 25</figureCitation>
), while in
<taxonomicName id="19B84D58FFFF326E9B2E4B28FE3E1546" authority="Fachin &amp; Hauser, 2022" authorityName="Fachin &amp; Hauser" authorityYear="2022" box="[273,415,1561,1586]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="1550" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
<emphasis id="ECCCEAC9FFFF326E9B2E4B28FE3E1546" box="[273,415,1561,1586]" italics="true" pageId="38" pageNumber="1550">F. trivittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7FF57B2FFFF326E9B924B28FDB11546" box="[429,528,1561,1586]" pageId="38" pageNumber="1550" rank="species">sp. nov.</taxonomicNameLabel>
, the medial band is narrowed at the level of the transverse suture and the posterior lateral bands are weakly connected to it, forming a triangular macula (
<figureCitation id="46832A5EFFFF326E9BA94B6FFE581503" box="[406,505,1630,1655]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (ab) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="38" pageNumber="1550">Figure 7</figureCitation>
(h)).
</paragraph>
</subSubSection>
</treatment>
</document>