457 lines
53 KiB
XML
457 lines
53 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1198.120353" ID-Pensoft-Pub="1313-2970-1198-279" ID-Pensoft-UUID="080972D94F1654EEB056AF1FE576D04B" ID-ZooBank="A5AC5067B58D44909B9818CC1CD0CDA9" ModsDocID="1313-2970-1198-279" checkinTime="1714135927740" checkinUser="pensoft" docAuthor="Harvey, Mark S. & Steinmann, David B." docDate="2024" docId="4E2E83EE476D5E83BC4BDD5AC3E7DEDA" docLanguage="en" docName="ZooKeys 1198: 279-294" docOrigin="ZooKeys 1198" docPubDate="2024-04-25" docSource="http://dx.doi.org/10.3897/zookeys.1198.120353" docTitle="Larca boulderica Harvey & Steinmann 2024, sp. nov." docType="treatment" docUuid="771E4C1C-56BD-4977-99BD-6DE45C97160A" docUuidSource="ZooBank" docVersion="1" id="080972D94F1654EEB056AF1FE576D04B" lastPageNumber="279" masterDocId="080972D94F1654EEB056AF1FE576D04B" masterDocTitle="A new troglomorphic species of Larca (Pseudoscorpiones, Larcidae) from Colorado" masterLastPageNumber="294" masterPageNumber="279" pageNumber="279" updateTime="1714135927740" updateUser="pensoft">
|
||
<mods:mods id="22E2CC5AF2A79C11CD355B8DA9950F99" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="96BFAC5D565F9F9A5CD56AA9553367FC">
|
||
<mods:title id="E1C27F75D33A055A7724B6C12113BD5A">A new troglomorphic species of Larca (Pseudoscorpiones, Larcidae) from Colorado</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5B4302DD20BA805AF2D26BCF63C6421C" type="personal">
|
||
<mods:role id="BBA1F7BBB1B4AD0468717A5FC1F0F028">
|
||
<mods:roleTerm id="2C2D529B863792D2DA00D6F680BE3BEA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="51E50A5221D217E244F339F0E30D8FB0">Harvey, Mark S.</mods:namePart>
|
||
<mods:nameIdentifier id="E3EA429DE6FADE199BB46593EB2079C5" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
|
||
<mods:affiliation id="D8C21BDCBC6F70F581F1198882F09AEE">Collections & Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia & School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia</mods:affiliation>
|
||
<mods:nameIdentifier id="B7E666BBCDEBC2598D1DE40A7D67AE38" type="email">mark.harvey@museum.wa.gov.au</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="2717D7AB6D71B3C8B905F2631B1D7644" type="personal">
|
||
<mods:role id="5D821B7E9708D0EF365DDB98BD64F76B">
|
||
<mods:roleTerm id="C0B65B895CBF6F84B7A3215A3AB62982">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="91E855138D3B1FB3E6C0CC93D7DC3703">Steinmann, David B.</mods:namePart>
|
||
<mods:affiliation id="3C0C25113B38B7BDBC401C2F415872E6">Department of Zoology, Denver Museum of Nature & Science, 2001 Colorado Boulevard, Denver, Colorado 80205, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="85C64E4E68B24C7E0715B5FB00EA697D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A42F4DA2FCAD17EB92252AA8D340D40B" type="host">
|
||
<mods:titleInfo id="EDF144C556BEA5DDFB35E763F223C1F8">
|
||
<mods:title id="B84BE1E247E29E9B7CDCE07FDFB4FDCF">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B9E3AF00F4342E02A39D5A2A7F868F74">
|
||
<mods:date id="298061D6259255512449352852B1402E">2024</mods:date>
|
||
<mods:detail id="B12499B6E3516CD8D0420F0A7C263085" type="pubDate">
|
||
<mods:number id="380FB68CB775BA24F2615669CADAE30A">2024-04-25</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="78991929E626466135722CFE679D9A7B" type="volume">
|
||
<mods:number id="51032A5D004C100A938F92C7ECAC8CF3">1198</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C0AAFB8BA6864A31B3EADE739D4738FF" unit="page">
|
||
<mods:start id="8D2930BF05F29AD68AADF8C82BF71CD9">279</mods:start>
|
||
<mods:end id="9D60297C94743132050B29D6FE362B1D">294</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="327A0AB0D66E1BEC09B18807249F9BB4">
|
||
<mods:url id="4953619F841D7E8C1C2D42764EF522CE">http://dx.doi.org/10.3897/zookeys.1198.120353</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="98E294AD390B24AE7FF4111CDB922B25">journal article</mods:classification>
|
||
<mods:identifier id="D4893C3FF44D78CACE200EB0C20A6BEA" type="DOI">http://dx.doi.org/10.3897/zookeys.1198.120353</mods:identifier>
|
||
<mods:identifier id="136ECB45147413918F1F215881191604" type="Pensoft-Pub">1313-2970-1198-279</mods:identifier>
|
||
<mods:identifier id="8037A4052671F02E8FCAE07B683ADF96" type="ZooBank">A5AC5067B58D44909B9818CC1CD0CDA9</mods:identifier>
|
||
<mods:identifier id="BEDDD33ED2D2D0F457C1975A8D5867E6" type="Pensoft-UUID">080972D94F1654EEB056AF1FE576D04B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="4E2E83EE476D5E83BC4BDD5AC3E7DEDA" LSID="urn:lsid:zoobank.org:act:771E4C1C-56BD-4977-99BD-6DE45C97160A" httpUri="http://treatment.plazi.org/id/4E2E83EE476D5E83BC4BDD5AC3E7DEDA" lastPageNumber="279" pageId="0" pageNumber="279">
|
||
<subSubSection id="C98AE09349788567B6284C9CE3F421E5" pageId="0" pageNumber="279" type="nomenclature">
|
||
<paragraph id="01F2049FBCAC8F4826733937C69908F0" pageId="0" pageNumber="279">
|
||
<taxonomicName id="039E0EFC39DC9ED6D73C74A730DE8F08" LSID="https://zoobank.org/771E4C1C-56BD-4977-99BD-6DE45C97160A" authority="Harvey & Steinmann, 2024" authorityName="Harvey & Steinmann" authorityYear="2024" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca boulderica" order="Pseudoscorpiones" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="boulderica" status="sp. nov.">Larca boulderica</taxonomicName>
|
||
<taxonomicNameLabel id="3D4A029C0D852F2D1AF2483871516D6E" pageId="0" pageNumber="279">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7E2C4BBE0B8672A11B24B9D2F3EB7E67" pageId="0" pageNumber="279" type="description">
|
||
<paragraph id="712F7B2E3A4C28C5F38755962034D602" pageId="0" pageNumber="279">
|
||
<figureCitation id="76003430BE3CECCFC75D1CDBAB87DC71" captionStart="Figure 6" captionStartId="F4" captionText="Figure 6. Larca boulderica sp. nov., in situ in Mallory Cave." figureDoi="10.3897/zookeys.1198.120353.figure6" httpUri="https://binary.pensoft.net/fig/1033627" pageId="0" pageNumber="279">Figs 6</figureCitation>
|
||
<figureCitation id="08F1DA35597AFFDBA4759FCFD511EC18" captionStart="Figures 7–9" captionStartId="F5" captionText="Figures 7 - 9. Larca boulderica sp. nov., paratype female (DMNS): 7 body, dorsal 8 body, ventral 9 cephalothorax, dorsal. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1198.120353.figures7-9" httpUri="https://binary.pensoft.net/fig/1033628" pageId="0" pageNumber="279">, 7-9</figureCitation>
|
||
<figureCitation id="543FA6DC5508CAAC399E7C48EE8C3703" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">, 10-18</figureCitation>
|
||
<figureCitation id="04448497281D8A21375AD4694D59BBBB" captionStart="Figures 19–21" captionStartId="F7" captionText="Figures 19 - 21. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 19 genital sternites, ventral, male 20 genital sternites, ventral, female 21 genitalia, ventral, female. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1198.120353.figures19-21" httpUri="https://binary.pensoft.net/fig/1033630" pageId="0" pageNumber="279">, 19-21</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4F63EA8B0F82E22866F503BE231DD290" pageId="0" pageNumber="279" type="materials_examined">
|
||
<paragraph id="4AE79B96D08D4D5400510E7E8D15DF82" pageId="0" pageNumber="279">Material examined.</paragraph>
|
||
<paragraph id="0A563213B87855EF330E788B2AFE712E" pageId="0" pageNumber="279">
|
||
<materialsCitation id="5B2E40A512C7236CA0A0CC7A57ADBA93" collectingDate="2008-11-29" collectorName="D. Steinmann" country="U. S. A." county="Boulder County" elevation="2134" latitude="39.974167" location="Mallory Cave" longLatPrecision="11" longitude="-105.2895" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
|
||
<emphasis id="DCDDD3C2CCCBE7BD874BF39E83E09D19" bold="true" pageId="0" pageNumber="279">Types</emphasis>
|
||
:
|
||
<collectingCountry id="1CE9B4AA788B1EBABAF77A57BAA877A7" name="United States of America">U.S.A.</collectingCountry>
|
||
:
|
||
<emphasis id="07B5B38D3ED0FDD3399D0263305861EA" bold="true" pageId="0" pageNumber="279">
|
||
<collectingRegion id="F2F84DC17DA4D916B2D63A58E54759E1" country="United States of America" name="Colorado">Colorado</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty id="54A7965F34039FFACDC5BFD3109BB958">Boulder County</collectingCounty>
|
||
:
|
||
<emphasis id="8FB558E8B94C2C4BDCF7E4965B40986B" bold="true" italics="true" pageId="0" pageNumber="279">
|
||
<typeStatus id="BC5C32B03D8C8F6C00F354D1873863E6" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
male,
|
||
<location id="400F36450013089866E8F577A59DF123" LSID="urn:lsid:plazi:treatment:4E2E83EE476D5E83BC4BDD5AC3E7DEDA:400F36450013089866E8F577A59DF123" country="U. S. A." county="Boulder County" latitude="39.974167" longLatPrecision="11" longitude="-105.2895" name="Mallory Cave" stateProvince="Colorado">Mallory Cave</location>
|
||
,
|
||
<geoCoordinate id="702BE0ABC193655CD1E538D6477875DB" degrees="39" direction="north" minutes="58.45" orientation="latitude" precision="9" value="39.974167">39°58.45'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CA3D0D4334B27EB3B89097BB2094B31E" degrees="105" direction="west" minutes="17.37" orientation="longitude" precision="9" value="-105.2895">105°17.37'W</geoCoordinate>
|
||
,
|
||
<quantity id="61345BA35FA3E45A2EDD07A55403F6EE" metricMagnitude="3" metricUnit="m" metricValue="2.1336" unit="ft" value="7000.0">
|
||
<elevation id="9BBE1DD35D7A9620E31473C6235AAD12" metricMagnitude="3" metricUnit="m" metricValue="2.1336" unit="ft" value="7000.0">7000 ft</elevation>
|
||
</quantity>
|
||
(
|
||
<quantity id="6C0439B44DF57156C403E4569439BF9B" metricMagnitude="3" metricUnit="m" metricValue="2.1399999999999997" unit="m" value="2140.0">
|
||
<elevation id="65EBFC546CCA047836EFE2D269DA6F47" metricMagnitude="3" metricUnit="m" metricValue="2.1399999999999997" unit="m" value="2140.0">2140 m</elevation>
|
||
</quantity>
|
||
) a.s.l.,
|
||
<collectingDate id="DACAA7171622E8145CDEB6850FE0EFF1" value="2008-11-29">29 November 2008</collectingDate>
|
||
, under rock, dark zone of cave,
|
||
<collectorName id="C1EC312E0800ED2FB8240C051456A922">D. Steinmann</collectorName>
|
||
(DMNS)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="CA3E9C83F9999B267D1418819604039E" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="D. Steinmann" country="United States of America" county="Boulder County" elevation="2134" latitude="39.974167" location="Paratypes" longLatPrecision="11" longitude="-105.2895" specimenCount="4" specimenCount-female="4" stateProvince="Paratypes" typeStatus="Paratypes">
|
||
<emphasis id="1AFDACA66E176450AC38FB4F37908841" bold="true" italics="true" pageId="0" pageNumber="279">
|
||
<collectingRegion id="10533B03CBC049B1D56AF2BD38A2C69F" country="United States of America" name="Paratypes">
|
||
<typeStatus id="542089B924F52367BADB1E2B31F71C56" type="paratype">Paratypes</typeStatus>
|
||
</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="DDCBC4C48813C74A1372B351A350FE3E" count="4" type="female">4 females</specimenCount>
|
||
, collected with holotype (DMNS)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="031A4CD754B02D14692F7B3D94FEF8DA" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="D. Steinmann" country="United States of America" county="Boulder County" elevation="2134" latitude="39.974167" location="Mallory Cave" longLatPrecision="11" longitude="-105.2895" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="paratype">
|
||
<specimenCount id="7A2D16151104F7BE5EAF8B171692371C" count="1" type="male">1 male</specimenCount>
|
||
, collected with holotype (WAM T162363)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="337CD2C07BEA11C65B219DB5002BF83A" collectingDate="2009-11-12" collectorName="D. Steinmann" country="United States of America" county="Boulder County" elevation="2134" latitude="39.974167" location="Mallory Cave" longLatPrecision="11" longitude="-105.2895" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="paratype">
|
||
<specimenCount id="D7E204E9CC9F37C29B1DC6B6497B90D6" count="1" type="female">1 female</specimenCount>
|
||
, same data as holotype except
|
||
<collectingDate id="9993F0337AFD2223CE9C420951CCE13F" value="2009-11-12">12 November 2009</collectingDate>
|
||
(DMNS)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8187CFF61C6B3C9FC4DC45EDDEC8E359" collectingDate="2009-11-12" collectorName="D. Steinmann" country="United States of America" county="Boulder County" elevation="2134" latitude="39.974167" location="Mallory Cave" longLatPrecision="11" longitude="-105.2895" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="paratype">
|
||
<specimenCount id="15CBBBBCE4CCB8D15C31D0498042F297" count="1" type="female">1 female</specimenCount>
|
||
, same data as holotype except
|
||
<collectingDate id="9816E98EBFF37FC1AE556B169FF42589" value="2009-11-12">12 November 2009</collectingDate>
|
||
(WAM T162059)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C96030C8D66DF5E7D9E318FA1010300" pageId="0" pageNumber="279" type="diagnosis">
|
||
<paragraph id="346EA8F2F05A33D748E5CD9D7858ADF7" pageId="0" pageNumber="279">Diagnosis.</paragraph>
|
||
<paragraph id="B7837752DDA87150C5380551B37B3292" pageId="0" pageNumber="279">
|
||
<taxonomicName id="9D5C2740F4D20E5B09D5266FBBB48857" authorityName="Harvey & Steinmann" authorityYear="2024" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca boulderica" order="Pseudoscorpionida" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="boulderica">
|
||
<emphasis id="B3D9ED535A0CFB45BF8F284B98B0CD38" italics="true" pageId="0" pageNumber="279">Larca boulderica</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName id="B6CFFB117E7C0A284891D7C725992748" lsidName="L. aalbui" pageId="0" pageNumber="279" rank="species" species="aalbui">
|
||
<emphasis id="2C06ED0046A81657AE5D10B219A7AA51" italics="true" pageId="0" pageNumber="279">L. aalbui</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DB013F8A4A6A8195E910669F3C3BEAF3" lsidName="L. rotunda" pageId="0" pageNumber="279" rank="species" species="rotunda">
|
||
<emphasis id="9C8E1308A81ADDEB9CC0FF999961BE73" italics="true" pageId="0" pageNumber="279">L. rotunda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="99B5CBBEA72CDEF983C182B07B60E7EE" lsidName="L. welbourni" pageId="0" pageNumber="279" rank="species" species="welbourni">
|
||
<emphasis id="551275E3434D691480B479B5A38FD1FA" italics="true" pageId="0" pageNumber="279">L. welbourni</emphasis>
|
||
</taxonomicName>
|
||
as all have four trichobothria on the movable chelal finger (Figs
|
||
<figureCitation id="7F6FC4C12A4E143386E91EB7A6671FE5" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">14</figureCitation>
|
||
,
|
||
<figureCitation id="1E0D4A25C094C95F888E5BED1D635EA3" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">15</figureCitation>
|
||
) and trichobothrium
|
||
<emphasis id="DF72908C5AD92A4D24C219A4EA3010C8" italics="true" pageId="0" pageNumber="279">ist</emphasis>
|
||
is closer to
|
||
<emphasis id="EAF41D5E2ED0E7944870B3F37DABE295" italics="true" pageId="0" pageNumber="279">ib</emphasis>
|
||
than to
|
||
<emphasis id="7724BBACBABA76F136CBFD8F1EBE78E6" italics="true" pageId="0" pageNumber="279">isb</emphasis>
|
||
(Figs
|
||
<figureCitation id="49324F01336E26B91A2A7CFBF08B6E98" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">14</figureCitation>
|
||
,
|
||
<figureCitation id="7BB2DEAB8EA6743CE9CB5525A5C31DA4" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">15</figureCitation>
|
||
). The only other species with four trichobothria,
|
||
<taxonomicName id="1DC250A561413B29638B31ADD9235772" lsidName="L. cavicola" pageId="0" pageNumber="279" rank="species" species="cavicola">
|
||
<emphasis id="D39E470263A0F0852FFE3F3A80BC844F" italics="true" pageId="0" pageNumber="279">L. cavicola</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AC5119CDA21D54DD6A2B4945C7F964DB" lsidName="L. guadalupensis" pageId="0" pageNumber="279" rank="species" species="guadalupensis">
|
||
<emphasis id="7C501A543A26280D9F0ACA49DB5DAA89" italics="true" pageId="0" pageNumber="279">L. guadalupensis</emphasis>
|
||
</taxonomicName>
|
||
, have trichobothrium
|
||
<emphasis id="4F3C210AA5C5648EC8EA8A9B55E085B2" italics="true" pageId="0" pageNumber="279">ist</emphasis>
|
||
situated midway between
|
||
<emphasis id="81BB6C82E6891B2A3C86F48EEE8CEF56" italics="true" pageId="0" pageNumber="279">ib</emphasis>
|
||
and
|
||
<emphasis id="855B93BC8DC8E5B2D980B30C96F515DD" italics="true" pageId="0" pageNumber="279">
|
||
isb.
|
||
<taxonomicName id="C7F7BC6085EAA010DFA7C15E0E4FFF01" authorityName="Chamberlin" authorityYear="1930" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca" order="Pseudoscorpionida" pageId="0" pageNumber="279" phylum="Arthropoda" rank="genus">Larca</taxonomicName>
|
||
<taxonomicName id="6E38B5848F41C237B95A7E09E134D119" authorityName="Hoff & Clawson" authorityYear="1952" class="Arachnida" family="Larcidae" genus="Archeolarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archeolarca rotunda" order="Pseudoscorpiones" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="rotunda">Archeolarca rotunda</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="FBB68A14CBF8B62AD1578BBE5A664863" lsidName="L. welbourni" pageId="0" pageNumber="279" rank="species" species="welbourni">
|
||
<emphasis id="3859A757F3150D03B796C7CFB7E12F76" italics="true" pageId="0" pageNumber="279">L. welbourni</emphasis>
|
||
</taxonomicName>
|
||
have a rounded chelal hand, whereas
|
||
<taxonomicName id="3344F218B7D15716F920A97F64D64097" lsidName="L. aalbui" pageId="0" pageNumber="279" rank="species" species="aalbui">
|
||
<emphasis id="C33A8FE3E87911D30E8608872C971250" italics="true" pageId="0" pageNumber="279">L. aalbui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5F7B6B3E3ADD95F0685C1C2D9A8A8C46" lsidName="L. boulderica" pageId="0" pageNumber="279" rank="species" species="boulderica">
|
||
<emphasis id="9308F67E1DCC7DB8B724C732A256253C" italics="true" pageId="0" pageNumber="279">L. boulderica</emphasis>
|
||
</taxonomicName>
|
||
have a thinner hand (Figs
|
||
<figureCitation id="463C24B33DBAA45A67F9D537321F43F6" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">12</figureCitation>
|
||
,
|
||
<figureCitation id="4BEA251B3092F282A5C07DF6F8F79047" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">13</figureCitation>
|
||
).
|
||
<taxonomicName id="BDB4034BFEA7C45C5A5A122357F8B24D" authorityName="Harvey & Steinmann" authorityYear="2024" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca boulderica" order="Pseudoscorpionida" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="boulderica">
|
||
<emphasis id="293A0DC58FB49B5D3C5E6442270E9BB1" italics="true" pageId="0" pageNumber="279">Larca boulderica</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C881048B99E0C6E9864FC52446EDB0F" lsidName="L. aalbui" pageId="0" pageNumber="279" rank="species" species="aalbui">
|
||
<emphasis id="F202C17CF217401D138C43C6CB2E9BE3" italics="true" pageId="0" pageNumber="279">L. aalbui</emphasis>
|
||
</taxonomicName>
|
||
by the positions of trichobothria
|
||
<emphasis id="31CFCFAB9F0605149DB8DDC8D79F75F8" italics="true" pageId="0" pageNumber="279">st</emphasis>
|
||
and
|
||
<emphasis id="EA3D65C28F623BD28C7790F10834C7EB" italics="true" pageId="0" pageNumber="279">t</emphasis>
|
||
(separated by at least one areolar diameter in
|
||
<taxonomicName id="F41E4A404B3B8794005710D574DED4F6" lsidName="L. boulderica" pageId="0" pageNumber="279" rank="species" species="boulderica">
|
||
<emphasis id="FC2E09D4CA3DB17858F78593C1498C25" italics="true" pageId="0" pageNumber="279">L. boulderica</emphasis>
|
||
</taxonomicName>
|
||
but by less than one areolar diameter in
|
||
<taxonomicName id="EB9779A4FCD27170E3A23411B90C3350" lsidName="L. aalbui" pageId="0" pageNumber="279" rank="species" species="aalbui">
|
||
<emphasis id="2722325E7067F3CFB800494DCBE9A040" italics="true" pageId="0" pageNumber="279">L. aalbui</emphasis>
|
||
</taxonomicName>
|
||
), and the less slender pedipalpal segments [e.g., 4.57-4.59 (♂), 4.07-4.71 (♀)
|
||
<normalizedToken id="EB70A2E1E3D007DA68C6A468ED5E3699" originalValue="×">x</normalizedToken>
|
||
and patella 3.09-3.22 (♂), 3.21-3.32 (♀)
|
||
<normalizedToken id="1B4F5E226AD6DCDE041634D1A99DE0C4" originalValue="×">x</normalizedToken>
|
||
longer than broad in
|
||
<taxonomicName id="04E512F4CE0694650BA59BF7AA531D93" lsidName="L. boulderica" pageId="0" pageNumber="279" rank="species" species="boulderica">
|
||
<emphasis id="A15954140C18592A70D24DA9F103CC1B" italics="true" pageId="0" pageNumber="279">L. boulderica</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation id="070C2F3DD3983442A971BC9FCE20E218" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">12</figureCitation>
|
||
,
|
||
<figureCitation id="9D03D8018EB1D6092E2CABFC98E526A3" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">13</figureCitation>
|
||
); femur 5.3-5.9
|
||
<normalizedToken id="E8BA7E7463524655B2C6541974C397E3" originalValue="×">x</normalizedToken>
|
||
and patella 3.9-4.35
|
||
<normalizedToken id="CE0FFFE2715CC866A01558034C3297C3" originalValue="×">x</normalizedToken>
|
||
longer than broad in
|
||
<taxonomicName id="A5B2227FAA670433680C65A07CDC4D27" lsidName="L. aalbui" pageId="0" pageNumber="279" rank="species" species="aalbui">
|
||
<emphasis id="D80B3F643D2CB3F764BDFBFAEDA4DB7E" italics="true" pageId="0" pageNumber="279">L. aalbui</emphasis>
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CB44B9BDF3D658E5859A6E0BD637F6DA" pageId="0" pageNumber="279" type="description">
|
||
<paragraph id="3CFC24BFE94D38E9DC3C1F21B20B01D0" pageId="0" pageNumber="279">Description</paragraph>
|
||
<paragraph id="5B9AE36AB0E1FBD1E3A41C7E6C57AD04" pageId="0" pageNumber="279">
|
||
<emphasis id="7152CEF92BFFDFE1FA9E29F797881E99" bold="true" pageId="0" pageNumber="279">
|
||
(adults).
|
||
<emphasis id="3D830DE6EBBE5E79A7B44638ED79264D" italics="true" pageId="0" pageNumber="279">Colour</emphasis>
|
||
</emphasis>
|
||
: most body parts pale yellow-brown, genital region of female and legs slightly paler (Figs
|
||
<figureCitation id="C63165D9A70CE3BD00DC2041D7DDB80E" captionStart="Figure 6" captionStartId="F4" captionText="Figure 6. Larca boulderica sp. nov., in situ in Mallory Cave." figureDoi="10.3897/zookeys.1198.120353.figure6" httpUri="https://binary.pensoft.net/fig/1033627" pageId="0" pageNumber="279">6</figureCitation>
|
||
-
|
||
<figureCitation id="DA37E410AD9F7914BE1DD6A453EAAF38" captionStart="Figures 7–9" captionStartId="F5" captionText="Figures 7 - 9. Larca boulderica sp. nov., paratype female (DMNS): 7 body, dorsal 8 body, ventral 9 cephalothorax, dorsal. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1198.120353.figures7-9" httpUri="https://binary.pensoft.net/fig/1033628" pageId="0" pageNumber="279">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="5B06180FD99EF0ED71251E087443DF82" doi="10.3897/zookeys.1198.120353.figure6" httpUri="https://binary.pensoft.net/fig/1033627" pageId="0" pageNumber="279" start="Figure 6" startId="F4">
|
||
<paragraph id="8E692C4366EAFA3F79E52C6CBA20FB58" pageId="0" pageNumber="279">
|
||
<emphasis id="022BB3558D58FF038C7206A914E73D8B" bold="true" pageId="0" pageNumber="279">Figure 6.</emphasis>
|
||
<taxonomicName id="B1F3BF50B9D406866D7B237BB6448BBB" authorityName="Harvey & Steinmann" authorityYear="2024" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca boulderica" order="Pseudoscorpionida" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="boulderica">
|
||
<emphasis id="13280C2DD589D40E9AC154E09521FEB8" italics="true" pageId="0" pageNumber="279">Larca boulderica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., in situ in Mallory Cave.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B1530B78F705FDF6622BAABC73E45623" doi="10.3897/zookeys.1198.120353.figures7-9" httpUri="https://binary.pensoft.net/fig/1033628" pageId="0" pageNumber="279" start="Figures 7–9" startId="F5">
|
||
<paragraph id="07F21F6A6DAA1DFD9E2D8B86B70A7053" pageId="0" pageNumber="279">
|
||
<emphasis id="535D1EE0F439D9A525A01E7A8073E99D" bold="true" pageId="0" pageNumber="279">Figures 7-9.</emphasis>
|
||
<taxonomicName id="7474348F300E41B5B64811632E5AEE97" authorityName="Harvey & Steinmann" authorityYear="2024" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca boulderica" order="Pseudoscorpionida" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="boulderica">
|
||
<emphasis id="371F929734B6C1D33BD2B0A6307973D9" italics="true" pageId="0" pageNumber="279">Larca boulderica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype female (DMNS):
|
||
<emphasis id="36DA5458D8782CF0F77B2A86937483A6" bold="true" pageId="0" pageNumber="279">7</emphasis>
|
||
body, dorsal
|
||
<emphasis id="2C121C6626070A53600CD2C86146F9A1" bold="true" pageId="0" pageNumber="279">8</emphasis>
|
||
body, ventral
|
||
<emphasis id="813EE5228476131F22A1A9735DB6B41C" bold="true" pageId="0" pageNumber="279">9</emphasis>
|
||
cephalothorax, dorsal. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="22C2D116DACD55C55B9ED6E563FA410A" pageId="0" pageNumber="279">
|
||
<emphasis id="C7AEE4B65916858CD3EDB8C980C610CF" bold="true" italics="true" pageId="0" pageNumber="279">Setae and cuticle</emphasis>
|
||
: setae long, usually curved, distally acuminate; most cuticular surfaces granulate.
|
||
</paragraph>
|
||
<paragraph id="AADB9E613FC844F9D4B7E62028737F41" pageId="0" pageNumber="279">
|
||
<emphasis id="89BE2215490E9C819C9ED59AF05CE058" bold="true" italics="true" pageId="0" pageNumber="279">Chelicera</emphasis>
|
||
: with 4 (rarely 5) setae on hand, with
|
||
<emphasis id="F9C5E1AA7E9B874E0593B5F9947F5844" italics="true" pageId="0" pageNumber="279">sbs</emphasis>
|
||
usually absent, and 1 subdistal seta on movable finger; all setae acuminate; seta
|
||
<emphasis id="8A4AD57959790322A59DBAF3FAF796B3" italics="true" pageId="0" pageNumber="279">bs</emphasis>
|
||
slightly shorter than others; with 2 dorsal lyrifissures and 1 ventral lyrifissure; galea of ♂ short with terminal bifurcation (Fig.
|
||
<figureCitation id="2F44E6C97065F021E13A5CE3A506574C" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">16</figureCitation>
|
||
), of ♀ long and slender with 3 terminal to subterminal rami (Fig.
|
||
<figureCitation id="3E669D1CDF7265C745863D654CB741EA" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">17</figureCitation>
|
||
); rallum of 4 blades, the most distal blade with several serrations on leading edge, other blades smooth; serrula exterior with 16 (♂), 18 (♀) blades; lamina exterior present.
|
||
</paragraph>
|
||
<paragraph id="88B0FEA73227FD9330C6E4F9A1718450" pageId="0" pageNumber="279">
|
||
<emphasis id="1AB77A99442BD329E216CA18A7799511" bold="true" italics="true" pageId="0" pageNumber="279">Pedipalps</emphasis>
|
||
: Pedipalp (Figs
|
||
<figureCitation id="93B904CD7F55E23B5474B368640B941E" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">12</figureCitation>
|
||
,
|
||
<figureCitation id="99B96A3C729FF0F21315EEEDBDFFA4AF" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">13</figureCitation>
|
||
): most surfaces of trochanter, femur, patella and chelal hand lightly and granulate, chelal fingers smooth; trochanter, femur, patella and chelal hand with prominent, curved, slightly denticulate setae arranged sparsely; patella with 3 small sub-basal lyrifissures; trochanter 1.85 (♂), 1.79 (♀), femur 4.57-4.59 (♂), 4.07-4.71 (♀), patella 3.09-3.22 (♂), 3.21-3.32 (♀), chela (with pedicel) 4.36-5.00 (♂), 3.83-4.03 (♀), chela (without pedicel) 4.09-4.78 (♂), 3.60-3.81 (♀), hand (with pedicel) 2.09-2.33 (♂), 1.83-1.87 (♀)
|
||
<normalizedToken id="4C84241868B178A2CC25949610281A2D" originalValue="×">x</normalizedToken>
|
||
longer than broad, movable finger (with pedicel) 0.96-1.05 (♂), 0.97-1.03 (♀)
|
||
<normalizedToken id="DEAD56951395A8FC04C7C7356C2542E6" originalValue="×">x</normalizedToken>
|
||
longer than hand. Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria (Figs
|
||
<figureCitation id="4348BF1CB1031F7EF7990C6D06590C6D" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">14</figureCitation>
|
||
,
|
||
<figureCitation id="BBC4E9856B3491E993541462A1FF1DC3" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">15</figureCitation>
|
||
):
|
||
<emphasis id="11D3E4AF475233C069CD47A789543851" italics="true" pageId="0" pageNumber="279">eb</emphasis>
|
||
,
|
||
<emphasis id="A24E897CAE005561B2B083C440B535A2" italics="true" pageId="0" pageNumber="279">esb</emphasis>
|
||
,
|
||
<emphasis id="608ED3A9101D876CEA5E0FC4C43BA364" italics="true" pageId="0" pageNumber="279">ib</emphasis>
|
||
and
|
||
<emphasis id="8667CEE90EB3B92DB8284CB92DD8ED3E" italics="true" pageId="0" pageNumber="279">ist</emphasis>
|
||
situated subbasally,
|
||
<emphasis id="5991D92D63C28B54C87F2B95E630089B" italics="true" pageId="0" pageNumber="279">est</emphasis>
|
||
,
|
||
<emphasis id="FF5F71C7C7E3FC4EF89A37FD21BB6CE8" italics="true" pageId="0" pageNumber="279">isb</emphasis>
|
||
and
|
||
<emphasis id="CAEE5F56D609100498687E2145202328" italics="true" pageId="0" pageNumber="279">it</emphasis>
|
||
submedially,
|
||
<emphasis id="0D3890841BCED8EC96D68FE98CC47FF1" italics="true" pageId="0" pageNumber="279">et</emphasis>
|
||
subdistally,
|
||
<emphasis id="31B9BE0F1A0D549923A373DCFD07AB13" italics="true" pageId="0" pageNumber="279">est</emphasis>
|
||
slightly distal to
|
||
<emphasis id="DD857EAFF6BBE249B16B56A031B3708D" italics="true" pageId="0" pageNumber="279">it</emphasis>
|
||
,
|
||
<emphasis id="DA072FCA0647B615498E6E031C7FF69C" italics="true" pageId="0" pageNumber="279">ib</emphasis>
|
||
opposite
|
||
<emphasis id="CFC6D0031FE78FBFD31F9AD623C1DCF1" italics="true" pageId="0" pageNumber="279">esb</emphasis>
|
||
, and
|
||
<emphasis id="38E0AB21E5A52845450EFD1C5BA7F965" italics="true" pageId="0" pageNumber="279">ist</emphasis>
|
||
distal to
|
||
<emphasis id="7E2E80D7C60B3DBEBB21DC4AC5905EC1" italics="true" pageId="0" pageNumber="279">esb</emphasis>
|
||
;
|
||
<emphasis id="E522A25D82BCE11B0C82FB9E9A609A5C" italics="true" pageId="0" pageNumber="279">b</emphasis>
|
||
and
|
||
<emphasis id="CFE75CF77470ADD1D1EFB0D4A5EF1DBA" italics="true" pageId="0" pageNumber="279">sb</emphasis>
|
||
situated subbasally, and
|
||
<emphasis id="D45AE9D49A04DF96C0D7D9B1A2DB9965" italics="true" pageId="0" pageNumber="279">st</emphasis>
|
||
and
|
||
<emphasis id="59C405D7BDA1CEA3DFE91B275773A6A4" italics="true" pageId="0" pageNumber="279">t</emphasis>
|
||
situated submedially, with
|
||
<emphasis id="C094463DA032372F60E2AB22ED797DD5" italics="true" pageId="0" pageNumber="279">st</emphasis>
|
||
situated very close to
|
||
<emphasis id="9267393628A3D50FDAA3442A9FE69F86" italics="true" pageId="0" pageNumber="279">t</emphasis>
|
||
, separated by slightly more than 1 areolar diameter; patch of microsetae not present on retrolateral margin of fixed chelal finger near et. Venom apparatus present in both chelal fingers, venom ducts not visible. Chelal teeth rounded, very low; fixed finger with 30 (♂), 30 (♀) teeth; movable finger with 29 (♂), 28 (♀) teeth; accessory teeth absent.
|
||
</paragraph>
|
||
<paragraph id="DC1D63DB09A048C035E1C8706306469F" pageId="0" pageNumber="279">
|
||
<emphasis id="9EDD391B2DD0EF2BDDE487FB9C84321E" bold="true" italics="true" pageId="0" pageNumber="279">Cephalothorax</emphasis>
|
||
: carapace (Figs
|
||
<figureCitation id="BCAD763E503B1A5EA5560795B812459E" captionStart="Figures 7–9" captionStartId="F5" captionText="Figures 7 - 9. Larca boulderica sp. nov., paratype female (DMNS): 7 body, dorsal 8 body, ventral 9 cephalothorax, dorsal. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1198.120353.figures7-9" httpUri="https://binary.pensoft.net/fig/1033628" pageId="0" pageNumber="279">9</figureCitation>
|
||
,
|
||
<figureCitation id="83B10AD2B1E8FC629B67346E4DBD4CE8" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">10</figureCitation>
|
||
): 0.73-0.75 (♂), 0.79-0.83 (♀)
|
||
<normalizedToken id="9902BB63EDAEE4CA63CBA1409E10B86A" originalValue="×">x</normalizedToken>
|
||
longer than broad; anterior margin straight; with 2 pairs of rounded corneate eyes, tapetum present; with 25-41 (♂), 24-30 (♀) setae, arranged with 6-8 (♂), 4-6 (♀) near anterior margin, 8-17 (♂), 10-15 (♀) in prozone, 7-8 (♂), 7 (♀) in metazone and 4-6 (♂), 3-4 (♀) near posterior margin; with 1 deep, broad median furrow. Coxal region: manducatory process rounded with 1 distal seta, 1 small sub-oral seta, and 12 (♂), 9 (♀) additional setae; median maxillary lyrifissure large, rounded and situated submedially; posterior maxillary lyrifissure rounded. Coxae I to IV becoming progressively wider. Chaetotaxy of coxae I-IV: ♂, 8: 8: 8: 12; ♀, 8: 8: 7: 11.
|
||
</paragraph>
|
||
<paragraph id="8E5DDDCBA0ACE5781BF6301C8FC9E0AD" pageId="0" pageNumber="279">
|
||
<emphasis id="06B2B38FAD83D4B30816551D0710DD21" bold="true" italics="true" pageId="0" pageNumber="279">Legs</emphasis>
|
||
: femora I and II longer than patellae; junction between femora and patellae III and IV very angulate; femora III and IV much smaller than patellae III and IV; femur + patella of leg IV 5.21 (♂), 4.81 (♀)
|
||
<normalizedToken id="F337F9FF60CA4F4452543C184FC43E00" originalValue="×">x</normalizedToken>
|
||
longer than broad (Fig.
|
||
<figureCitation id="0019B8E1EC40D3F71B853DC03C61B3F3" captionStart="Figures 10–18" captionStartId="F6" captionText="Figures 10 - 18. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 10 carapace, dorsal, male 11 left pair of eyes, dorsal, male 12 right pedipalp, dorsal, male 13 right pedipalp, dorsal, male 14 left chela, retrolateral, male 15 left chela, retrolateral, female 16 left galea, dorsal, male 17 left galea, ventral, female 18 left leg IV, retrolateral, male. Scale bars: 0.5 mm (12, 13); 0.25 mm (10, 14, 15, 18); 0.1 mm (11); 0.05 mm (16, 17)." figureDoi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279">18</figureCitation>
|
||
); metatarsi and tarsi not fused; tarsus IV without tactile seta; subterminal tarsal setae arcuate and acuminate; claws simple; arolium much longer than claws, not divided.
|
||
</paragraph>
|
||
<caption id="CA1A65B683A362F8BF05D22CDD390D7E" doi="10.3897/zookeys.1198.120353.figures10-18" httpUri="https://binary.pensoft.net/fig/1033629" pageId="0" pageNumber="279" start="Figures 10–18" startId="F6">
|
||
<paragraph id="34A8E1A47041F64A8B8F587DF206D336" pageId="0" pageNumber="279">
|
||
<emphasis id="7E38C62CFA745FB6E58B73F5D2FB2F01" bold="true" pageId="0" pageNumber="279">Figures 10-18.</emphasis>
|
||
<taxonomicName id="FFC608E401CFB8B83774C5A7BE9208DB" authorityName="Harvey & Steinmann" authorityYear="2024" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca boulderica" order="Pseudoscorpionida" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="boulderica">
|
||
<emphasis id="6AB80A5923F68149FAC14B3C8BB99BE1" italics="true" pageId="0" pageNumber="279">Larca boulderica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male and paratype female (DMNS):
|
||
<emphasis id="6141B0589141C6E30C97A53C4C8E4AD4" bold="true" pageId="0" pageNumber="279">10</emphasis>
|
||
carapace, dorsal, male
|
||
<emphasis id="B28F332426032ADBFF8A2CC31531CCCB" bold="true" pageId="0" pageNumber="279">11</emphasis>
|
||
left pair of eyes, dorsal, male
|
||
<emphasis id="BDC01F4751AB23AFB8B6D5215350C493" bold="true" pageId="0" pageNumber="279">12</emphasis>
|
||
right pedipalp, dorsal, male
|
||
<emphasis id="D64B5548F5C84C961412055246ED7466" bold="true" pageId="0" pageNumber="279">13</emphasis>
|
||
right pedipalp, dorsal, male
|
||
<emphasis id="E29FBC6FF938D17F35029961C9416B80" bold="true" pageId="0" pageNumber="279">14</emphasis>
|
||
left chela, retrolateral, male
|
||
<emphasis id="34D7B93F1E300784019D4D9718F27514" bold="true" pageId="0" pageNumber="279">15</emphasis>
|
||
left chela, retrolateral, female
|
||
<emphasis id="084C78F74B009C381D101F78A4A96A98" bold="true" pageId="0" pageNumber="279">16</emphasis>
|
||
left galea, dorsal, male
|
||
<emphasis id="907DF739610FAAD66A4579D0ED06F582" bold="true" pageId="0" pageNumber="279">17</emphasis>
|
||
left galea, ventral, female
|
||
<emphasis id="E8DE653DF2CD327F7705A751E066FCBE" bold="true" pageId="0" pageNumber="279">18</emphasis>
|
||
left leg IV, retrolateral, male. Scale bars: 0.5 mm (
|
||
<emphasis id="C0936431C68DDCEC2F3035874406F274" bold="true" pageId="0" pageNumber="279">12, 13</emphasis>
|
||
); 0.25 mm (
|
||
<emphasis id="5034569E198AC15BDEC0A824D76176EC" bold="true" pageId="0" pageNumber="279">10, 14, 15, 18</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis id="02F9E7080385D397D8B81AE2DFE9CDAF" bold="true" pageId="0" pageNumber="279">11</emphasis>
|
||
); 0.05 mm (
|
||
<emphasis id="4015C41830C6FB5910373680B462721D" bold="true" pageId="0" pageNumber="279">16, 17</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC0B7CFB656B646790412B9FB099B12D" pageId="0" pageNumber="279">
|
||
<emphasis id="8D3B5AD8E2487436060A3C76B32E8E02" bold="true" italics="true" pageId="0" pageNumber="279">Abdomen</emphasis>
|
||
: tergites II-VIII and sternites IV-VIII of male and female with medial suture line fully dividing each sclerite. Tergal chaetotaxy: ♂, 7: 8: 8: 11: 11: 11: 11: 10: 6: 6 (arranged T4T): 6: 2; ♀, 4: 6: 10: 11: 12: 11: 12: 11: 10: 8 (arranged T6T): 4: 2; tergites I-X uniseriate. Sternal chaetotaxy: ♂, 22: (0) 7 [3 + 3] (0): (0) 21 (0): 8: 9: 8: 8: 6: 6: 4: 2; ♀, 13: (0) 12 (0): (0) 7 (0): 8: 7: 8: 9: 7: 6: 4: 2; sternites IV-X uniseriate; ♂ and ♀ sternite II with all setae situated near posterior margin (Figs
|
||
<figureCitation id="1A530BA950BA7A0C214A54EB30018575" captionStart="Figures 19–21" captionStartId="F7" captionText="Figures 19 - 21. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 19 genital sternites, ventral, male 20 genital sternites, ventral, female 21 genitalia, ventral, female. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1198.120353.figures19-21" httpUri="https://binary.pensoft.net/fig/1033630" pageId="0" pageNumber="279">19</figureCitation>
|
||
,
|
||
<figureCitation id="10FB75839F709B15B317C589D6B04AD5" captionStart="Figures 19–21" captionStartId="F7" captionText="Figures 19 - 21. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 19 genital sternites, ventral, male 20 genital sternites, ventral, female 21 genitalia, ventral, female. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1198.120353.figures19-21" httpUri="https://binary.pensoft.net/fig/1033630" pageId="0" pageNumber="279">20</figureCitation>
|
||
); most setae of male sternite III clustered near posterior margin (Fig.
|
||
<figureCitation id="15F392451665D5060F745B0AC2578DBB" captionStart="Figures 19–21" captionStartId="F7" captionText="Figures 19 - 21. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 19 genital sternites, ventral, male 20 genital sternites, ventral, female 21 genitalia, ventral, female. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1198.120353.figures19-21" httpUri="https://binary.pensoft.net/fig/1033630" pageId="0" pageNumber="279">19</figureCitation>
|
||
). Spiracles with helix. Anal plates (tergite XII and sternite XII) situated between tergite XI and sternite XI, and surrounded by desclerotized region of tergite XI and sternite XI; sternite XI with 26 (♂), 22 (♀) small lyrifissures. Pleural membrane finely wrinkled-plicate; without any setae.
|
||
</paragraph>
|
||
<paragraph id="63ECD6186AE3E172BAD2C3F060E40A60" pageId="0" pageNumber="279">
|
||
<emphasis id="E084B791AF061691639AFEDEE00F3978" bold="true" italics="true" pageId="0" pageNumber="279">Genitalia</emphasis>
|
||
: male: very similar to that described for
|
||
<taxonomicName id="F36D236E7DC53077F99A6854505C8554" lsidName="L. laceyi" pageId="0" pageNumber="279" rank="species" species="laceyi">
|
||
<emphasis id="7F1FCFB039B0611C17CFF00BC5D7587D" italics="true" pageId="0" pageNumber="279">L. laceyi</emphasis>
|
||
</taxonomicName>
|
||
Muchmore, 1981 by
|
||
<bibRefCitation id="7824C12E6E1CF1CFD5435A763E6F9145" author="Muchmore, WB" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="279" pagination="47 - 60" refId="B38" refString="Muchmore, WB, 1981. Cavernicolous species of Larca, Archeolarca and Pseudogarypus with notes on the genera, (Pseudoscorpionida, Garypidae and Pseudogarypidae). The Journal of Arachnology 9: 47 - 60" title="Cavernicolous species of Larca, Archeolarca and Pseudogarypus with notes on the genera, (Pseudoscorpionida, Garypidae and Pseudogarypidae)." volume="9" year="1981">Muchmore (1981)</bibRefCitation>
|
||
. Female with 1 pair of lateral cribriform plates and 2 median cribriform plates, one of which is larger than the other (Fig.
|
||
<figureCitation id="5F9F3093803D73381A7B601E3F114BDC" captionStart="Figures 19–21" captionStartId="F7" captionText="Figures 19 - 21. Larca boulderica sp. nov., holotype male and paratype female (DMNS): 19 genital sternites, ventral, male 20 genital sternites, ventral, female 21 genitalia, ventral, female. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1198.120353.figures19-21" httpUri="https://binary.pensoft.net/fig/1033630" pageId="0" pageNumber="279">21</figureCitation>
|
||
); spermathecae absent.
|
||
</paragraph>
|
||
<caption id="F0800C98FAA9593A2AB213CC30A55383" doi="10.3897/zookeys.1198.120353.figures19-21" httpUri="https://binary.pensoft.net/fig/1033630" pageId="0" pageNumber="279" start="Figures 19–21" startId="F7">
|
||
<paragraph id="774F84906CE8E287F8D8FB9AA75B60F9" pageId="0" pageNumber="279">
|
||
<emphasis id="246E3E4044301C6CDD573A598AC3C8B6" bold="true" pageId="0" pageNumber="279">Figures 19-21.</emphasis>
|
||
<taxonomicName id="DA5D5473F8F2FACFD197F2524EA80B42" authorityName="Harvey & Steinmann" authorityYear="2024" class="Arachnida" family="Larcidae" genus="Larca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larca boulderica" order="Pseudoscorpionida" pageId="0" pageNumber="279" phylum="Arthropoda" rank="species" species="boulderica">
|
||
<emphasis id="4080BC775995DC0CA2BA749B3CBA9AFF" italics="true" pageId="0" pageNumber="279">Larca boulderica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male and paratype female (DMNS):
|
||
<emphasis id="88A114496C52EC9869D57FA39D1A7FA8" bold="true" pageId="0" pageNumber="279">19</emphasis>
|
||
genital sternites, ventral, male
|
||
<emphasis id="9B171488E72EA47DAD490AF450B91FF0" bold="true" pageId="0" pageNumber="279">20</emphasis>
|
||
genital sternites, ventral, female
|
||
<emphasis id="3028CF01B56E8F9FD654511AF0557B5D" bold="true" pageId="0" pageNumber="279">21</emphasis>
|
||
genitalia, ventral, female. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC286C11065C271D3F9A74B1363E3FD2" pageId="0" pageNumber="279">
|
||
<emphasis id="9BD475D885EBA40130C45221E95975FB" bold="true" pageId="0" pageNumber="279">
|
||
<emphasis id="C98182C4688D461BAAA2D8AEC36DE789" italics="true" pageId="0" pageNumber="279">Dimensions</emphasis>
|
||
(mm)
|
||
</emphasis>
|
||
: Male: holotype, with 1 other male in parentheses (when measured): Body length (excluding chelicerae) 2.37 (2.25). Pedipalp: trochanter 0.370/0.200, femur 0.895/0.195 (0.845/0.185), patella 0.725/0.225 (0.650/0.210), chela (with pedicel) 1.200/0.275 (1.225/0.245), chela (without pedicel) length 1.200 (1.170), chelal hand (without pedicel) length 0.575 (0.570), movable finger length 0.550 (0.600). Carapace 0.640/0.850 (0.580/0.800), anterior eye diameter 0.055, posterior eye diameter 0.050. Leg IV: femur + patella 0.730/0.140, tibia 0.530/0.100, metatarsus 0.265/0.065, tarsus 0.250/0.055.
|
||
</paragraph>
|
||
<paragraph id="9F6C0B7E57379D9DE094F01D6C0EFBF4" pageId="0" pageNumber="279">Female: paratype, with 3 other females in parentheses (when measured): Body length (excluding chelicerae) 2.51 (2.51-2.74). Pedipalp: trochanter 0.375/0.210, femur 0.895/0.205 (0.895-0.990/0.205-0.220), patella 0.755/0.235 (0.765-0.795/0.230-0.240), chela (with pedicel) 1.205/0.315 (1.190-1.255/0.295-0.325), chela (without pedicel) length 1.135 (1.125-1.255), chelal hand (without pedicel) length 0.575 (0.550-0.595), movable finger length 0.565 (0.565-0.575). Carapace 0.640/0.815 (0.645-0.655/0.805-0.815), anterior eye diameter 0.065, posterior eye diameter 0.060. Leg IV: femur + patella 0.745/0.155, tibia 0.530/0.095, metatarsus 0.270/0.065, tarsus 0.250/0.055.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8D56782356D38B22CE1292F4DD0908AA" pageId="0" pageNumber="279" type="etymology">
|
||
<paragraph id="02D44AF93388AE1EA64B038E866119A7" pageId="0" pageNumber="279">Etymology.</paragraph>
|
||
<paragraph id="92F730E3CDA345DBD338C889713F07D6" pageId="0" pageNumber="279">The species epithet is a noun taken from the type locality of Boulder County, Colorado. Mallory Cave is situated on City of Boulder, Open Space and Mountain Parks property.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |